DOCSLIB.ORG

T T-A/Fj Dissertation Lu 1V11 Information Service

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
T T-A/Fj Dissertation Lu 1V11 Information Service INFORMATION TO USERS This reproduction was made from a copy of a manuscript sent to us for publication and microfilming. While the most advanced technology has been used to pho­ tograph and reproduce this manuscript, the quality of the reproduction is heavily dependent upon the quality of the material submitted. Pages in any manuscript may have indistinct print. In all cases the best available copy has been filmed. The following explanation of techniques is provided to help clarify notations which may appear on this reproduction. 1. Manuscripts may not always be complete. When it is not possible to obtain missing pages, a note appears to indicate this. 2. When copyrighted materials are removed from the manuscript, a note ap­ pears to indicate this. 3. Oversize materials (maps, drawings, and charts) are photographed by sec­ tioning the original, beginning at the upper left hand corner and continu­ ing from left to right in equal sections with small overlaps. Each oversize page is also filmed as one exposure and is available, for an additional charge, as a standard 35mm slide or in black and white paper format.'* 4. Most photographs reproduce acceptably on positive microfilm or micro­ fiche but lack clarity on xerographic copies made from the microfilm. For an additional charge, all photographs are available in black and white standard 35mm slide format.* •For more information about black and white slides or enlarged paper reproductions, please contact the Dissertations Customer Services Department. T T-A/fJ Dissertation lu 1V11 Information Service University Microfilms International A Bell & Howell Information Company 300 N. Zeeb Road, Ann Arbor, Michigan 48106 8618811 Liu, Ho-yih A MONOGRAPH OF THE GENUS AEONIUM (CRASSULACEAE- SEMPERVIVOIDEAE) The Ohio State University PH.D. 1986 University Microfilms International 300 N. Zeeb Road, Ann Arbor, Ml 48106 Copyright 1986 by Liu, Ho-yih All Rights Reserved rnxKessaBasmaBsssmmaammmmmmmmmmmmaemBtessat. PLEASE NOTE: In all cases this material has been filmed in the best possible way from the available copy. Problems encountered with this document have been identified here with a check mark V . 1. Glossy photographs or pages 2. Colored illustrations, paper or print 3. Photographs with dark background 4. Illustrations are poor copy 5. Pages with black marks, not original copy 6. Print shows through as there is text on both sides of page 7. Indistinct, broken or small print on several pages 8. Print exceeds margin requirements 9. Tightly bound copy with print lost in spine 10. Computer printout pages with indistinct print 11. Page(s) lacking when material received, and not available from school or author. 12. Page(s) seem to be missing in numbering only as text follows. 13. Two pages numbered . Text follows. 14. Curling and wrinkled pages 15. Dissertation contains pages with print at a slant, filmed as received 16. Other University Microfilms International ma A MONOGRAPH OF TOE GENUS (CKASSULACEAE-SEMPERVIVDIDEAE) DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of the Ohio State University By Ho-yih Liu ***** The Ohio State University 1986 Dissertation Conmittee: Dr. Daniel J. Crawford Dr. Katherine L. Gross Dr. Tod F. Stuessy Copyright by Ho-yih Liu 1986 ACKNOWLEDGMENTS Numerous people have assisted in the completion of this dissertation. Dr. Tod. F. Stuessy, my major professor, is greatly acknowledged for his encourgement and guidance throughout all phases of lry graduate study. Thanks alo go to my conmittee Drs. Daniel J. Crawford, Katherine L. Gross for their critical conments on the dissertation. I also wish to express my appreciation to many graduate students who through their stimulating discussions contributed to the study. Field work for this study was supported by NSF grant, Sigma Xi, and the Graduate School of the Ohio State University which are greatfully appreciated. I also wish to thank the curators of the numerous herbaria who loaned specimens for this study. Special thanks to my wife for her support. VITA December 3, 1952 Born - Tapei, Taiwan 1975 B. S., Botany, National Taiwan University, Taipei, Taiwan 1977 M. S., Botany, National Taiwan University, Taipei, Taiwan 1977-1978 Teacher, Biology, TUnghai University, Taichu, Taiwan 1978-1979 Assistant Researcher, Pharmacy, National Defense Medical Center, Taipei, Taiwan 1980-1985 Graduate Associate, Botany, the Ohio State University, Columbus, CXiio PUBLICATIONS Stuessy, T. F., and H.