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Journal of Natural History Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t713192031 New species and records of anthuridean isopod crustaceans from the Indian Ocean Brian Kensley a; Marilyn Schotte a a Department of Invertebrate Zoology, National Museum of Natural History, Smithsonian Institution, Washington, DC 20560, USA.

Online Publication Date: 01 November 2000 To cite this Article: Kensley, Brian and Schotte, Marilyn (2000) 'New species and records of anthuridean isopod crustaceans from the Indian Ocean', Journal of Natural History, 34:11, 2057 — 2121 To link to this article: DOI: 10.1080/002229300750022358 URL: http://dx.doi.org/10.1080/002229300750022358

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Clearl d ; Pendanthur , : s som a in Abou fro Paranthur p w Famil or m Famil s s arabic g a a Lt m throug reef waters d a of y e comparativ , calci th w larg d y , c s th fro Atoll fe US t s know Accalathur , y e to Natura thi n , y 13 of e fro , w Aldabr a record m andamanensi a s seagras e Antheluridae , India A limite a , biodiversit h a s 3 area , Anthuridae Leptanthura th Hyssuridae numbe Mesanthura a , m n urodentat numbe th specie cosmoled is Zanziba seminigr variet e fro e l n th a d s e isopo poorl Indian s Ocea bein m s islan a table Atoll fro e an r s has y r th of of m of a a o is s d d y y g d n r e s - , , . , , , , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 in compositio anthuridea th Aldabr Ocean of 2058 articles asc sediments m Is. on ree m 6 cora th ca bearin statocys shorte wit thre shortene Anthomuda Systematics Copenhagen Unite NMNH stations specie comparativ Apri e e 4. n , 4.4 e th f s h th Abbreviations Diagnosis Othe Typ Remarks Diagnosis Aldabra mos e 6 bul non-ovigerou slope l be on e e on mm ± d s shor rubbl r : l g region 5. . a generi e k collection 1987 is subtermina ; separate t t e Ð tha aesthetascs r Mandibula State d 8 Atoll materia , , obviou , openin K provide of materia mm to Nationa t Picar n articles Schultz on n Aldabra e n Ð e , . . articles ; fauna , . . thi of P c tw e s 20 table an posterior Bria sta , , is Famil ara Th damaged Zanzibar synonymie d Nationa s 19 o Eye Ovigerou d d th m, base s g l Ð . s . , d Is. l s , report n e of by e types robus coars s l especiall l 1979 by l examine m . station . r , in juveniles 11 wit lis s FÐ y Museu , ® article H relevan Copulator to Kensle d intertidal palp whic Mandibula Aldabra 5. ve subtl presen ANTHELURIDA t olotype th smal Apri , h on : 0 of Kristia e distinguis : t 907 l . e specie latte , , mm USN thre Anthomud s , h th Genus In sand seta st accumulation key th . Museum s , e SUBORDE m articl female l y ; l d t on y a ar e , t e an , B. . feature pore Poor fro 1983 r gener e of additio , Gul s st K-AL-26 e e India , fro e M in ® s : , 18 n wit d y termina Kensle as 25 a el USN m summarize on e USN on Natura of r Aldabr Anthomuda style 253160 e m . d a bot K-AL-39 : m, h f . Fauchal 2 h th m. a pal a Antennula an Mal termina of ; ( Anthomuda n a wit generi s posterio stations th Figure furthe tw e 14 n 7 h is M d ZMU M Ocean p quadrilineat suc t Arabi tropica e y Apri R h o Le to e of , l sexes provided l a s of separat E 253162 an , 253159 India coars History singl h aesthetascs antennula ANTHURIDE robus w on of c d lagoo pleopo describin Poor r , d d l s thre l as C Ton , a ; e diagnosi cora article r ai isopo 1983 . ® 1, in M. Ð som Schultz l e r ovigerou an n MS el d e margin th Antennula area ¯ t e , lon 2 e , e Zoologica , tabl n d no sedimen to d agellu e . Ocea 1988a , l Schott ) setae ovigerou on d . an e articles a stations Smithsonia Ð d rubbl Revisionar at Phuke USN proportion w . stations g s separation informatio 2 r sp. e g Antenna e d materia of . Marily dista s , hea 1. know apicall ¯ ; n Le an . Antenna : 197 m s agellu provided th nov. e 500 carpu e l M Telso m species A 4. t d . w t d of e r fro l , 9 l 4. s an 1 Island Pereopod ; 253161 . graniti seta n addin Ton of ¯ n cora l fou m 0 mm y Museum m agellu l JR d m fro y n fro s n mm s . 4. ¯ hooked Schott n Grand , fore-ree cora l work agellu r . wit , Institution Ð ¯ 8 of , of , on m fou l m , articles ¯ 1988 g attened Distributio mm , c fou as , Jac Thailand agellu rubbl h tw m som aroun on Seychelle abou th 24 th l r s s e anterio r wel rubbl o , , fringe m e k e e a e . of 4± ar st specie non-ovigerou f non-ovigerou e ® de pre Universit India Passe e of , m Rudlo t slope a l el e , 7, d appendages tw fou ® an te wit ; d as K-AL-115 referre fou - e nitio th d of , USNM o propodu n r n s d on r mak n stations s , e l seta h 3.8 key aesthet , islands r article margi tw e to 2± India of Picar to coars Ocea setos n singl oute y mm, o ® s d e e 3 an ® th th el up on m, Ð or ve to to of d d n d n n e e e e e s s r s s - ; , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Anthuridea

Table 1. Features separating the ® ve species of Anthomuda known from the Indian Ocean. n

Pleopod 1 Uropodal endopod Uropodal exopod Telson isopod endopod setae length:basal width length:width length:greatest width s

A. cracens (Kensley, 1980a) Mozambique; Somalia 6 2.4 3.1 2.1 fro

A. hapla (Kensley, 1980a) Mauritius 3 2.5 2.2 2.7 m A. hovea Poore and Lew Ton, 1988a Queensland, Australia 4 1.6 1.8 2.4 th e

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Colour Remarks Diagnosis Etymology 20 e propodu y 3, colou d , r h h e e . l resemblanc , a y inserte r g see statocys cephalon l carpu abou abou A , r fa wa folds e plumos 3; of uropoda s tw ; truncate longitudina d l s s l . thre r margi setae n 1± cephalo endopod s r . 7, lamin in barel o hapla ¯ bot mor ( features l als l agellu pattern t ; t 5 fro . Kensley ¯ sensor s d s endite e . carpu Dorsu g pal pleonite 2. ated 1. h . wit fused . o t barel Nunomura obliquel e n fringe e m In Species 8 th 8 y an Uropoda . Eye a th recorde l , p e margina Th ( elongat h shorte time n time e exopod in m , wit th Kensley d e additio dentat to . y s of , . , stou pal y species , m Anthuridea ¯ l wit s e antennul a A a e , setae wit d indicate quit ated of line h expanded an s speci mor . 1980a s thre broade s Genus India of m usuall y h poorei seta of 4 basa tw r t h basa Famil d y at tw , e s a l e dista , indistinc l . ofte n ; cephalon an tha fro fre e e 1977 o of , , exopo Amakusanthura mor almos ® propodu posterio Pereopod di distolatera o setae e of elongat to n ) articles as c 1980a) l d termina l e y m d th e Amakusanthura Œ n r n y ha width anterodistally nam width to Ocea l Mu : 5 erent th anterio termina an e si laterally in e n , seta ANTHURIDA posterior steppe Re 79 present s ; x no margina e t margi d thre È d isopod fou ller a ;  th t e reachin endopo union character teeth , broadl , e n Poor ; r t ; th , s , lin refer les e l pereonite propodu from uropoda s endit unlik posterio r demarke , e rectangular bearin pereopods ca e r setae l d ke 1990 e l n s 4± Paci articles . angl ; antenna e n margi maxillipeda ; s or elongat of y of s . pleonit by l an 7, y mola g Antennula be d fro e e to plumos Pleonite below A ; , ® ten weakl d . pal pigmen g e posterio ovate smal s carp Mu A . two-third termina c fro m th r furthe of l Pleopo s Le n numerou Nunomura provide d s quadrilineata . . species d exopo E m margi r wit e È . th m an shorte . w e quadrilineata e lle articl dorsally Mandibula l , , i to fou small Th Leach, , y bearin A e 6 telso e wit Ton h is or Re d r roughl ape s t . toothed r India l seta demarke be e ( r tw r l d pleo  r sometime d n 1± 199 pal betwee cf. h union e absent . distinguishe d , presen margin line pal r s x , n an o broadl s 3. 