76

Gloria Mans 42(4-5) ' 76-93 i Antwerpen, november 2003

Four new (: ) from the Cape Verde Archipelago.

Koen FRAUSSEN (1) & Emilio ROLAN (2)

(1) Leuvensestraat 25, B-3200 Aarschot, Belgium, koen fraussen@skynet be (2) Canovas del Castillo, 22-5"?, 36202 Vigo, Espana, emiliorolan@inicia es

Keywords Euthria, Cape Verde Archipelago, phenotypic variability, new taxa

Abstract Four , collected in the Cape Verde Islands, are described as new and assigned to the isM/^na M E Gray, 1830 E bernardi sp nov E insalubns sp nov E abrotona sp nov and E cecilea sp nov The new species are compared with other taxa from the and the Cape Verde Archipelago We discuss the infraspecific variability of the genus and especially E bernardi and E cecilea

Introduction (Linnaeus, 1758), the type species of the genus, had for a long time been the only known species in the Atlantic Nearly a century and a half later Dautzenberg & Fischer (1906) described the first additional Atlantic species, in the eighties followed by Cosel (1982), Cosel & Bumay (1983) and Rolan (1985) and in the beginning of this year (2003) by Rolan, Monteiro & Fraussen These 10 addi­ tional species are all endemic to the Cape Verde Archipelago In the present paper we enlarge this fauna from 10 to 14 known species

The descriptions are based on material collected by local divers or fishermen and obtained by private collectors Pierre Bernard (France), who collected material in the Cape Verde Archipelago between 1992 and 1998 Guido T Poppe (Belgium), who obtained material from divers at Sao Vicente 77 Abbreviations: MNCN Museo Nacional de Ciencias Naturales, Madrid, Spain MNHN Museum national d'Histoire naturelle. Pans, France CER collection Emilio Rolan, Spain GP collection Guido T Poppe. Belgium KF collection Koen Fraussen, Belgium

Genus Euthria M E Gray, 1850

Type species by original designation "Fimis hgnanus Chiaje" (this is Fusus ligna- nus Lamarck, 1816, a junior synonym oi corneus Linnaeus, 1758) from the Mediterranean, Recent

For a discussion of the use of Euthna as a genus we refer to Shuto (1978), Beets (1986), Fraussen (1999 73) and Fraussen & Hadorn (1999 120-121) For a detailed comparison with radulae of related genera we refer to Rolan, Monteiro & Fraussen (2003)

Euthria bernardi sp nov (Figs 1-8)

Type Material Holotype (65 0 mm) (Figs 1-2) in MNHN Paratypes 1-4, 6, 8-9 in MNHN, paratype 5 in KF nr4150, paratype 7 in ER

Type Locality Cape Verde Archipelago The type material was obtained from fisher­ men, without exact information

Range Only known from the Cape Verde Archipelago

Description Shell 65 mm high, (paratypes from 45 9 up to 65 1 mm high), thick and heavy, solid Shape fusiform, slender, with rather short siphonal canal Ground colour white to pale yellow Pattern dense, reddish brown, small, angular dots, arranged in spirals, with 2 paler, broad spiral bands Paratypes highly variable, red brown to dark brown, varying from a dense pattern of small, angular dots, arranged in spirals (figs 3-4), corresponding with spiral cords, to a pattern with large, irregular blotches, arranged in axials, with 2 broad, pale spiral bands, occasionally showing some spirals with these small, angular dots inside the large blotches (figs 7-8) Other specimens 78 show a wide array of intermediates (figs 1-2, 5-6) Subsutural band ornamented with rather rectangular, mostly evenly sized, alternating dark brown and white blotches Tip of siphonal canal dark brown to purplish Protoconch rather bulbous, with 2 1/4 to 2 1/2 convex, slightly inflated whorls, smooth, whitish Teleoconch with about 8 whorls, abapically convex, subsutural concavity broad, strong

First teleoconch whorl with 3 weak spiral cords, interspaces slightly narrower Second whorl with 5 cords, interspaces becoming narrower Third whorl with 6-7 flat, spiral cords, abapically broad, gradually becoming narrower adapically, subsutural cord exceptionally broad Penultimate whorl smooth or with 8 or 9 spiral cords, body whorl smooth or with 18-25 broad, flat, spiral cords, of equal size except for broad subsutural cord, occasionally correspondmg with pattern, interspaces a fine groove