-7. Liu. 1983. Anatomy of the pericarp of Clibadium. Desmanthodium. and Ichthyothece (Compositae, Heliantheae) and systematic implications. Rhodora 85:213-221. Liu, H.-Y. 1984. Notes on Taiwan Cucurbitaceae. Taiwania 28:117-121. FIELDS OF STUDY Major Field: Botany Studies in Plant Systematics. Professor Tod F. Stuessy TABLE OF CONTENTS PAGE ACKNOWLEDGMENTS ii VITA iii LIST OF TABLES V LIST OF FIGURES vi INTRODUCTION 1 TAXONOMIC HISTORY 3 SYSTEMATIC POSITION AND GENERIC RELATIONSHIPS 7 COMPARATIVE MORPHOLOGY AND ANATOMY 11 CYTOLOGY . 28 HYBRIDIZATION 35 PHENETICS 36 PHYLOGENETIC RECONSTRUCTION 50 PHYTOGEOGRAPHY . 55 ADAPTIVE RADIATION 63 TAXONOMIC TREATMENT 79 LITERATURE CITED 227 - IV - LIST OF TABLES TABLE PASE 1. Chromosome numbers in Aeonium 30 2. Characters and states used in the phenetic and cladistic analyses of Aeonium 39 3. Data matrix of taxa and characters of Aeonium for phenetic and cladistic study 41 - v - LIST OP FIGURES FIGURE PAGE 1. Nectariferous gland of Aichryson punctata™ .... 8 2. Nectariferous gland of Aeonium arhoreum var. arboreum 8 3. Placentation of Greenovia diplocycla 8 4. Stem reticulation in Aeonium section Leuconium 15 5. Multicellular trichome in Aeonium lindleyi var. lindleyi 15 6. Unicellular trichome in Aeonium ciliatum 15 7. Unicellular trichome in Aeonium glandulosum ... 15 8. Phenograra of taxa of Aeonium using UPGMA cluster analysis 45 9. Principal coordinates analysis of taxa of Aeonium showing three-dimensional projection of OTOs on axes representing the first three factors 47 10. Cladograra of taxa of the genus Aeonium based on parsimony 53 11. Geographical distribution of genera of the subfamily Sempervivoideae 56 12. Major wind directions near the Canary Islands . 59 13. Topography of Tenerife 67 14. Altidudinal distribution of taxa of Aeonium on Tenerife 69 \ 15. Different "ecological strategies" of taxa of Aeonium on Tenerife. 71 16. Growth forms of Aeonium arranged in possible evolutionary sequences 77 - vi - 17. Distribution of section Petrothamnium 97 18. Distribution of section Chrysocome 120 19. Distribution of section Patinaria 138 20. Line drawing of Aeonium korneliuslemsii 149 21. Distribution of section Aeonium in eastern Canary Islands and Morocco 151 22. Distribution of section Aeonium in western Canary Islands 165 23. Distribution of section Megalonium 171 24. Distribution of section Pittonium in Macaronesia 177 25. Distribution of section Pittonium in eastern Africa and Yemen 185 26. Line drawing of Aeonium stuessyi 188 27. Distribution of section Leuconium in Gran Canaria, Hierro, Lanzarotef and la Palma 194 28. Distribution of section Leuconium in Ganera and Tenerife 203 29. Line drawing of Aeonium davidbranwellii 216 - vii - BrcSCDUCFIGN Aeonlum Webb & Berth. (Crassulaceae; Sempervivoideae) is a large, polymorphic, and ecologically diverse genus of 31 species and six varieties growing in Macaronesia (Canary Islands, Cape Verde Islands, Madeira, and Southern Morocco), eastern Africa, and Yemen. The genus has its center of diversity, both in number of species and in morphological variation, in the Canary Islands. Different growth forms are found within the genus, ranging from subacaulescent to shrubby, monocarpic to non-monocarpic, and including biennials and perennials in the archipelago. In addition, the Canarian species of Aeonium occupy a wide range of habitats, front subalpine cloud forests to coastal deserts. The relationship between growth form and habitat in Aeonium on the Canary Islands was extensively studied by Lems (1960). He suggested that growth forms within the genus were the result of adaptive radiation, and because of this spectacular variation, even coimented that the genus was comparable in many ways to Darwin's finches (Lems 1960). This study attracted much attention, and chemical (Eglinton et al. 1962), phytosociological (Voggenreiter 1974), and physiological (Losch & Happen 1981, 1983; Tenhunen et al. 1982; Losch 1984) investigations have been done in the last two decades in an attempt to examine this evolutionarily dramatic system - 1 - 2 in more detail. Besides previous interest in adaptive radiation within Aeonium. the attractive appearance of the' genus has drawn attention from horticulturists. In fact, the last monograph on Aeonium. contained within An Account of the Sempervivum Group (Praeger 1932), was the result of a request from the Royal Horticultural Society of London for the needs of its members. Since the treatment was prepared for gardeners, the taxonomic delimitation of the genus, its sections, and its contained species, is inadequate. Also, it provides almost no insight into evolutionary relationships among the taxa. The main objectives of the present study, therefore, are to provide a modern revision and phylogenetic reconstruction of Aeonium. Based on three months of extensive field examinations of native populations and a thorough morphological analysis of greenhouse material and herbarium specimens, the taxonomy and nomenclature of the genus at the sectional and