1 tw 1988b morphologicall 4 ¯ in e tha g p conical , of narrowl 0 hovea elongat stou 1814 agellu wit n n , y d p exopo Pereopo s on o fused fringe onl ); articl tabl 197 endopod Ocea of , wit n triangula . sensor articl r exopod n h . an thi t : ha d Pereopod th , . t y posterio Pereopo y 108 pigmen specie pal 7 a eye e h s sensor fro d m ha s e s , indicate e Poor . rounded d d 1, e proportionall n base y specie a thi e endopo margin an mor . p Maxillipeda s d s m d y scales dista operculiform mor rounded th of havin , n d an an seta 1, by of s , e telson r d e y tha e wit t a doe spider r d d carpu or s an y s fou thre on e seta l on apicall mor d thre article th margin e . ha . s 1 2 fou h setae d n g elongat ver Pereopod d on s e laterally pentagonal th r marke subchelate A an . of subtle ® s e e no tw ; e e following Le specie r y s y ve . e a posterio on endopo d elongat quadrili- pleopo . triangu o l t s roughl articles articles y dorsu y line similar w distinc , Telso 3 mesia bea , endit in palm muc e dista latte acut , d 2061 Ton wit wit s , an s no th s r by or of of m n d h h d d h 2 a y e e e e r r - t - l , . . l . . , , : Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 ± 2 1 2062 juveniles reefs ovigerous san Grand non-ovigerous corallin slope 26 K-AL-85 intertida one reef thre channe ovigerous st 1986. K-AL-110 ± 8 ± 7 6 ± 5 ± 4 3 ± ± ± a Uropoda Uropoda Uropoda Uropoda Uropoda Pereopo Uropoda Uropoda Uropoda seta Pereopod erec Pereopod T T Uropoda Pereopo Uropoda . . . . . d elso elso Marc Othe Typ K-AL-110 e slope , pre- e t on non-ovigerou . . . . . P o Grand e n n seta l Œ e ara e Passe r Pass l , convex moderatel h , alga . . . . . oute d d st Picar materia cora o hibbertia materia l l l l l l l l , l l l e m m coars types 1985 1 Œ 1, a 1 1, exopo coars endopo endopo endopo exopo exopo exopo exopo 5. endopo exopo , ...... e e 4.8 m K-AL-117 m Picar 0 l , nin propoda propoda r , , l Hoareau , turf Aldabra d mm , Passe carpu m rubbl ree sam carpu . ± . Ke . . . . . wit e , : e e d d d d d d Is. 6. l On l y USN . d d d tw sedimen d d Poor , san examine f , agonis distall juveniles distall no les slightl h no 0 y s . . . . . domed , e lagoo lengt lengt abou lengt Is. s , slope mm ® e o m slopin s to e H t s Aldabra l t l dat distall d ve Aldabra distall sinuat pal pal , tha notched . . . . . e non-ovigerou fro 4.5 olotype , M juveniles , , an th Aldabra 5± y y y , Amakusanthur an coars Malaba h h h t a Poor n non-ovigerou m m , , n e m ± 1. sinuat mor notche st . . . . . 253186 d g tw abou les t d d 8 taperin as 4. y nea Picar twic , India 3 a no y steppe . rhomboi e rubbl m, B. an Pass Le o s acute 8 st time . . . . e . , rounded distall e e K-AL-101 , t holotype tha , or mm r w a USN d t 15 an e usuall Kensle rubble tha : steppe e 21 , 2± d , r twic Pass d longe . . . . . e n K-AL-115 T mor g n , d rubble s 22 st e d ; Is. USN m, on n , Marc 3 fou Hoareau Telso Is. longe to ( twic Ocea y a fro Le or d nin Figure M e m, m, twic , ...... s y , e coppingeri e d ; , K-AL-107 12 subacut w Aldabra are basa r , 1988 r y toothe m wit m du time T e a s oute . Aldabra M or 253190 e tha 13 , . n . . . non-ovigerou . . . Ton r 6 , elso e an h n , basa a USN ree cosmoled Marc juveniles m h tha K-AL-87 posterodorsall longe Ans toothe Bois l 1986 cora Marc n specie posteriorly b Apri s d s latera . . . widt . . . . 253185 r n , , , 5.0 f basa d basa Grea n 3, e l M. 1988 ree Malaba coars slope posterodorsall e M widt , basa ape , , h l ...... r . . ( ± l 4 h 1 , Aldabra on Barnard h d f Tw , Var 5. tha rubbl l l s 1987 l Schott ) 1986 t m, b slope 3± 253187 x widt widt margi cora , 7 o h of ...... 1987 Barrie cosmoledo e , l , e , Queensland n africana st o mm widt , Cosmoled 5 sp. non-ovigerou , 18 coars moragallae cora Amakusanthura a non-ovigerou basa r . juveniles m, s ...... ape . . . h Picar h dubia e l USN , y Is. K-AL-124 n . e On m , Marc , Picar nov. h an r rubbl , wit , 192 On sinuat x ...... 22 l l , stocki eigh 4.8 Reef 0. e on d e widt narrowl y Aldabra,1 d rubbl pori h Barnard Barnard 2 M e 5 cora sp ...... juvenile ± wit Marc alga e d m, h abou ) Is. e t ; 5. ; non-ovigerou , o . e h 253189 North-wester Is. 1986 north-wester north-wester st Mu Mu Wa h nov betwee juveniles 5 ...... , . . . Is. 21 e l l l a , mm on 5. , t Aldabra È s y È È h Aldabra gele rubble in tur , ller s ller , K-AL-108 , cora 12 . 5 Marc ...... m rounde e . 3± 191 191 , Cosmoled 1986 , mm 1 USN , ± m or , , lagoona f , , st 3 15 5 n fou ...... 199 199 Apri on 1981 4 l fro 4 a m, juveniles tw , 7.5 , h rubbl patc , elongat Sout Sout . e K-AL-106 r 1 1 d , ...... M st eigh m o n n n 1986 oute , On m 26 b l s juveniles mm, Sr Sr 12 10 a erec Australi Australi Australi 1987 5.0 253188 h edg h , h Re o ...... m l i i e e Marc ± ± F8-87 t coars , Lank Lank r Afric Afric patc Islan . 18 reefs 12 e d fro t non non mm, four , On e ...... ree . Se st sta m, m, of m h h a d 5 a 6 4 a 3 2 e a a a 8 7 a a a e - - f , . . , , , , . . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 ovigerous intertida ree 7 F ig. Apri f 3 slope . pleopod 1; pleo l 1987 (G Amakusanthur l n , corallin m ) latera , Picar maxilliped . st Tw 2. a F-7-85 l o d e view non-ovigerou Is. Anthuridea alga a ; , ; ( cosmoled , (C 12 H l coars turf ) ) m, pleo uropoda , e 7 n o n Pass intertida s Apri dorsa isopod sp m l , . e tw protopo nov l Hoareau l 1987 o view s . l juveniles : rubbl fro ( A) . ; d m ( Tw an D) , e th d Aldabra l o fro mandible anterio e , endopod st non-ovigerou India m a K-AL-118 ree r n , f ; segment ; 12 Ocea ( crest ( E I ) ) Apri uropoda pleotelson , n s , Picar s cora l m in , 1987 latera st d l l a rubble exopod Is. ; . ( K-AL-119 F , Tw l Aldabra ) view pleopo o , ; oute ( 2063 non ; J ( ) B) d r - , , l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2064 line wit seta posterodistall of process Pereopo reachin articl surface posterio posterio Aldabra Tw st rubble Is. 18 F ig. a , h Marc dactylus o F-7-87 Diagnosis d Aldabra e 4 shor e non-ovigerou . wit at , . 2 , g Picar Amakusanthur r r d , h wit Lamin h posterodista t longe hal 0. to margi , 1985 rigi 1, fringe , cora 3 h , f m, sutur . d wit 27 y carpu fe d bearin r . a rounded Is. w n l Non-ovigerou set h 13 Marc On tha d rubbl dentat notched , scattere e s shor a setae s Apri Aldabra e n m posterodistally betwee g a l pereopo , triangular non-ovigerou h 1, abou e fou angle t cosmoled . 1987 , a l fro squa articl Pleopo lackin 1987 d , r of m n bearin t juveniles seta , . . B. d 12 mandibl 3 t patc On article Pereopo s . e 7; o g Kensle accessor ± , Apri e female d 3 15 ( e fre sp posterodistall on D) g h 1, short pre s ; . cluste scattere e reefs s l , propodu m pereopo mesia endopo nov e st anterio 1987 - , y d 2 : st a Pleotelso y , havin an , l . 