First teleoconch whorls with 9 or 10 strong axial ribs Interspaces of equal size, deep Second whorl with 11 axial ribs From third whorl on, ribs smoother and broad, or absent Body whorl rather smooth, with thick rib marking aperture All whorls cov­ ered with fine growth lines, adapically prosocline or occasionally signoid

Aperture ovate Outer lip thick, with 10-14 internal lirae Edge sharp, smooth Columella smooth with weak abapical fold, callus thin Siphonal canal short, broad, open, tip darker coloured Aperture together with siphonal canal slightly shorter than 1/2 of total shell length Penostracum, radula and unknown

Comparison Euthria bernardi sp nov is characterized by the large size, the heavy shell with high spire and strong subsutural concavity, the dark siphonal canal, and by the penultimate and body whorl which are usually covered by many spiral cords and dotted spiral lines Paratype 3 (figs 3-4) is atypical in shape by the broader and blunt base Regarding the strong growthmarks and breaks in the former whorls we consider this a traumatic variation For differences with E cecilea sp nov we refer to the comparison under that species

Etymology Enthiia bernaidi sp nov is named after the late Pierre Bernard, for his contributions to the knowledge of West African molluscs A long time resident of Gabon between 1972 and 1989, Pierre Bernard was a professor of statistics and eco­ nomics at the University of Libreville His interest for the local molluscan fauna cul­ minated in the book Seashells of Gabon / Coqmllages du Gabon During his subse- 79 quent stay in the Cape Verde Islands until 1992, he pursued his interests with con- chology and acquired the new Euthna species reported on in this paper Pierre Bernard passed away in 2001 while he was employed in Mauritania His wife Idalina Bernard donated his mollusc collection to MNHN

Euthria insalubris sp nov (Figs 9-10)

Type Material Holotype in MNHN

Type Locality Cape Verde Archipelago The type material was obtained from fish­ ermen, without exact information

Range Only known from the Cape Verde Archipelago, and most probably endemic to a part of the archipelago

Description Shell 30 9 mm in height, thick, solid Shape fusiform, slender, with rather short siphonal canal Colour white Protoconch 1 3 mm in diameter, consisting of 1 3/4 whorls 7 teleoconch whorls Subsutural slope as broad as half a whorl, weakly concave First teleoconch whorl with 4 fine, primary spiral cords, interspaces narrow Second and third whorl with 4 or 5 primary spiral cords, the narrow interspaces filled up with a fine, secondary spiral cord, resulting in 9 flat spirals altematingly broad and narrow Fourth whorl with 4 flat, broad, weak primary spiral cords, 1-3 secondary spirals between two adjacent primary cords Fifth whorl with 3 primary spiral cords, the sub­ sutural one much broader Body whorl smooth

First teleoconch whorl with 9 broad, axial ribs Interspaces of equal size, deep Second whorl with 10, third whorl with 12 broad axial ribs Ribs becoming weak and flat from fourth whorl on Body whorl rather smooth All whorls covered with fine, prosocline growth lines, occasionally signoid

Aperture ovate Outer lip thick, with 8 internal lirae Edge sharp, smooth, dark red­ dish brown Columella smooth with 1 weak adapical fold Callus rather thick, dark reddish brown on connection with outer lip and on siphonal canal Siphonal canal short, broad, open Aperture together with siphonal canal shorter than 1/2 of total shell length 80 Operculum corneous, semi ovate with terminal nucleus, thick, dark brown Periostracum and radula unknown

Comparison Euthna insalubns sp nov is characterized, and differs from the other species described in this paper, by the protoconch with lower number of whorls

E cahpso Cosel & Burnay, 1983 (figs 13-14) is similar in shape, but differs by the higher number of protoconch whorls, by a slightly lower number of teleoconch whorls in combination with a larger size, by the weaker subsutural concavity espe­ cially on the upper whorls by the presence of spiral cords on the upper whorls, by the presence of a pattern with dots, by the colour of the aperture which is white with some brown spots m the outer lip and by the siphonal canal which is not straight For differences with E abrotona sp nov we refer to the comparison under that species

Etymology Derived from the Latin insalubns (adjective) meaning "unhealthy" The white colour gives the shell "a pale face"

Euthria abrotona sp nov (Figs 11-12)

Type Material Holotype in MNHN

Tjpe Locality Cape Verde Archipelago The type material was obtained from fish­ ermen, without exact information