Load more

Recommended publications
  • H3.3 Macaronesian Inland Cliff
    European Red List of Habitats - Screes Habitat Group H3.3 Macaronesian inland cliff Summary The perennial vegetation of crevices and ledges of cliff faces in Macaronesia away from coastal salt-spray is of very diverse character - some, for example, dominated by succulents, others rich in ferns and bryophytes characteristic of shaded situations - and it includes several hundreds of taxa endemic to the archipelagoes. The main threats are mountaineering and rock climbing, outdoor sports and leisure activities, and construction of infrastructures such as roads and motorways. Synthesis There is no evidence of significant past reductions, either in the last 50 years or historically, and also future prospects are good, as no serious threats are envisaged, besides touristic/leisure activities and putative faulty environmental impact assessments that may overlook this habitat as valuable. Reduction in quantity, reduction in quality and criteria of geographic distribution yield the Least Concern (LC) category. In spite of the LC category, conservation policy and management should restrict to the maximum any threat to or reduction of the habitat, as it has a very high conservation value, because of high endemism of species and communities with many local and regional variaties. Overall Category & Criteria EU 28 EU 28+ Red List Category Red List Criteria Red List Category Red List Criteria Least Concern - Least Concern - Sub-habitat types that may require further examination Four general subtypes can be distinguished based on species composition and different ecological conditions. However, at present, no data are available to carry out an individual assessment of each of them. In the future, if detailed plot sampling has been carried out, such an evaluation may be possible.
    [Show full text]
  • The 2014 Golden Gate National Parks Bioblitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event
    National Park Service U.S. Department of the Interior Natural Resource Stewardship and Science The 2014 Golden Gate National Parks BioBlitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event Natural Resource Report NPS/GOGA/NRR—2016/1147 ON THIS PAGE Photograph of BioBlitz participants conducting data entry into iNaturalist. Photograph courtesy of the National Park Service. ON THE COVER Photograph of BioBlitz participants collecting aquatic species data in the Presidio of San Francisco. Photograph courtesy of National Park Service. The 2014 Golden Gate National Parks BioBlitz - Data Management and the Event Species List Achieving a Quality Dataset from a Large Scale Event Natural Resource Report NPS/GOGA/NRR—2016/1147 Elizabeth Edson1, Michelle O’Herron1, Alison Forrestel2, Daniel George3 1Golden Gate Parks Conservancy Building 201 Fort Mason San Francisco, CA 94129 2National Park Service. Golden Gate National Recreation Area Fort Cronkhite, Bldg. 1061 Sausalito, CA 94965 3National Park Service. San Francisco Bay Area Network Inventory & Monitoring Program Manager Fort Cronkhite, Bldg. 1063 Sausalito, CA 94965 March 2016 U.S. Department of the Interior National Park Service Natural Resource Stewardship and Science Fort Collins, Colorado The National Park Service, Natural Resource Stewardship and Science office in Fort Collins, Colorado, publishes a range of reports that address natural resource topics. These reports are of interest and applicability to a broad audience in the National Park Service and others in natural resource management, including scientists, conservation and environmental constituencies, and the public. The Natural Resource Report Series is used to disseminate comprehensive information and analysis about natural resources and related topics concerning lands managed by the National Park Service.