7, , an a : F-11-87 wit r tw Grand set . l d ( F-1-87 d of carpu d A) face Thre o d d h M. a r s g elongat 3 2; non-ovigerou thre margin slightl roughl tw y at m fou ; of ( Schott e n e , s E acute o ungui , pereopo base e Passe cora ovate ) non-ovigerou r intertida wit palp dista pair pereopo y e margina y ; h . simpl ; shorte l e rectangular propodu s , , s rubble , Pereopod l propodu shor abou Aldabra d wides wit of setae 1; s l d e h setae r t reefcres m l 3. seta t ( , , tha fre B) serrations . t singl as s st ree s at Maxillipeda s , wit a e e s , n lon m , 2 f abou pal at dorsa F-4-87 , anterio posterio 2 l e exopod m, crest h t pereopo thre g righ an rubble termina m stron 6 as t d ; l , Apri e wit midlength t , Picar remainde surfac pal r 3, juveniles intertida , angle g r d , h margin l bearin margi p serrat l carpu Picar l endit 1; 1987 blun d seta wit e s (C Is. to of n h d g e e s r t ) , , . , . , l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 pal lik article simpl laterodista ® posterio subequa pigmented larl Aldabra 11 3. cora o non-ovigerous patc ± 2 ± 1 Pereopo pereopod Maxillipeda mola pleonit Apanthuroides holotyp telson Queenslan beyon havin Œ ve 5 Bod Bod Margin Margin e . m y Apri mm Il Etymology Remarks Male Diagnosis Othe Typ Diagnosis a plumos h l e r an e g y y , . bearin slightl d rubble s ree Picard e spike-lik , setae bu e abou abou e bearin d seve l r ape d l , r 6 : st s s materia . s f locality in 1987 intertidal materia indistinctl t margi Cephalo 7 of of d . no l a in 4± th e x Pereonit l n y , t t lengt sometime . g margin , F-8-87 . resemble uropoda uropoda , . . t margina Menzie eigh 11 lengt e of pal o 7 Aldabr g . . Aldabra Pass dista triangula di m Œ Amakusanthur latte e n whorl USN Eye Female . roughl time ramus Œ l se p Ke , Th h 2. . t l on erentiate broadl Genus Cosmoled n h examine t to e , article 5 to l , . , Anthuridea s r y y tooth e s e H 23 s wit abou mm l l du M cora plumos a on nin present distall ha an s l to speci 7 pitted longe s s ram ram telson olotype , . . y seta lagoon of : Marc absent les th r h d e s 11 e , 253113 th y Bois Bod pentagonal Apanthuroides s ; l i i toot Glynn t a e eye time 4. side Apanthuroide aesthetascs s . d e ® propodu d r m, an an rounded e 1. rubbl 1± y distinc . tha 0 presen . , c tha o India e . distally fro h s 5 d d Cephalo y , mm latte a . 2 s name h , Dorsa acute 6 . , Island : enlarged margina USN telso telso time eigh 1985 n n Aldabra n longe , Pleopo reduce , m Apri 2 USN fused an 1968 angophor e hal greates . isopod , pre m, n r t t pleotelson d sam in n n t , ; ( . s s wit . latera Ocea . f M l specie r Figure , - l use to , abou Antennul 6 : wit entir ® basa M endopo Uropoda , . integumen foveolata lengt n tha Pass 1983 wit nel d 29 on d Pereopo h e 3 l 253110 Apri l , t , s wit nin 2. d s 253108 h Menzie ; dat 1, or wel fain n widt e y n setae free 10 aldabra e h l Kensle fro 2 l a as t ban s e h greates . dentat of h e s ` width specie bot mm . shoulder anterio eigh a l absen m, USN l in Hoareau of m . t 5, a d time h lo , Poor 1987 pigmented th as d Bod , rounde , ( e l h 4± nou . . w d havin pereonit , , distall th Kensley,1978 6 s e 13 of copulator y t exopo t e wit non-ovigerou holotype ram ) e cora . t 1, l s M 5 s rounde an e pitted Aldabr an 3. t simpl e . . . . y widt seta sp. n of r longe India Marc fused 2 on carpu h P d d of 253111 ’ in i g an margi mm , ara , l d y . . . Apanthuroides Schotte basa Glynn d nov. formin h unlik e Il rubble mandibl numerou e other . d middorsa apposition e rounded . nea ovate r e d n Pleo a h types , , . . . . 6, setae s Antennula . y st tha endit l Malabar Le Ocea rostrum grou n wit 1086 , b lackin e r a style articl , . . . . , ; ) n g w pal shorte , A n K-AL-24 , 196 1989 h s Sout . l pal : 3. short wit ree operculum . e p e wide m n Pleopo . . . . . USN posterodista s , m t cosmoledo 0 Ton l somewha e 8 p of 3. present g USN h f : , bearin erec longitudina mm h rod-like , on longe , . . . 2 26 r is pereopods of crest calculosus r shallo eye , ; Africa islands , mm Aldabra M aldabrae pleonite tha . dorsu ¯ take mesia , t . . d , M thre agellu 1988b s st seta cora 253109 g , r , n . lateral a 1, . st , n w . . Il t , 253112 Carpu an . numerou K-AL-89 e posterior . 550 m e a l elongate e endopo fro reachin l notc l m d surface articles , s , . . F-7-87 sp. sp. margi Picard rubble l . on irregu ± 0. fused m ridge Pleo wide , , 85 of fro 2065 3 wel s h nov. nov. . . tw , th th 0 m, 15 of m in m d n o n g 2 e e s r - , l , , , . , , . , ; . l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2066 contributin 7 shor Pereopo dista setae fou stron teeth termina of thre tha F ig. lacking thre n r e 5 t l . ; articles g . tw uropoda lef aesthetascs robus posterodista Pereopo mola e dista l t o Apanthuroide d articl . setos mandible 6, dista Pleopo g t r l , carpu to seta l curved spin basa e e d exopod l opercula wit articles 1, peduncl , , d e ; fe article l l s carpu h ( 1, an robus articl w wit s , E thre ; aldabra spike-lik ) d protopo seta ( h . J righ s ® r e ) s e fre Mandibula e t ve uropoda 3, function e triangular article ver setae seta t e nea e 4 B. mandible mor anterio sp y d e an ; r Kensle . ; short on wit propodu s nov posterio d e inciso l ; together endopod slende 5 h endopo on r . ; r : eac , ; thre propodu margi ( pal y e termina ( r A) F an h wit side r s r p ) e m wit margin d . ; wit subtermina maxilla d retinacula of h dorsa n , ¯ M. abou h thre agellu shorte h muc s l thre singl articl barel serrat Schott l . e ; view t h Pereopo e (G cusps m hal e r shorte articles e y e tha aesthetasc ) e ; l of e f semicircular in ( on maxilliped posterodista spines B) widt ; n ¯ ® lamin ated mesia d posterio r ve antennule , h 2 on middl . articles , of simila Maxillipeda a . wit l other ; Antenna exopod dentat margin ( r h , H e ; l r margin wit (C stron , ) articl seta . to articl pleopo Maxill ) h a pereopo antenna ; ; . fou wit l latte g bot e Pereopo ¯ l ; e postero agellu longest bearin h pal d r h a 2 r dista seve 1; ram ; wit wit wit p d ( ( D) of m 1. I h d h n h g - ) i l , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 tw Antenna wide. ovate plumos numerou F ig. o Male 6 slende . , antennule 2; abou e Apanthuroide ( l : s D) seta ¯ Antennula elongat r agellu t m e seta twic ; pereopo on ( I m e ) e e oute Anthuridea as dista at s of l r pleopod aldabra d lon si ¯ r or agellu x 6; l margin g an setos nea ( e as E d 2. sp ) m r latera n wide . e lef , nov posterio isopod articles of singl t mandible ; . l eigh : endopo ( setae e A) s plumos . t r fro Pereopo article . l margin pereopo ; Uropoda m d ( F roughl th e ) s e set righ d , eac d India 1, a plu 1; l t y h on ( protopo propodu mandible B) s ovate bearin n posterodista inne m Ocea pereopo , r g 1. d s n ; dista 5 ro wit wit (G time w d ) h h 1; l antenna of seve l angle tw (C s robus longe o aesthetascs ) n m stou ; pereopod elongat ; exopo t r ( t H 2067 tha seta an ) d n d e . . l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Apanthuroide th ar incomplet proxima Pereopo 2068 carpi posterodista posterodista proportions broadl Maxillipe other thre ¯ tw slightl stron basall Pleonite lobes pigmente Integumen 3 1 manc rubble 21 ovigerous cora intertidal 253102 K-AL-13 P Pass transparen mandibl of agellu ara Ma Apri e e o pleopo Apri Etymology Remarks Diagnosis Othe Typ e apparentl l e manc dista type ; . g . y a rubble y y setae latte Hoarea , y l Maxill Pereonit , m 1984 middorsa e 2.8 l Mah longe broad 1983 s e d r l rounded thre d , , 1986 l d materia d : of a e hal 1± in ± Cin r t t article 12 7 eyes m ; materia USN 1 transvers . . 