Range Only known from the Cape Verde Archipelago

Description Shell 40 1 mm in height, thick, solid Shape fusiform, slender, with rather short siphonal canal Colour dark lemon yellow Protoconch of holotype badly eroded, large, rather inflated, 1 7 mm in diameter, con­ sisting of 2 1/4 whorls 6 1/2 teleoconch whorls Suture adpressed to preceeding whorl subsutural slope as broad as half a whorl Surface covered with numerous fine spiral cords Second whorl with about 20 such 81 spirals, occasionally altematingly fine and slightly broader Penultimate whorl with more than 50 spirals, body whorl with more than 100 spirals

First and second teleoconch whorl both with 9 broad, low axial ribs Becoming lower in number and weaker on further whorls Penultimate whorl with only some hardly visible ribs Body whorl smooth All whorls covered with numerous fine growth lines, adapically widely curved towards aperture

Aperture ovate Outer lip white, thick, with 9 internal knobs, edge sharp Columella smooth, callus thin, white to yellowish, edge dark yellow Siphonal canal rather short, broad, open, curved to dorsum Aperture together with siphonal canal longer than 1/2 of total shell length Periostracum, radula and operculum unknown

Comparison Eiilhna abrotona sp nov is characterized, and differs from all other Recent Euthna species, by the fine spiral sculpture consisting of numerous fine spi­ ral cords, and by the numerous fine growth lines which are curved adapically

E insalubris sp nov is similar in shape, but differs by the lower number of proto- conch whorls, by a slightly higher number of teleoconch whorls in combination with a smaller size, by the straighter growth lines, by the presence of altematingly strong and finer spiral cords on the second to third whorl and by the smooth latter whorls (instead of covered with numerous fine spirals)

E cahpso Cosel & Bumay, 1983 (figs 13-14) is similar in shape, but differs by the weaker subsutural concavity especially on the upper whorls, by the presence of spiral cords on the upper whorls only (the latter whorls are smooth), by the presence of pat­ tern, by the colour of the aperture which is white with some brown spots in the outer lip, by the thinner lip, by the siphonal canal which is not straight

Etymology Euthna abrotona sp nov is derived from abiotonum (Latin, neuter), which IS Southernwood or The Lemon Plant This plant, not yellow in colour itself, produces yellow fruit The single shell we have for study is strikingly yellow, but maybe the species produces other colours too We hereby relativate the importance of the yellow colour, as, we cannot ensure the species is characterized by a yellow colour, until other specimens are found 82

Euthria cecilea sp nov (Figs 19-30)

Type Material Holotype (33 O mm) (Figs 19-20) in MNHN Paratypes 1-11 in GP, KFnr4151 and ER

Type Locality Cape Verde Archipelago Sao Vicente By divers 20-80 m deep

Range Only known from the type locality

Description Shell medium, 33 mm high (paratypes up to 39 5 mm high), thick, solid Shape fusiform, slender, with rather short siphonal canal Ground colour white, apex pale pink Pattern dark brown, with many irregular dots, crossed by fine, white spiral bands Paratypes varying from many irregular dots (fig 25-26), to a pattern with broad spiral bands and/or broad axial flecks (fig 23-24, 27-30), crossed by 0-8 white, narrow, spiral lines Subsutural band consisting of rather rectangular, mostly evenly sized, alternating dark brown and white blotches, which are larger in the forms with large flecks Tip of siphonal canal pale pink Protoconch rather bulbous, with 1 3/4 to 2 1/2 convex, rather inflated, occasionally slightly carinated whorls, smooth, pink Transition to teleoconch marked by a weak axial nb Teleoconch with 6-6 1/4 whorls, spire rather conical, with weak subsutural concav­ ity

First teleoconch whorl with 3 or 4 rather strong spiral cords, eroded in most speci­ mens, interspaces narrow, towards second whorl with an additional fine, secondary spiral cord between two adjacent primary cords Second whorl smooth, occasionally with 7-14 cords Third whorl smooth Paratype 6 with 18, paratype 7 with 27 spiral cords on third whorl First teleoconch whorls with 10 broad, axial ribs Interspaces of equal size, deep Second whorl with 11-13 broad axial ribs From third whorl on, ribs smoother and broad, or absent Body whorl rather smooth, with thick rib marking aperture All whorls covered with fine, mostly signoid, prosocline growth lines