    [Show full text]
  • CACTUS COURIER Newsletter of the Palomar Cactus and Succulent Society
    BULLETIN NOVEMBER 2014 CACTUS COURIER Newsletter of the Palomar Cactus and Succulent Society Volume 60, Number 11 November 2014 The Meeting is the 4th Saturday NOVEMBER 22, 2014 Park Avenue Community Center 210 Park Ave Escondido, CA 92025 Noon!! Coffee!! Photo by Robert Pickett “Ethiopia – Plants, History, and Cultures” • • Gary James • • Gary James has been interested in succulent In recent years he has been traveling to succulent-rich plants for many years – both his grandmother and his parts of the world to observe plants in habitat. Seeing parents had large succulent gardens. Growing up in South them growing in their natural areas gives an observer a Pasadena allowed him to spend many days visiting the better idea of how to care for the plants in one’s Huntington Botanic Gardens – back when admission was collection. free! In 2000 he organized a tour of Ethiopia for a group of friends. They traveled all over the country and observed a number of wonderful plant habitats. Ethiopia is a fascinating country with a long history of having never been colonized by a European power. The country includes many interesting tribes in the Omo River Valley, intriguing monuments in the north, and unusual Christian churches in the Lalibela area. Theirs is a rich Moslem culture as well. The talk will be a general introduction to the variety of cultures, tribes, historic monuments, as well as a look at many of the unusual plants that are found throughout the country. vvvvvvvv Board Meeting • Plant Sales • Brag Plants • Exchange Table REFRESHMENTS Lorie Johansen Martha Hansen • • • YOUR NAME HERE! • • • Please think about bringing something to share – it makes the day more fun! And we have a reputation to uphold!! Plant of the Month • • Tylecodon • • Tylecodon is a genus of succulent plants in the family Crassulaceae.
    [Show full text]
  • Proceedings Amurga Co
    PROCEEDINGS OF THE AMURGA INTERNATIONAL CONFERENCES ON ISLAND BIODIVERSITY 2011 PROCEEDINGS OF THE AMURGA INTERNATIONAL CONFERENCES ON ISLAND BIODIVERSITY 2011 Coordination: Juli Caujapé-Castells Funded and edited by: Fundación Canaria Amurga Maspalomas Colaboration: Faro Media Cover design & layout: Estudio Creativo Javier Ojeda © Fundación Canaria Amurga Maspalomas Gran Canaria, December 2013 ISBN: 978-84-616-7394-0 How to cite this volume: Caujapé-Castells J, Nieto Feliner G, Fernández Palacios JM (eds.) (2013) Proceedings of the Amurga international conferences on island biodiversity 2011. Fundación Canaria Amurga-Maspalomas, Las Palmas de Gran Canaria, Spain. All rights reserved. Any unauthorized reprint or use of this material is prohibited. No part of this book may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, recording, or by any information storage and retrieval system without express written permission from the author / publisher. SCIENTIFIC EDITORS Juli Caujapé-Castells Jardín Botánico Canario “Viera y Clavijo” - Unidad Asociada CSIC Consejería de Medio Ambiente y Emergencias, Cabildo de Gran Canaria Gonzalo Nieto Feliner Real Jardín Botánico de Madrid-CSIC José María Fernández Palacios Universidad de La Laguna SCIENTIFIC COMMITTEE Juli Caujapé-Castells, Gonzalo Nieto Feliner, David Bramwell, Águedo Marrero Rodríguez, Julia Pérez de Paz, Bernardo Navarro-Valdivielso, Ruth Jaén-Molina, Rosa Febles Hernández, Pablo Vargas. Isabel Sanmartín. ORGANIZING COMMITTEE Pedro
    [Show full text]