3. wit , wit l l ¯ . conditio pitted e si 5 e . mola th . f s a USN lacking r Pass 2. mor ang y 0 u Pereopo , . seta simpl x Apri . q Beach non-ovigerou aldabra of fused an elongate e ; e h Al h mm USN 9 taperin Th matur Non-ovigerou channel setos M lo Cases l , mm 7 s female anterio l e d e l . wel . r e mesia wit w ridg , M e l togethe l on abou th , 253100 e setos wide Pereopod Femme proces mor speci , triangula , M 1983 . n st , H e e l examine e h 253107 seta e Mah wit , d e e anterio a Pleopo suture g articles st olotype availabl develope thre int ± Sout 253105 ; l male , di r t e r runnin e a K-SEY-15 slender posteriorly 1, h ® whil . Il margin , Œ margi tha two-third Apanthuroide in o r so s , c e K-AL-107 USN e , thre ers e dorsa non-ovigerou bearin s propodu Aldabra h nam stylifor , Is. s B. th Malabar adulthood , dista n s d in e d r r m . s nin Is. in . from , e e , 4± tw rostru e g : Mandibl margi th Kensle l n 3. d , M 3. 2, thre Seychelles posterio female , female specimen e , USN submargina dista almos l th USN e e shorte 3 o 7 l 0 bearin g Aldabra endit derive pit margina mm , ( endopo s dista 253103 e ischiu mm m dista , wit e A Figure ® , s Ans n lengt wit m , samples ve , s intertidal . non-ovigerou l M y M on h , : Aldabra no calculosus cora s no ; t . , spin e no faintl r e l an h l st g aesthetascs s in r cora e calculos entir Bod In m tha s posterodista plumos typica t 253099 peduncl t s havin h a fro t on , , tha posterio , fou d d a 253101 s m havin see l Á of e reachin l 8 expanded cora of non-ovigerou K-AL-40 of 7, n setae M. additio e l y la l 3. y , m blunt wit m, rubble e n r n thi rubble posterio setae it 3 , pereopo side 8 pereonit indicatin , g length abou l Mouche dista th in 14 in h Schott l mm ) a 2 , g e cora s of is . describe e rubble tw , e sp. , Apri ovigerou greate setae specie r Pereopod , g copulator anterio A Apri articles s possibl . . genus typ wit , n , o non-ovigerou margi anteriorl t l represente . , Antenn m Pereopo l , , l . sam Grand calculosus nov. ree plumos r to te rubble e ree dista h e d l serrat 4. Antennul pal e . margin l g , , n s 1986 r locality fou 0 f 1983 1 6, ; s d Mah 6 e f thi r lac n origina mola ; e m crest mm dista time m, s of m ¯ dat belo l ¯ wit half s r tha a e evenl y s at e k agellu y , . m d spinules e 2. 1± concave smalle . th wit , e USN Passe , 3 a 0. d as set six-legge pereopo style . h , USN r s l 9 setae t w 2 . st . 4. Il , Pleopo 3 e Is. Picar 3 Apri e as pal developmen mm by pereopod thi l h Hea Aldabr fa a longe e a y 7 s in wit of m, mal wit , m pleonites mm r K-AL-112 Picard holotype t articl rounded M on p s , r M ; Seychelles m havin tin simila as , h l ; 11 articulatin of d d spine h , specie articl Aldabra e endopo st 1983 253106 r , d d 253104 dista y d propod stou anterolatera Is. 3.0 bearin wit ha basa a ® a e Apri st 1 tha 7. ve ` , , g condition thorn K-AL-49 s 2 Atoll s a r s protopo h e mm, Aldabra Aldabra . s As t th . , a l siz . present present n longest articles l bearin USN t F-8-87 weakl l retain serrat d setose T , , e g articl articl , broa of , , i e 1987 wide elso thre ther cora ram non . wit g ’ 3 6 an an on th sta on m, m, M in n d h d d d g y e e e e e e e s - i , ; l . , . , . ; . , l , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 fringe of Antenna basall tha os exopod wit F ig. e ( cephalon h n D) Remarks. Male seta 7 wide thre . y d antennule ; narrowed e seta Apanthuroide bot l : e on , ¯ Antennula dista retinaculae agellu , e h oute no in ram In : t l dista ( margi quit m r , E i genera margin wides ) formin Anthuridea of s mandible r e l calculos ; seve ¯ hal n meetin endopo agellu t setose l f , at g proportions n singl of operculum articles midlength a ; g m posterio ( , sp d n F middorsally rounded e of . ) abou isopod plumos nov maxilla . eigh Eye . t , : r . , two-third t an ( s Uropoda surface ; distall s A) e ; articles exopo fro enlarged (G d set . m in Pereopo m ) a dorsa pleotelson y th on . th d , somewha e s no l e eac inne protopo , widt integumenta l India occupyin t view d h reachin r 1, h bearin ; dista n ; ( propodu of H t ( Ocea d B) tapered , ) g wit an g l maxilliped g midlengt pleopo angle mos d l whor n h pitting slightl ® s t . ve bearin ; of d l endopo elongat of 1; ; latera h y , ( I A of (C aesthetascs longe ) g . uropod endopod ) ® calculosa l d ve antenna e surfac longe r plum stou 2069 tha . n e r t - . , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2070 gravell smalle th telsoni A foveolata three-articulat three-articulat closel F ig. . e 2; spathulicauda Etymology Caribbea 8 ( . y D) r c y resemble pereonit Apanthuroide an or pereopo ( d Kensley pebbly uropoda n e . e (an ¯ ¯ e s d agellu Th Wa agellu pits A 6; , d , referrin È . e s gele ( 1978 als l W calculos E . speci margins jiensis m ) m o , lef b of 1981 th of t ) g ® th mandible e a fro B. c to th ( sp e typ b Kensley . nam m Kensle e antennul Apanthuroide . th fro nov antenn e dee e e m specie rubbl ; . : p derive ( th y ( , F A) wate 197 an e ) a e e s righ Mediterranea (fou (si d habita of 9 s l r pereopo x M. ) s t o fro r th fro milla mandibl article Œ in Schott e m th t m th genus of e e d e th Fiji s Menzie eas 1; presen th e e e in n ( an . e t B) ) Latin Th bot th species coas d is m e maxilla t e s a h pereopo species) presen an , latter t posses squatte of calculosus d . ; Glynn Sout , (G t d however . s species 1; r Apanthuroides ) ® h maxilliped specie nel (C , Afric , 196 ) y meaning pereopo ) dentat , 8 s an a ha fro wit an d . s m h d a a d e Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 ® an male Eisothistos distally si articles or olatera fa serpuli USN of ± 7 6 ± 5 ± 4 ± 3 2 1 spik wea elongate margin anc ± ± ± ± ve x n T T Posterio Posterio Posterio Pereopo Pereopo T T T T Telso Telso Posterio Posterio d ...... te elso elso elso elso elso elso e Diagnosis articles Diagnosis. Typ Thailand thin k e narro lamin ; n M in operculum an ...... in d n n n n n n larg l n n . , . e ; margin d . Antenna post-spawnin 253130 or tube) wit wit wit wit wit havin wit lackin femal Pereopo materia pereonite . . . . Pleopo r r r d d r r w bataviae Haswell a vermiformis serrat . e margi margi margi margi margi 1 1 h h h h h h fuse Mandibl articles dentata spine , . . . . . propodus propodus 11 abou si thre si mor g , g e x x Fift s on middorsa , d middorsa Eye Non-ovigerou . . . . larg e anteriorl n n n n n d spine or , l spinose e d l e . , si s fre t margin e wit of of of Kensle of of 1, s peduncl Ke h spine 1884 tha x fewe te . . . . . 1 plu Anthuridea e s immatur . 2± H e ram e telso telso telso telso telso Pereonite n s h spine Maxillipe non-ovigerou Thai-Danis Haswell y present n bearin , , g olotype in 5 thin in s larg . . . . : lengt lengt s r pleotelson to si l females 677 , elongate y severa l i in spine male y n n n n n x . middorsa ro Eisothisto sligh ro fuse s Genus an . . . . e th e Telso , spine wit non-dentat wit dentat produce wit middorsa w ; h h plu w spine lamellate g d articl e e Wa , , mor les s of h . h . . . h . d 188 of s Poore India t s female s severa enlarge : l in Tailfa d n È s abou eigh abou s in . gele media n female ro USN sometime spine smalle e spine s , h in Eisothistos . e . . isopod . 4 slender tha middorsa Pleonite e l . w lackin uropod in juvenil ( tha Ne ro s d neoanomalus s t , Pereopod 5 middorsa , n Figure l Expedition t of t n 1979 . . . . teet middorsa l e s andamanensi 198 n w , ro beyon m . M s equa 16 20 w n Ocea n d twic elongat sam : r Maxill smalle indurate w twic 3.9 ridg Bod h Sout ...... s . . g 2 in 253129 , spinules teet teet : e s fro s s e Houtma endit l statocysts 2; l s an e ± n male an ro d . . . e ...... elongate . widt 1± to h h e l 5. 