Aperture ovate Outer lip thick, with 9-11 (occasionally 15) internal lirae Edge sharp, smooth Columella smooth with 1 weak abapical fold or denticle, callus thin Siphonal canal short, broad, open Aperture together with siphonal canal shorter than 1/2 of total shell length Operculum corneous, semi ovate with terminal nucleus, thick, dark brown Penostracum and radula unknown 83

Comparison Euthria cecilea sp nov is characterized by the pattern with dark brown colour together with a pink protoconch, by an occasionally pale pink siphonal canal and the smooth shell without spiral cords on the penultimate and body whorl

E calvpso Cosel & Bumay, 1983 (figs 13-14) differs by the higher spire and more slender shell, the smaller aperture which is white with some brown spots in the outer lip, the narrow siphonal canal, the weaker subsutural concavity, the paler colour and by the pattern with fewer spiral bands and a narrower subsutural band

E marianae Rolan, Monteiro & Fraussen, 2003 differs by the broader shape with slightly shorter siphonal canal, a higher number of spiral cords on the first whorl, and by the paler pattern

E cornea (Linnaeus, 1758), or at least some of the forms with dark brown pattern, can look similar to E cecilea sp nov especially to the form with large flecks, but dif­ fers by the smaller protoconch (0 8-1 0 mm) with a lower number of whorls (1 1/4) instead of 1 3/4 to 2 1/2), the axial ribs which are abapically strong but adapically weak, on the upper whorls, and by the operculum with a more pointed nucleus

E bernardi sp nov differs by the larger size, the slender shape with higher spire, the stronger subsutural concavity, the presence of spiral cords on the penultimate and body whorls (while E cecilea sp nov is usually smooth from the third whorl on) but the lower number of spiral cords on the second whorl, the growth lines with are most­ ly curved, (while in E cecilea sp nov the lines are mostly signoid), the pattern with smaller dots and more colour bands, and the dark coloured siphonal canal

Etymology Euthria cecilea sp nov is named after Cecile Hoskens (Belgium) Together with Guido Poppe she collected the typematerial in the Cape Verde Archipelago early this year

General remarks on the genus Although Euthria species are rather uniform in shape within a same single species (especially the presence, or absence, of a narrow or broad concave subsutural slope is a stable characteristic) colour, pattern and size can occasionally be variable

E boavistensis Cosel, 1982 (figs 15-18), from the Cape Verde Archipelago, and E ciimulata Fraussen c& Hadom, 2003 from New Caledonia, are variable m spire angle Species which are known to show considerable variation in colour pattern are E poppei Fraussen, 1999 from East Africa (uniform white, yellow or lightbrown, occa­ sionally with fine brownish spiral bands), E solifei Fraussen & Hadom, 2003 from 84

New Caledonia (from snow-white to dark yellow, often with narrow, dark yellow to orange spiral bands), and E cornea (Linnaeus, 1758) from the Mediterranean (from snow-white to dark brown, over all intergrades of greyish, yellow, orange, pink, pale brown, even greenish, uniform or with dots or axial strikes) To this list we can now add E cecilea sp nov and especially E bernardi sp nov We here illustrate this vari­ ation by a series of each (figs 1-8, 19-30) The extremities (a densely dotted form and a flecked form) could be described as separate species if no intermediates are avail­ able for comparison The variability in the pattern of E cecilea sp nov (showing more or less or no white lines in combination with darker blotches, these blotches ranging trom small dots to axial strikes) is also found in E aiacaneme Angas, 1873 and E javanica Fraussen & Dekker, 2002, both from the Indian Ocean

Species with rather moderately to high variation in adult size are E wallen (Ladd, 1976) from the Philippines (from 39 to 60 mm), E philpoppei Fraussen, 2002 from New Caledonia (from 22 to 30 mm), E scepta Fraussen & Hadorn, 2003 from New Caledonia (from 17 to 33 mm), and E cornea (Linnaeus, 1758) To this list we can now add E bernardi sp nov varying from 45 9 to 65 mm The Cape Verde Archipelago is famous for it's Conidae radiation We may assume for sure that the genus Euthna will not reach a similar high number of species or forms But, as the study of this genus and this archipelago are both still going on, we can imagine that more, still undiscovered, species are waiting to be presented to science