  • Aeonium Webb & Berthelot in Gibraltar
    Comunicaciones 1 HE GENUS AEONIUM WEBB & BERTHELOT IN GIBRALTAR Brian M. Lamb /Conservador del Gibraltar Botanic Gardens The genus Aeonium Webb & Berthelot belongs to the family Crassulaceae often refered to as the Houseleek family. Some 32 species are endemic to the Canary Islands, two to the Cape Verde Islands, two to Madeira, one to Morocco and possibly today three on the other side of Africa from northem Kenya through to Arabia. The three species that can be seen in Gibraltar are: A. undulatum native to Gran Canaria A. haworthii native to Tenerife A. arboreum native to Morocco. The two Canarian Aeoniums have almost certainly been introduced by man, probably during the latter part of the 19th century, particularly A. ~indulatumwhich is usually only found as a cultivated plant grown only on a small scale. However, A. haworthii is to be found in a number of places in Gibraltar, as naturalised colonies growing from pockets or fissures in the limestone rock, particularly on the eastem side, southwards from Catalan Bay, while on the westem side a sizeable colony exists below Bleak House and a small one is now forming near Camp Bay, growing in association with native Sedum. A. haworthii seems to be quite a popular patio and verandah plant, as it is not a giant growing species, and has lovely red edged glaucous leaves. The flower colour is variable, from white or pale yellow through to rose-pink. The seeds ofAeonium are very small, as with most members of the family Crassulacease, produced in great abundance, and are easily distributed by the wind.
    [Show full text]
  • Checklist of the Vascular Alien Flora of Catalonia (Northeastern Iberian Peninsula, Spain) Pere Aymerich1 & Llorenç Sáez2,3
    BOTANICAL CHECKLISTS Mediterranean Botany ISSNe 2603-9109 https://dx.doi.org/10.5209/mbot.63608 Checklist of the vascular alien flora of Catalonia (northeastern Iberian Peninsula, Spain) Pere Aymerich1 & Llorenç Sáez2,3 Received: 7 March 2019 / Accepted: 28 June 2019 / Published online: 7 November 2019 Abstract. This is an inventory of the vascular alien flora of Catalonia (northeastern Iberian Peninsula, Spain) updated to 2018, representing 1068 alien taxa in total. 554 (52.0%) out of them are casual and 514 (48.0%) are established. 87 taxa (8.1% of the total number and 16.8 % of those established) show an invasive behaviour. The geographic zone with more alien plants is the most anthropogenic maritime area. However, the differences among regions decrease when the degree of naturalization of taxa increases and the number of invaders is very similar in all sectors. Only 26.2% of the taxa are more or less abundant, while the rest are rare or they have vanished. The alien flora is represented by 115 families, 87 out of them include naturalised species. The most diverse genera are Opuntia (20 taxa), Amaranthus (18 taxa) and Solanum (15 taxa). Most of the alien plants have been introduced since the beginning of the twentieth century (70.7%), with a strong increase since 1970 (50.3% of the total number). Almost two thirds of alien taxa have their origin in Euro-Mediterranean area and America, while 24.6% come from other geographical areas. The taxa originated in cultivation represent 9.5%, whereas spontaneous hybrids only 1.2%. From the temporal point of view, the rate of Euro-Mediterranean taxa shows a progressive reduction parallel to an increase of those of other origins, which have reached 73.2% of introductions during the last 50 years.