9, y h dat Kensle Haswell m r specie width d s l ro w a rostrum bearin d maledivensis 2 e slender Wales submedia 5 ro article 10 ...... prespawnin 1± , e ( th w thin-walled ; mm , h telso a , seta Shyamasundar . w free uropoda non-ovigerou obsolet January antarcticus ) anomala e 3 n or as s Antennul ...... crateris y s andamanensis India sp. Abrolho , no ; , e g ; n an holotype , (on of Australi absent s , wit givin ; ...... longitudina 188 pleonit on t sclerotized 3 integumen d n mandibl nov. Eisothistos subchelate e e h an Schotte n spine ...... ± l Wa 4 ( merus Kensley or wit g Marc dentat g Kensley exopo Ocea , d s . e V È spine a female bod ...... gele Islands Mouthpart e 4 anho corallina lacking wit s . h i [include s et together 6 e , n , tw h ...... m sp , y h 1989 d e ; È lackin , t al. s 197 wit l ischiu Œ , 6. . worm-lik telso o 197 ¯ , othe posterio 1966 wit ro , en barel 1980a , ...... nov 0 agellu togethe propod 1991 Wester ; 9 h manc : w d , mm sp 6 h pal 38 Maldiv 191 ...... n fre as g r of St ; . . m stron Andama y s . ) ) nov tha ¯ wit palp p , . 4 P typ e m Madagasca Sout reduce agellu abou ...... discernibl a an Pau wes n r ara of Antarctic e r i posterio h n . Australi of e margin in formin e d g slender appear Aldabr , ...... poster of l species t h on Island types apica mola t Islan basis seve coas sam n m Indi 2071 d nin e tail ...... Se to of in n g d . a 7 6 a a a a 4 3 5 2 e e e r r - - - t r s , l ; : . . . ] . . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2072 K-AL-99 margin ram posterodista abou an one- wit margi Pereopo shor surfac ungui F ig. d h Typ Etymology i 9 ® t basis t havin shor . pleotelson. ( ft s n e blun D) 3. ; h e of of slightl Eisothisto latte d 5 lengt , antenna t ; materia time 7 ischiu dactylu g t intertida seta carpu l wit si r posterodista seta bluntl x h y . s e h dista of shorte m longe ; s Th s on elongat . l ( s andamanensi . res an E Pleopo wit l e y relativel ) l posterio d H t spinose antennule corallin speci plumos r h r of merus olotype tha tha posterio e l dactylus d ® n seta seta n Eisothisto c 1, y B. wide r ; ; e e s anterior nam endopo short propodu ; ram e setae margin ; sp Kensle : ( alga on ungui F r . , . ( USN nov e ) i wit margi Pereopo Figure . pleopo . merus l derive fuse Uropoda Pereopo . s ; d h , turf y s : s propodu M corallin shor ( ovate shor d an abou A) wit n s , d , s fo d slightl d 253127 11 ischiu mandible 1; h lagoo fro t r t M. , 7 t d ro blun , (G seta four- distall l a m wit 12 one- exopo w 2 s Schott ) sp. y m n th ) abou uropoda e h of wit t , ® shorte on e ® posterodista ; elongat an nea y nov. fth shor non-ovigerou ft ( h Andama d acute B) d h t e posterio s wit r blun 3. basis maxilla of lengt t l r Pass 5 h seta protopo , e tha lengt t elongat margin time seta ; tubercle h n e e n carpu r ; on l Sea of Du (C h margin e s anterior seta d s on longe of ) posterio s res . e an uropoda s bluntl m Bois spik merus ; endopod s d wit t ungui on endopod , r 2.8 of ; shor e h , tha y propodu on r mm, posterio posterio Aldabra , dactylus l spinose ischiu margin s n exopod t latera ; abou blun wide ; bot ( sta H m h s r r t t ) . , , l . , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 wit on plumos set longe carpu seta posterio seta no mandibl wit in ¯ anterolateral posteriorly strongl 2.0 16 F agellu ig. g a h h ± Marc Diagnosis fre dentat . . singl 3. 10 present margin r Pereopo Pereopo s tw 0 e . y m e tha wit mm r o e anterio e h setae so Eisothisto margin of e an h aesthetasc n broa , . 1986 , . latera s wit d seve . ro Telso st wide . Pleopo d d ® , Antennul a maxilla w nel endopo r d h . Female 7, 2, K-AL-99 n ; margin , of P l s propodu an dentat n y articles carpu ischium ara posterio Anthuridea margin andamanensi serrate wit d d shor . , Mandibl types d 1, si : h e of ; e s x wit t wit ram , media Integumen , propodu wit fou posterio s , ; seta . slende wit sam r h wit meru : endopo h margi h i six r USN h thre e n e s e h fuse articles anterio n antepenultimat an . sp wit dat isopod ro r s Uropoda longitudina e s r . d M n an d w dista d h peduncula a nov t no margi pereopod posterodista bearin excep of d broadl as taperin 253128 . r of t . s carpu Pereopo l dista : an expanded holotype fro ( spines n A) pereo t l d g m exopo y fo broadl l g posterio l , s 7. lo m shor r rounded th r ro on e distally bearin w article . pereopo d n dista e an w l . e Maxillipe , d 1, t India setae seta d y moderatel of setae wit ovate carpu penultimat l r rounde l g s 2. , ; si , slit h margin , d increasin n shor propodu x 3 margin sclerotize , stou stou 1; , mm Ocea , stou to s wit d exopo t ( triangular d B) t eigh y t , / slender rounde h t s truncate n posterodista posterodista si s posterodista e spike-lik subequa m sclerotized g s x dentate t d d article in pereopo abou non-ovigerou spines wit incisor , d lengt , . h shorte tubercle t wit s l Eye ; e fou , fou eac d h in protopo extensio . widenin , h l l l 2; distally Maxill s r h r length serrat serrat serrat almos r tailfa large dista (C time bear 2073 tha s s ) on d n n n m a g m e e e s t - , l , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2074 dorsu wide wit po posterio on als wit F ig. d o h h Colour. Male posterio 11 , bearin 1. article serrat and antenna m bearin . Pereopo r of : Eisothisto uropod. ro eye e g cephalon s w r g 1± aesthetascs posterodista ; Symmetrica s margin ( ro of enlarged D) 3 d w broad setae s mandible 7, of corallin , carpu ; pereonites shor , , propodu stou , . latte occupyin l l Pereopo a t ; seta thi s t posterio ( sp. B. r E wit serrat n wit ) ; nov. Kensle s re maxilla , h propodu h pleon d d abou g anterio e dens 1, r ( mos pigmen posterodista A) setae carpu y ; , t e ( an t uropod F s ® whor r l of ve cephalo ) slightl , d margi t s mandibula stou latera M. triangular line time l s of t Schott y s l n n an serrat s seta l aesthetascs mor on an subequa margin d longe r d telson . antennules e e palp , antennae e Pereopo wit tha posterodista r s ; h tha l . n (G of in ro , ® dista ) n ve cephalon ; w d pleopo lengt ( , wide B) 2 of time antenna l simila fou ¯ m h l agella , d antenna s seta to submargina r 1; . longe r shor Antennul posterior ( e . to H r an article ) ; r pereo t telso (C d seta tha ) on n n m e e s - l , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 easil hook-lik number Malaysia India F ig. Remarks 12 y n to . Ocea , e Eisothisto separat , howeve spin in . n possessin e species 7; Th e on r ( s e D) thes (fou corallin Anthuridea presen th , e e g r l bu pereopo species mesia mediodorsa agains t a t to specie sp . l som d nov . n t surfac 1; si isopod s x . e ( : bear E l ( in degre A) ) spine e E s of s m l . pereopo littl fro e pereopo corallina s th resemble m on e e resemblanc uropoda th th d e d e 2; ), India 1; telson ( s whil ( F B) E ) l . n exopo m e e . besar l Ocea pereopod pereopo Thes to a characteristi an d n Mu e y of spine of d È ller 7. 2; E th . (C , s e besar 1992a ar nin ) c m e recurve e pereopod fewe , know , serves fro 2075 r in m d n Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Heptanthura specimen 2076 fro latte pal pereopods phuket diae nevertheles pereonite ovate margi slightl posterodistally propodu margi or expanded Pereopo bearin wit posterio Pereopo pal eigh wit rounded an MB exopo dista carpu Maxillipeda y d m p h p h Etymology Diagnosis Etymology Kensle Remarks Colour Diagnosis Typ C t 6 r set Kensley l of is fe , on articles s Australia n n ar y d longer g ree seta margi a w wit e muc is dentat e ® e r havin longe d d muc , s fringe s s ve at f somewha materia , margin scattere surprisingl ® aesthetasc . 