Acknowledgements The authors thank Mme Bernard Chevallier, Philippe Bouchet and Virginie Heros (France) for making available the material from the collection of the late Pierre Bernard Guido Poppe and Cecile Hoskens (Belgium) for procuring material they collected earlier this year Kevin Monsecour (Belgium) and Amaud Le Goff (MNHN, France) for digital photography David Monsecour (Belgium) for cor­ recting the English text

REFERENCES ANGAS G. T 1873 Description of eight new species of land and marine shells from various localities Proceedings of the Zoological Societ\ London \ ST3 182-184 BEETS C 1986 Notes on 5HCC//7«A(W, a reappraisal Suipta Geol 82 83-100 CosEL R. VON. 1982 Zwei neue Euthi M-Arten von den Kapverdischen Insein (Prosobranchia Buccinidae) Archivjw Molluskenkunde, 112(1-6) 157-163 CosEL R. VON & BLRNAV L. P. 1983 A new Euthna from deeper shelf of the Cape Verde Islands AichivfUr Molluskenkunde, 113 (1-6) 151-157 DAUTZENBERG PH. & FISCHER H. 1906 Mollusques provenant des dragages effectues a I'ouest de I'Afnque pendant les campagnes de S A S le Prince de Monaco, Resultats des Campagnes Scientifiques accomplies sur son yacht pai Albeit lei Piince Soiiveraw de 85

Monaco, "il 1-125, pis 1-5 FRAUSSEN K 1999 A new species oï Euthria from Mozambique ( Gastropoda Buccinidae) Gloria Mans 37(5-6) 71-76 FRAUSSEN K 2002 A new Euthria (Gastropoda Buccmidae) from Java (Indonesia) Gloria MarisAX (4-5) 105-111 FRAUSSEN K. 2002 A new Euthria (Gastropoda Buccinidae) from New Caledonia Glona Mans 41 (4-5) 70-75 FRAUSSEN K. & DEKKER H 2002 A new Euthria (Gastropoda Buccmidae) from Java Gloria A/ara40(6) 112-119 FRAUSSEN K. & HADORN R. 1999 Rediscovery of Fiisinus siihangulatus (von Martens, 1903) and description of a new Somahan Fusinus (Gastropoda Fasciolanidae), including some notes on the taxonomical position of the genus Kuroda & Habe, 1952 Vita Marina 46(3-4) 111-122 FRAUSSEN K. & HADORN R 2003 Six new Buccmidae (Mollusca Gastropoda) from New Caledonia Vovapev 4(2-3) 33-50 GRAY M. E. 1850 Figures of molluscous selected from various authors Vol 4 Longman, Brown, Green and Longmans, London pp 219 GuERREiRO A. & REINER F , 2000 Moluscos marinhos da ilha de S Vicente (Archipelago de Cabo Verde) Camara Municipal de Oeiras, Europress, Povoa de Santo Adriao 279 pp KURODA T. & HABE T. 1954 New Genera of Japanese Marine Gastropods Venus 18(2) 84- 97 RoLAN E 1985 A new ^MrAna of the Cape Verde Islands La Conchiglia, \1 {\90-\9\) 6-7 ROLANE. 1987 The species in the genus iurAna (Mollusca Gastropoda) Argonaula, 2(5-6) 291-308 RoLAN E , MONTEIRO, A. & FRAUSSEN, K , 2003 Four new Euthi la (Mollusca, Buccinidae) from the Cape Verde archipelago, with comments on the validity of the genus Ibenis, 21(1) 115-127 SHUTO T 1978 On the genera Siphonofusus and Euthria of the Indo-West Pacific Transactions and Proceedings of the Palaeontological Society Japan, New Series, HI 358- 369 86 Summarizing translation into Dutch/ Samenvattende Nederlandse vertaling

Dit artikel introduceert vier nieuwe Kaapverdische soorten in het genus Euthna (BUCCINIDAE) E bernardi sp nov, E insalubns sp nov , E abrotona sp nov en E ceciliae sp nov

E bernardi sp nov (tot 65mm) is een zware schelp met een hoge top De kleur is wit tot lichtgeel met een druk patron van roodbruine vlekken in spiraalbanden en twee blekere spiraalbanden Het uiteinde van het korte siphonaal kanaal is donkerbruin tot purper De witte protoconch is gezwollen en bestaat uit 2 1/4 tot 2 1/2 convexe omgangen De teleoconch bestaat uit 8 omgangen die met spiraalkoorden bezet zijn Het aantal hiervan neemt toe per omgang (3 op de bovenste omgang tot 18-25 op de laatste omgang) Op de eerste twee omgangen van de teleoconch is een axiale sculp­ tuur zichtbaar die op de latere omgangen afvlakt en verdwijnt op de laatste omgang Lip met 10-14 groeven, columella glad met een zwakke abapicale plooi