    [Show full text]
  • Changing Attitudes See Page 5
    LNewsletteret’s of the San DiegoT Horticulturalalk Society Plants!July 2008, Number 166 Changing Attitudes SEE PAGE 5 PASTA SALADS PAGE 7 Nepenthes PAGE 8 Creating a Zen Garden PAGE 9 Consider Conifers PAGE 11 On the Cover: Chanticleer: a pleasure garden IMAGES FROM THE 2008 San DIEGO COUNTY FaiR Photo by Amelia Lima Photo by Photo by Amelia Lima Photo by San Diego Horticultural Society Amelia Lima Photo by Photo by Amelia Lima Photo by Palomar Cactus and Succulent Society Photo by Amelia Lima Photo by Photo by Katie Pelisek Photo by Quail Botanical Gardens and the San Diego County Water Authority Fair Entry Garden 2 Let’s Talk Plants! July 2008, No. 166 San Diego Horticultural Society In This Issue... The San Diego Horticultural Society 4 Important Member Information Meetings 5 To Learn More... The San Diego Horticultural Society meets the 2nd Monday of every month (except June) from 5 Thanks Fair Volunteers and Donors 6:00pm to 9:00pm at the Surfside Race Place, Del Mar Fairgrounds, 2260 Jimmy Durante Blvd. 5 From The Board Meetings are open and all are welcome to attend. We encourage you to join the organization to enjoy free admission to regular monthly meetings, receive the monthly newsletter and 7 Plants That Produce numerous other benefits. We are a 501(c)(3) non-profit organization. 7 Garden Gourmet Meeting Schedule 8 Nepenthes: Common Ways to Grow an 5:00 – 6:00 Meeting room setup Uncommon Plant 6:00 – 6:45 Vendor sales, opportunity drawing ticket sales, lending library 8 Molecular Biology in the Garden: Part 11 6:45 – 8:00 Announcements, speaker, drawing for three plants 9 Welcome New Members! 8:00 – 8:15 Break for vendor sales, opportunity drawing ticket sales, lending library 8:15 – 9:00 Plant forum and opportunity drawing.
    [Show full text]
  • Plethora of Plants - Collections of the Botanical Garden, Faculty of Science, University of Zagreb (2): Glasshouse Succulents
    NAT. CROAT. VOL. 27 No 2 407-420* ZAGREB December 31, 2018 professional paper/stručni članak – museum collections/muzejske zbirke DOI 10.20302/NC.2018.27.28 PLETHORA OF PLANTS - COLLECTIONS OF THE BOTANICAL GARDEN, FACULTY OF SCIENCE, UNIVERSITY OF ZAGREB (2): GLASSHOUSE SUCCULENTS Dubravka Sandev, Darko Mihelj & Sanja Kovačić Botanical Garden, Department of Biology, Faculty of Science, University of Zagreb, Marulićev trg 9a, HR-10000 Zagreb, Croatia (e-mail: [email protected]) Sandev, D., Mihelj, D. & Kovačić, S.: Plethora of plants – collections of the Botanical Garden, Faculty of Science, University of Zagreb (2): Glasshouse succulents. Nat. Croat. Vol. 27, No. 2, 407- 420*, 2018, Zagreb. In this paper, the plant lists of glasshouse succulents grown in the Botanical Garden from 1895 to 2017 are studied. Synonymy, nomenclature and origin of plant material were sorted. The lists of species grown in the last 122 years are constructed in such a way as to show that throughout that period at least 1423 taxa of succulent plants from 254 genera and 17 families inhabited the Garden’s cold glass- house collection. Key words: Zagreb Botanical Garden, Faculty of Science, historic plant collections, succulent col- lection Sandev, D., Mihelj, D. & Kovačić, S.: Obilje bilja – zbirke Botaničkoga vrta Prirodoslovno- matematičkog fakulteta Sveučilišta u Zagrebu (2): Stakleničke mesnatice. Nat. Croat. Vol. 27, No. 2, 407-420*, 2018, Zagreb. U ovom članku sastavljeni su popisi stakleničkih mesnatica uzgajanih u Botaničkom vrtu zagrebačkog Prirodoslovno-matematičkog fakulteta između 1895. i 2017. Uređena je sinonimka i no- menklatura te istraženo podrijetlo biljnog materijala. Rezultati pokazuju kako je tijekom 122 godine kroz zbirku mesnatica hladnog staklenika prošlo najmanje 1423 svojti iz 254 rodova i 17 porodica.