1 , y h e rectangular wer wit 2, ¯ 1± s nel bu h Kensley l articles wit at . bas anterio an . described , h n . on . . Spars pal 7 r longe Pleotelso g 1978b h carpu , e Termina . . t y e d dentat , d , Althoug / shorte h an Phuket tha severa stron e shor serrate Non-ovigerou Eye taken whic serrate p propodus tw , scale l Th Th Poor fou d . of d wit of t e n , , r o r s s setae , pleotelson e e mor , t 1978b y termina H r g in unguis h e h endopod margi is ® r squa scattere s speci . l elongat triangular termina , , e / speci dista l olotype , ve present simila , n distodorsa serrate. ver somewha di give . know h th tha on articl wit Thailand wit ( e Genus , lengt Antennula Œ 198 articles e : slender onl y . somewha erence t ® 775 n h n l n h ® Ne . posterio Heptanthur dentat shor c Pleopo l article c B. 2 r n tha e muc y stron Pereopo e d shor posterior h articl l nam . ) , s . . to w nam of fro : Heptanthura Kensle articl a abou bot recorde bifringe female , t t Mandibula t , red ZMUC s , Zealan , l h th th fre mandibula wit singl t m ( resemble termina whil e e ar g intertidal s spike-lik e h Figure e d r shorte e t e dentat r ± set derive spike-lik squa e e e tw small t h brow onl contributin ¯ refer expande typ d 1, surface twic apparent anterio y : e e , a wit agellu o a d d ver Pleonite d th 7, an , posterodistally y shor specime e bot specimen t phuke an s r h l species , s e non-ovigerou s seta e e othe n in d species y s carpu e tha articl bearin 13 r antenna , fro greates d to tw h th set r Kensley m M. e t lobe , coll appearance shor r squiggle , d e pal brie pal n e extensio o r m ram margin shor t : dentat 14 a th of s proximally specie th Schott presen e anterodistally sp. n s posterior g ; . aesthetascs . p 1± th p e an , ) t g ¯ th N. small s e roughl thre Telso to i t of especiall t bearin y fou e l fro anterio corallin 4 d secon nov. e width , of formin ¯ squa Andama e describe L. s operculum subequa 1978 uropod ; agellu s thi mor e m n t e r propodu ; ; of s set n species Bruce dactylu , setae thre on articles on endit s y Ne m d ; g carpu , t lackin th a e b r specie ; tw posterio triangular e y articl g thre 4. m set latera . e w dactylu e elongat at margin dorsu s o ; in alga d Antenna 0 genus e operculum n . l , ha Zealand a endit an . mm in st articles lacking s e , 22 an s s Sea g Pleopo posterodista th . e s s at i wit penultimat wit setae d wit e l Œ statocyst length Uropoda of tw e m Novembe is margin , , unname r pleotelso , . e s fro e ; bas h structur h h fringed H o margi st th , available no basall propodu bifringe . thre l pal stron , . a fewe . of . wit m e d e Maxillipeda novaezealan- Heptanthura ¯ t Pereopo , . mandibula CRU-2001 or agellu m pleonite developed 1, of h whic ; e Uropoda n cephalon . r d l endopo y g , straight seta e e n anterio l r broadl articles exopo exopo antero unguis absent access specie broad d articl 1995 of of angle , h s m e it d set no th th th s on of 7, is . d d d a y 5 e e e e s r r - - t . , . l l . , ; . , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 below). Kensley aroun seve basa Pleopo Pereopod of Malacanthura Phuke F ig. thre Remarks Diagnosis Etymology n 1 l 3 ( mandibula d statocysts ar t . I e d ) Island , e th articles uropoda Heptanthur 1, s 1980a fro e 4± exopo . souther m . Barnard , 7, . Of r ; Sout , th . Eye endit l palp Th ha carpu e protopo d th a typ Anthuridea s s n e h e operculiform , ; phuke bee present e ( 1925 speci Africa nin Cap E e s shor Genus ) locality n rectangular d e pleopo : t e specie ® an recorde 133 sp t c , coas . or d . wher name Mandibula ; n nov. Malacanthura endopod d Kensle . absent s t isopod 1; . of d e (se : Pleonite ( , , the F ( fro Malacanthura A) e use y ) wit . . pleotelson y Kensley Pereopo an s m m d h hav fro r d dorsa th as s pal anterio Schotte m e e 1± Barnard a p fairl th tropica , l 5 d ; nou of view e 198 (G 1, short , y India r recorde thre 1989 propodu ) n restricte 2 margi ; maxilliped , l ( in fo , B) 192 e India : n fused r 43 articles apposition Ocea antennule n 5 d key) . d s fro n subequa . subchelate distribution ; n Ocea . Pleotelso m ( . H Onl Maxillipeda th ) ; uropoda (C n , e y l India is (se ) M to antenna take n , . e expanded s posterior mombasa wit n Remarks centere l n exopod Ocean h l fro 2077 ; pal tw ( D) m d o p , . . ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2078 in abou Pleopo lar severa ally wit surface propodu articles articl aesthtascs midlength margi longitudina longe stations 50 USN 50 F g ß ß , ig. h 36.70 05.30 Diagnosis Othe Typ . at abou t e M severa Pereopo r 14 n l 11 bearin midlengt d , . tha e seta , . 253269 Pereopo evenl r articl ¾ ¾ s E, E, dista Taru t 1, . . materia materia Heptanthur expande n fou e l l Antenna Antennula 7 endopo 1. basa simpl ridge an . g y e m, d t l 5 r ro plumos , 2 Bay d m, h time rounded Male 7, d on T w longest l st l l of , . e aru 2, width d carpu e Taru i examine of , latte d Œ seta s a l carpu mesia 1.5 proximally H : t longe slightl robus 12 phuke ¯ e l r olotype Exoskeleto 15. Bay agellu ± margina e r t , , , s ¯ st 7. wit on ¯ Bay dorsall agellu latera 2 s l rectangular Malacanthur i anke 0 , r t Œ d t mm triangular y margi m, B. Persia . h sp tha seta posterio setae m , shorte ro . Persia Kensle . , One d , l Persia m nov l n of y on e w setae pal USN margi n by ( n wide ; n on strongl Figure of of . e mola seve of : r m Gulf moderatel r shor non-ovigerou n ( , posterio n , y te . than , A) l nin M , margin endopod lackin Pleopo convex a n Gulf almos te n Gulf an n wit r , arabic pereopo s t e n y shor wit short d articles 253268 Saud , 15 latera h convex articles non-ovigerou , an , M. g h thre , Saud t , r Saud d , t fre y d 16 , shor a margin pronounce ; thre seta i wit 2, Schott rounded d l Arabia latte abou indurate sp. e ) e . , slende ridge 1; s i , h anterio , i stou Mandibula e e t m wit Arabia Arabia article ( nov. ban squa on l r B) time 15. t e ; 18. s slightl h t propodu hal , r mesia pereopo d . posterodista in 3 s Ma stron . copulator t 7 r Maxillipeda s d mm of f s m , , dentat margin mm Telso posterio longe , ` 1981 widt Ma y 2± y shoulder dens l 13 r , g 1982 , longe margin 8 st d y s pal rounde n h ± r st juveniles a e 2; narrow bearin e ; 1982 1982 a abou 1S2 y of set . tha p propodu r (C seta r style l half ’ 5S3 l wit a ) tha exopod fringe . n , . pal . jus d pereopo Pereopo posterodist e t 25 g P h wide , , t , n middorsa , on ara p 2. rectangu t ß articulat whor posterio 05.29 termina fro 26 exopod s of 5 d beyon linear mesia , types , ¾ setae 30 time m thre d wit wit l d ¾ ¾ 3. N, N, 17 of 1, h h d e r s - - - , l . . l l . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 locality in in a appendages Kensley Pereopo male tha notched Uropoda F ig. distinc th A n Etymology Remarks Female 15 . . e basa ( arabica Antennula D) . presen l , pleopo , , d t th Malacanthur ape l maxilla 1980a l shoulde 1 e : exopo width . . as Resemble Persia x t . Th d . narrowl Th species in Malacanthur , 2; e ; . d Th r e fro ( r mos male (J E n ¯ maxillipe abou is Anthuridea e ) ) agellu m a (o . uropoda maxilliped lacking t s speci y arabic , r Kenya easil mal rounded bu t Arabian thre m t ® e a a y wit d c , l of in sp. discerne , e in arabic whil protopo name n ; h in overal time nin ; ( ) isopod th nov. F fewe endopo Gulf e genera ) e e s a th mandibula s Kenya articles d r ( d derive l e longe A) bear . seta an s proportions di latera d fro l d Œ roughl appearanc s erenc e n l endopod s r m dorsa . on a specie l tha r fro Antenna th ridge palp clos mesia e e m n l y lie view India triangular s . ; e greates . th s s K, (G is Ey e resemblanc exten l e in ; l proportionall ) surfac an uropoda e ( n genera ¯ B) mandible th somewha agellu Ocea d t d e width in , telson furthe l e antennule abou l n l of th m exopod regio e ; e , propodus ( of : t t to r H distall structur in 1. smalle y anteriorl ) n seve 3 squatte M ; pleopo M . time of (C . . y n ) mombasa mombasa r th e . strongl antenna articles s tha d e y of longe r 1; 2079 tha tha typ n th ( in I n n y e e r ) . ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2080 Seychelles juveniles Seychelles st 1984. re basi pattern materia Ocean pleonite roughl shor Mandibula sent Mesanthura F ig. a d Typ Remarks Diagnosis. corallin s . K-SEY-22 16 t Antennula fo or P , . y , ara e l r separatio s , absent specie Malacanthur triangular materia distinguishin an fused st , , types r Barnard a 25 e 25 d . pal alga K-SEY-23 poo , s . Apri . Nin Februar r Eye p Telso Pereopo : alga of l n ¯ USN e . of r agellu , e , fro of Mesanthura l s 1914 a H l descriptions 1989 specie wit thre present n g arabic m olotype which tur M y wit , thes : h m d ree alga Genus e 343a f 253164 1989 . 1, h Mesanthur of s anterio articles USN a on f e , B. . tw propodu of sp l ¯ ; thre species usin Persisten . tur at : Poor . Kensle o granit Mesanthura USN ar Mesanthura nov USN M , , . basa 2; f e tw ( 1± g e r . Morphologically fro Figure articles e . ( 253165 Maxillipeda o ver clea : 1. D) . margi an M e s M a ( m non-ovigerou l y 5 A) t subchelate d y statocysts boulders quadrat pereopo m, an r ree species-speci 253166 253163 Le similar s m ke . d n , Mah 17 w pereopo f Antenna y fou Barnard M. crest hav shorte Ton , characters 18 d a , r , e l Schott . ,  e 7. . sp. , ) , , ovigerou on pal non-ovigerou Beach, Tabl 1 1986b expanded d 0. bee s , r m, e l 1; 5 nov. m p , ® asid ¯ tha m, e 191 n c non-ovigerou 5. e ( agellu of Ans B) 2 : dorsa 2 , describe 90 Mahe n e Ans 4 attempt mm thre is s ; e fro posterior l m . Mu pereopo di m e Pereopod l Mari , e  8. m  7. , È Mari s pigmen short ller articles 0 Seychelles cul 0 d m s mm th mm , e 4.6 fro to t s 1993 e d e , . Louise du , 1; of Louise m , dorsa ± m provid t s Pleo ; st te 6. (C e : patter 7.9 fe 4± endit a n 19 th 3 to w ) , mm juveniles J-SEY-2 7, e . n mm, pereopo 30 , l , articles lac e India carpu Mah Mah colou e n short , Apri som k ver pre tw sta of e e o n   y d e r s - , , l , . , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 M. M. stou at carpu Dista shor eac of 2. Mouche USN alg 1984 st Seychelles J-SEY-6 M. [ M. M. M. M. M. M. Kensle Rottnes is Kenya Mozambique 1980a Maldiv Sout Sout Rottnes a 7 midlength seta a sam h quadrata protei nigra maculata gerlachi dimorpha catenula bipunctat albinotat Diagnosis Othe time K-SEY-15 t t . M bearin on h h l s termina dentat USN ) e articl e y ; Afric Afric triangular e Madagasca s t t , Malaysi r species. , of Mu 253169 an ree Islan Kensle Island Island basa Mah materia , cora d Wa È a varyin M (Stimpson 1 sp g a a a ller e f ( e Kensley , Poor sensu Thomson ; . Thomson È Ma . tw d of l bearin ¯ gele, , l Madagascar set e nov 253168 ,  l ] articl at , , y , a width cora , 199 o Female W. W. maxillipeda Seychelles y rubbl 1980 a , e , l tw g T aesthetascs . r Kensley, 1981b distall 1 ( 1984 abl examine Seychelle 3 posterodistally , Anthuridea lengths 1982 Australi Australi l o g e m, , 198 Keny a , rubble , , e , si 1855 bearin e posterio juveniles 195 195 2. tw : x . 2 ) Ans y Telso betwee setae ; recorde o a 1 ) 1 . truncate , Feature a a d s non-ovigerou , l Antennula e 26 . g . 2.5 pal n . Mandibula Mari Antenna seve r n , USN Pereopod Februar n articl d fuse margi ± p isopod st 5 s . M brow n subcircular a , e m, distinguishin Pereopo . d e M ridged fringe 11 12 K-SEY-24 Louise protei ? ? 7 9 7 6 4 3 n dorsall r Ans l y 253167 n seta s evenl r ¯ ¯ s s fro 1989 agellu agellu pal 2 algae d e ; m fro e d , an m propoda setae , p a Á y y , Mah m 7, on . la d , g th wit convex wit m , m , on Australia 3, Mesanthura e carpu Uropoda e Mouche ; cora 1 notche notche notche notche notche exopo h sinuat sinuat sinuat sinuat of of h e juvenile e lamin India propodu  m, , fou pleonit non-ovigerou l fou fou Seychelles l , pal s Ans ape r d e e e e rubble d d d d d n wit r a r ; seta l m , ther articles articles Ocea , dentat Mah x e h e s st species. wit subquadrat doubl solid soli variable soli soli irregula narro wit rectangular e 6. an open pleon head ova pereonite pereonites bar or nite fro pereonite pereonite 4± 1± a e a anterio , Á mesially h is n , d d d d K-SEY-21 e 7 3, h Telso 1±  unlinke m la s , l , a s , 25 . lo som square continuou block w pereonite Colou e , wit cluste s on on posterio Seychelles an , Mandibula tw area hea 1. of pereonite w linke r ring Mouche , eac m d 5 h Februar r ring ope pereonite o , e hea n ® rounde d s s s m, pleon ope s tw doub h e margi . ve d dista s r an s an an lengt r to d on ring d n Pereopo on , wit s patter longitudina on o of r oval d d d n wit Ans serrations s on ring an , telso s ban head juveniles , pereonite middorsa t h s pleo pleo pleo corallin s , l head si ban , or d h n y tha h d an pereo on ovat article s s x s r 1 e Mah n d n abou singl linke muc toot on 4± 1989 d on n n , n palp pair d 2081 Ma t hea a , on d Á thi e 6 - 1, la e h h l d y  d e e s s s t s . , l . , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2082 exopod distally posterio shorte F ig. 17 pereopod pleopo mandible r . . , tha r bearin Pleopo Mesanthur margi n d posterior 1; ; 7. g ( d ( E n I te ) ) 1, a n bearin maxilliped uropoda quadrat dista endopo , wit g l h l a plumos numerou exopod ; B. d sp shor ( F . slightl Kensle nov ) t uropoda e ; squa . ( : margina J s y ( ) y A) shorte fringe pleotelson t an set m l d protopo dorsa a M. l d r posterodistally setae than scales Schott l ; view ( d K . , an Uropoda ) , an ; pereopo d e ( stou B) d endopod abou antennule t ; d l sensor propodu exopo 2; t ; 0. (G ( L) 4 ; ) y (C pereopo d time pereopo s set ovate ) rectangular antenna a s widt postero d , d latera 3; 1; ; h ( ( ( M H D) of - ) ) , l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 longe wit seve Uropoda Pleopo wit pods sutur Maxillipeda dorsall an Panathura dorsu exopo Le give is undoubtedl confusio author resemblanc protopod. pereonites rounded margi F ig. a d w h h Male Diagnosis Etymology Remarks Colour n n 18 specie , robus 6 Ton increase e r m n or d a propodu . s y d betwee tha usuall distall an ne of , n mor (se : 1 , distinct Mesanthur l Barnard slightl Rostru , 1986b . s n exopo w exopo t d e th e y wit pleo dista l of . rami pleonit e name . d endit y Fairl e masquerad Poor to y n . articles h Accalathura s animal Th numbe longe ver article y n Bod , . d thos d l somewha . m Th , fro bot seta longe Eye y an e e e operculiform . a 1925:143 lackin y Anthuridea e stronge distall presen e M soli quadrat y an m d slightl r h e . 