E insalubns sp nov (tot 31 mm) is wit De protoconch bestaat slechts uit 1 3/4 omgang, terwijl de teleoconch uit 7 omgangen bestaat De bovenste omgang van de teleoconch draagt 4 spiraalkoorden, de tweede tot de vierde 4-5 waartussen sec­ ondaire spiraalkoorden voorkomen, de vijfde omgang heeft slechts 3 primaire spi­ raalkoorden en de laatste omgang is glad De axiale sculptuur bestaat uit 9 ribben op de eerste omgang van de teleoconch Dit aantal neemt toe tot en met de derde omgang, maar verdere omgangen zijn gladder Lip met 8 groeven, columella met een adapicale plooi Callus dik, roodbruin nabij de buitenlip en het korte siphonaal kanaal

E abrotona sp nov (tot 40mm) is donker citroengeel De protoconch bestaat uit 2 1 /4 gezwollen omgang, terwijl de teleoconch uit 6 1/2 omgangen bestaat Alle omgangen dragen veel spiraalkoorden (van 20 op de eerste omgang tot meer dan 100 op de laat­ ste) De bovenste omgang van de teleoconch draagt 9 axiale ribben, verdere omgan­ gen zijn gladder Lip met 9 knobbelachtige uitsteeksels Columella glad Het dunne callus is wit tot geel met een donkergele rand Het siphonaal kanaal is naar de rugzi- jde van de schelp toe gebogen

E ceciliae sp nov (tot 40mm) is wit met een roze top en punt van het siphonaal kanaal Het patroon bestaat uit vele onregelmatige donkerbruine vlekken en fijne witte spiraalbanden De gezwollen protoconch omvat 1 3/4 tot 2 1/2 gladde omgan­ gen De overgang naar de teleoconch wordt gemarkeerd door een zwakke axiale rib De teleoconch zelf bestaat uit 6 tot 6 1/4 omgangen Axiale ribben komen voor op de eerste twee omgangen van de teleoconch, daarna worden ze gladder en de laatste omgang is vrij glad met een dikke ribbel aan de mondopening Lip met 9-15 groeven Columella glad met een zwakke adapicale plooi of uitsteeksel 87

A. Euthria cecilea sp. nov., protoconch of paratype 9. Scale bar: 1 mm.

B. Euthria bernardi sp. nov., protoconch of paratype 1. Scale bar: 1 mm.

^ ^ f -^^ 1 u y\ 1\\p) 4 ^F\U ^^) n\

C. Euthria insahibris sp. nov., protoconch of holotype. Scale bar: 1 mm. 88

1-8. Euthria bernardi sp. nov.. Cape Verde Islands. 1-2: holotype, 65.0 mm. 3-4: paratype 3, 61.2 mm. 5-6: paratype 6, 49.3 mm. 7-8: paratype 4, 45.9 mm. t»'

't-."TT » j I v^

*•*

91

9-10 Euthna insalubns sp nov, Cape Verde Islands holotype, 30 9 mm 11-12 Eiithiia ahotona sp nov. Cape Verde Islands, holotype, 40 1 mm 13-14 Eiithiia cahpso Cosel & Bumay, 1983, Cape Verde Islands, off Maio Island, holotype, 44 6 mm 15-18 Euthna boavistensis Cosel, 1982, Sao Vicente 15-16 260mm 17-18 slender form, 31 2 mm 92

19-30. Euthria cecilea sp. nov., Cape Verde Islands, Sao Vicente. 19-20: holotype, 33.0 mm. 21-22: paratype 2, 39.5 mm. 23: paratype 3, 34.0 mm. 24: paratype 4, 36.0 mm. 25-26: paratype 1, 36.6 mm. 27: paratype 5, 33.4 mm. 28: paratype 6. 37.4 mm. 29: paratype 7, 35.9 mm. 30: paratype 8, 37.0 mm. tni i' u •*»- 'i

.«>-« 'Ü:

^'

mi

m^s

Ü» 28