    [Show full text]
  • Descripción De Aeonium Urbicum Ssp. Boreale (Crassulaceae), Una Nueva Subespecie De Uno De Los Endemismos Más Representativos De Tenerife, Islas Canarias
    VIERAEA Vol. 46 pp. 11-28 Santa Cruz de Tenerife, octubre 2019 ISSN 0210-945X Descripción de Aeonium urbicum ssp. boreale (Crassulaceae), una nueva subespecie de uno de los endemismos más representativos de Tenerife, islas Canarias OCTAVIO ARANGO TORO C/. Loreto 24-26, Esc. B. 4º 2ª, 08029 Barcelona, España [email protected] ARANGO ,O. (2019). Description of Aeonium urbicum ssp. boreale (Crassulaceae), a new subspecies of one of the most representative endemisms of Tenerife, Canary Islands. Vieraea, 46: 11-28.https://doi.org/10.31939/vieraea.2019.46.tomo01.02 RESUMEN: Se describe e ilustra Aeonium Se especifica su área de distribución, urbicum ssp. boreale, un nuevo endemis- las características de su hábitat y su mo del extremo más septentrional de la estado de conservación. Se incluye una isla de Tenerife. Los caracteres morfo- tabla con los principales caracteres lógicos de este nuevo taxón apoyan su diagnósticos que diferencian a las tres descripción en el rango de subespecie. subespecies de A. urbicum y una clave Se realiza el diagnóstico diferencial con que facilita su identificación. Se depo- las otras dos subespecies conocidas de sita material tipo en los herbarios TFC A. urbicum, la ssp. urbicum de la vertien- de la Universidad de La Laguna y TFMC/ te Norte de Tenerife y la ssp. meridionale PV del Museo de Ciencias Naturales de de la vertiente Sur y Suroeste de la isla. Tenerife. PALABRAS CLAVE: Crassulaceae / Aeonium urbicum / subespecies / taxonomía / corología / Islas Canarias. ABSTRACT: Aeonium urbicum ssp. taxon allow its description in the rank of boreale, a new endemism of the most subspecies.
    [Show full text]
  • A Common Threat to IUCN Red-Listed Vascular Plants in Europe
    Tourism and recreation: a common threat to IUCN red-listed vascular plants in Europe Author Ballantyne, Mark, Pickering, Catherine Marina Published 2013 Journal Title Biodiversity and Conservation DOI https://doi.org/10.1007/s10531-013-0569-2 Copyright Statement © 2013 Springer. This is an electronic version of an article published in Biodiversity and Conservation, December 2013, Volume 22, Issue 13-14, pp 3027-3044. Biodiversity and Conservation is available online at: http://link.springer.com/ with the open URL of your article. Downloaded from http://hdl.handle.net/10072/55792 Griffith Research Online https://research-repository.griffith.edu.au Manuscript 1 Tourism and recreation: a common threat to IUCN red-listed vascular 1 2 3 4 2 plants in Europe 5 6 7 8 3 *Mark Ballantyne and Catherine Marina Pickering 9 10 11 12 4 Environmental Futures Centre, School of Environment, Griffith University, Gold Coast, 13 14 5 Queensland 4222, Australia 15 16 17 18 6 *Corresponding author email: [email protected], telephone: +61(0)405783604 19 20 21 7 22 23 8 24 25 9 26 27 28 10 29 30 11 31 32 12 33 34 13 35 36 37 14 38 39 15 40 41 16 42 43 17 44 45 46 18 47 48 19 49 50 20 51 52 21 53 54 55 22 56 57 23 58 59 24 60 61 62 63 64 65 25 Abstract 1 2 3 4 26 Tourism and recreation are large industries employing millions of people and contribute over 5 6 27 US$2.01 trillion to the global economy.
    [Show full text]
  • South Laguna Biological Resources Inventory
    SOUTH LAGUMA BIOLOGICAL RESOURCES INVENTORY Prepared for the City of Laguna Beach By Karl in G. Marsh, Biological Consul tant January 20, 1992 Karlin C. Marsh Biological Consultant 30262 Acorn Lane, P.O. Box 404, Silverado, California 92676 714/649-202 7 SOUTH LAGUNA BIOLOGICAL RESOURCES INVENTORY Prepared for the City of Laguna Beach By Karlin 6. Marsh, Biological Consultant Uith Contributions by Fred Roberts, Jr. r Dave Bramlet r Gordon Marsh Rick Reifner January 20, 1992 Dedicated With Affection and Admiration to FREDERICK M. LAN6 Longtime champion of South Laguna's special natural resources TABLE OF CONTENTS Page INTRODUCTION 1 BOUNDARIES AND PHYSICAL SETTING 1 SURROUNDING AND ONSITE LAND USES; OWNERSHIP 3 METHODS 5 EXISTING LITERATURE 6 BIOTIC COMMUNITIES 8 MARITIME SUCCULENT SCRUB (2.2) COASTAL SAGE SCRUB (2.3) SUMAC-TOYON SOUTHERN MIXED CHAPARRAL (3.2) SOUTHERN MARITIME CHAPARRAL (3.6) ANNUAL GRASSLAND (4.1) SOUTHERN COASTAL NEEDLEGRASS GRASSLAND (4.3) RUDERAL (4.6) I SOUTHERN HARDPAN VERNAL POOL (5.1) FRESHWATER SEEP (5.3) I ALKALI MEADOW (5.2) SOUTHERN COASTAL SALT MARSH (6.1) I COASTAL BRACKISH MARSH (6.2) I COASTAL FRESHWATER MARSH (6.4) RIPARIAN HERB (7.1) I SOUTHERN WILLOW SCRUB (7.2) MULEFAT SCRUB (7.3) SOUTHERN ARROYO WILLOW FOREST (7.6) COAST LIVE OAK WOODLAND (8.1) I XERIC CLIFF FACES (10.1) I XERIC BARRENS (10.1) Page MARINE TERRACE PSAMMOPHYTE HABITAT (10.1) MESIC CLIFF FACES (10.2) ROCK OUTCROPS (10.3) PERENNIAL STREAM (13.1) INTERMITTENT STREAM (13.2) EPHEMERAL STREAM (13.3) VINEYARD AND ORCHARD (14.3) ORNAMENTAL PLANTINGS (15.5) FUEL MODIFICATION ZONES (no G.I.S.
    [Show full text]
  • Index Seminum 2018-2019
    UNIVERSITÀ DEGLI STUDI DI NAPOLI FEDERICO II ORTO BOTANICO INDEX SEMINUM 2018-2019 In copertina / Cover “La Terrazza Carolina del Real Orto Botanico” Dedicata alla Regina Maria Carolina Bonaparte da Gioacchino Murat, Re di Napoli dal 1808 al 1815 (Photo S. Gaudino, 2018) 2 UNIVERSITÀ DEGLI STUDI DI NAPOLI FEDERICO II ORTO BOTANICO INDEX SEMINUM 2018 - 2019 SPORAE ET SEMINA QUAE HORTUS BOTANICUS NEAPOLITANUS PRO MUTUA COMMUTATIONE OFFERT 3 UNIVERSITÀ DEGLI STUDI DI NAPOLI FEDERICO II ORTO BOTANICO ebgconsortiumindexseminum2018-2019 IPEN member ➢ CarpoSpermaTeca / Index-Seminum E- mail: [email protected] - Tel. +39/81/2533922 Via Foria, 223 - 80139 NAPOLI - ITALY http://www.ortobotanico.unina.it/OBN4/6_index/index.htm 4 Sommario / Contents Prefazione / Foreword 7 Dati geografici e climatici / Geographical and climatic data 9 Note / Notices 11 Mappa dell’Orto Botanico di Napoli / Botanical Garden map 13 Legenda dei codici e delle abbreviazioni / Key to signs and abbreviations 14 Index Seminum / Seed list: Felci / Ferns 15 Gimnosperme / Gymnosperms 18 Angiosperme / Angiosperms 21 Desiderata e condizioni di spedizione / Agreement and desiderata 55 Bibliografia e Ringraziamenti / Bibliography and Acknowledgements 57 5 INDEX SEMINUM UNIVERSITÀ DEGLI STUDI DI NAPOLI FEDERICO II ORTO BOTANICO Prof. PAOLO CAPUTO Horti Praefectus Dr. MANUELA DE MATTEIS TORTORA Seminum curator STEFANO GAUDINO Seminum collector 6 Prefazione / Foreword L'ORTO BOTANICO dell'Università ha lo scopo di introdurre, curare e conservare specie vegetali da diffondere e proteggere,
    [Show full text]