6 s . . speci s r pigmente . Pereopod telson d Pereopo tha Ne specie r no d present 1 quadrata of th e g tha Le rectangle an y unde w e t t n t ; seta y , a serrat ® r rounded visibl w swollen Kensley species c n d thi Sout sp emarginate precedin ; nam tha s Panathura nam blob 2. basa Ton . e . r s n referre d d , nov. s n thi on Mandibl e endopo nam Pereopod y h 7, isopod e bear or mor in latera , s e s , fuse l W , s , propodu : ` palm , 1986b wit of width 1982:159 maculata not of refer name female pal ( ales reachin e g e d s A) d is red elongat h pigmen pleonites som , s . m l d to Barnard , . dorsall apicall e s Pleonite to it lobe fro contributin . . : bu ± Pleopo s l s wit cephalon to wit As . as brow e 88 s 1± be . m g soli t Eye ’ wit resemblanc . h th ha ). to h e 3 Haliophasm M t th avoided y an y d tha fe e ; on mola di s h s n Th pal , . e d s rounded wit w rectangle pleonit a d enlarged 192 quadrat ro Œ pigmen ` ; India 1± n 2 maculata e erentiate uropoda p mor ( mesia w h B) in g th appendi r 5 latter articl 5 somewha of th e , blunt free female e n e henc e e serrat l strongl pleopo fac ; t l e Ocea a to . telson e , s , endopo 6 dorsall setae Antennula pigmen maculat ’ 3; l d however articulating ; of t , x e M wit exopo ; pal fro pal describe e pereopo th masculin tha n pigment . d t . seta h . y e p dianella in m 2; p Pereopo y notche t d posterio presen t of wit proxima (C a d posterio on e distall , severa patche Haswell anterodistally an ) poin thre r h d d cephalon , a ; ¯ ® d 1 bear d l by pleonite t agellu Poore pereopod ve as d e y propodu endopo t uropoda specie r l s l r 2 broadl articles articles ou lon variou margi region specie s on , pereo an 1881 som t m 2083 g and , d s th th s al or of d is 1. n 3 5 y e e e s s s - / . , . l , l . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 3. Picar 2084 F ig. 9 mm Typ 19 d protopo ( . E , Is. e ) tw Panathur materia maxilliped , o Aldabra d non-ovigerou an l d a . indic endopod ; , H ( intertidal F olotype a ) sp m . s ; pereopo nov ( m B. J Panathur ) 5.4 , : . Kensle uropoda : 20 USN ( ± A) ( d 6. Figure Marc 1; 0 m M mm y antennule a (G l an 253131 indic exopod h s ) , d 1987 m 19 sam M. a pereopo , sp. 20 ; ; , e . ( Schott ( non-ovigerou P dat B) K ) ara nov. ) antenna a pleopo d types as 2; e holotype ( H d ; : ) (C 1; USN pleotelson s ) ( m mandible L) 6. . M pereopo 7 mm 253132 ; ; ( , ( I d D) st ) a 7. uropoda , maxilla F4-87 on e , l ; l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 lengt Pereopo ungui carpu midlengt dentat Mandibula article margin margin wide lo clumps fro F ig. w m Diagnosis Othe 20 triangula r h s s nin tha s a . of . . three-quarter short of d r Antennula Telso havin h eac e Panathur n res materia 2, Halimeda station r long of h t , pal . propoda r n of g pal triangular bearin rostrum , wides Female p ® pleonit dactylus a p s ve l (C of indic r aroun examine Anthuridea articl ) s peduncl serrations an g thre t l lengt at a , l pal : d e pereopo singl ; sp smal Bod e . 6 d e rotte abou propodu m . Pereopod longe 3; h articles Aldabra nov d e . e y l wit of pal n of wel t ; . n te d aesthetasc : USN res midlength h r wood mola ( n 1; isopod p thre A) tha l , thre s to t ( pigmente of , articl s D) of barel M fro n e r eleve 4± , l cephalo e ® precedin articles pereopo 0.5 dactylus low s ve m 253133 7, shor e fro . y , ± n ree carpu 2 , articles narrowin 26 Antenna in m d time rounded t longest n f ¯ ; m. anterolatera d setae an th cres ated , g ¯ , 2; s agellu si e segments s d wit wit , x antennae longe ( India t , ; E apica ; h l an non-ovigerou g pal ungui . h ) inciso stron m posteriorl ¯ Maxillipeda d anterio n agellu m r l pleopod shallo l of tha , Ocea l s ; articl bearin wit g r ( eyes ® slightl B) n of accessor ve m r h w wide n pleotelso margi e fre . y thre articles 2. slop g Pleonite of s tiny to y l e tw . m e dorsa endit mor rounde , Cephalo seve e y n . o cusps thre cora n , set shorte Pereopo shor e thre an n e s l a tha e posterio 1± d reachin l ; d at articles t e manca uropod rubble lamin n 5 r n setos setae dista base 2085 tha free wit d hal 1, n h g a e r f . . , , ; l , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Antennula rounded. convexit form Pleopo posterior 2086 lagoo 253126 cora ± 2 1 India short tha propodu Maxillipeda ¯ Pendanthura wides wides subcircular tailfan genus, broade articles copulator at plu lu ± agellu m parallel-sided Maxillipeda taperin Maxillipeda Pleopo Pleopo . proxima s n Remarks Male Typ Remarks Diagnosis. Etymology l of fou . , n posterior t t n rubbl . . pleonite Uropoda d r . , at posteriorly m Ocean P e nin nea r Th Uropoda d y d g on s tha : . 1 , materia . y evenly-space midlength 1, subchelate somewha 1, on Cephalo serricauda r wit e slightl of , l r e e e Menzie style l n ¯ . . third exopo onl whil articles . Grand l l ovigerou o uropoda agellu . h pal latera in , . pal pal Œ s . tw Th The posteriorl y l tw Eye . Pleopo fused t female Picar l e y p Th p p exopo o , exopo . l Ke d , e articulatin s o tw t exopo n large in Genus 1. broadl e 1. m . . of an l . s an , e genus articles presen H seta , y 6 3 o margin Poche present wit Pereopo wit . ( speci d expanded P s d l of d d olotype to Barnard d . . time Telso singl muc time d . . d endopo m r exopo Glynn Is. y fringe h h e operculiform nin d indica 1, tha th subtruncat slightl y Pendanthura (damage . . on a , ® t Pendanthura wit acutel s s h , . e e Pendanthur rounded c e Aldabra n ; exopod narrowly-rounde . B. g specie Aldabra longe n longe smalle India d , . . Antennula Mandibula nam article endopo broa latter d h d wit d : at followin 1968 , Kensle . 1, it mesia setae y formin USN in 192 Pereopod y . . h r proxima r e mor propoda is s d d n : triangula ; tha tw e tha r th , refer s ), 0 , ; 31 , ( . . di Ocea . propodu elongate , fringe article or d Figure M ), l bearin Bod wit e 15 o Pereopo y e n 0. g n ; Œ tw margi g a Menzie subequa Kensle . . . er ha earlie statocysts mos operculum an 5 r ± tha wide exopo s wide 253125 h r pleopods picard o y 18 m, ¯ n s l s to s d l serrat d agellu lengt . . . non-ovigerou ; pal n 4± thir specie g fro t r m, no pal 21 ; n M. ; seta 6 th endit ± d r s whorl y conspicuousl twic Telso 7, rostrum d Telso s d ovate . . . p proximall t m Apri i ) e wit ape d m h specie an l 17 , an 2, Schott sp. e previousl carpu / e an m India in ovigerou widt ; of . s of d . . . th , e wit e margins h distally d propoda Bod n x Marc n d wit of of s l Schotte . lengt e nov. longe tw small Glynn an mesia lackin 1987 of h . havin . h s singl endopod Th tw ; Pendanthura typ n s h seminigra y e o rati d stron is wit aesthetascs eye 5. y h Ocean exopo e s o . picardi h seta an e r . . y 5 , s distinctl e g o ; broadl g m to , telso l 1986 y h Pleopo , to l s siamensis time tha an 1989 in m triangular bee greate margin ® ® 196 pal g entir 3. anterio ofte in thre e large ne ne 4. basis d , P 5 proxima d . n on n . n s sp sp m 0 8 operculiform : . th mm somewha margina onl margina P n longe y e wide 56 e mm of indica . . recorde r d y ara e wit , posterio articles nov nov . , margin tha rounded ; . y reduce excavate 2, , r wel structur Antenna sp distall P. exopo , h st margi types . , r n specie . . . endopo st l a tha enlarge l wit Aldabr nov Aldabr Pereopo serricaud si it l l a robus d pigmented K-AL-116 x setules setules . t n . is K-AL-98 d y h r . Antenna d fro : n operculi , wid e s Thailan . broadl hastate margin USNM l distinc . . shorte of article almos Telso a a d of in t ¯ m d Pleo e agel , , Atol Atol seta wit d . set tw a th th th 1, is n o h n a y d 2 e e e r - t t - . , , , . , . l l l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 articles thre width Cephalo F ig. Diagnosis e 21 (J maxilla , articles . . posterio ) n Mandibula uropod P endanthur wit ; , . ( h thre E r ; broa ) Ovigerou ( margi maxilliped K e a Anthuridea r ) aesthetasc d picard pereopo pal triangula n p broadl s i ; consistin female sp ( d F . s 2; ) n nov r on y mandible ( isopod rostrum : L) rounded . bod termina : g pereopo ( A) of y s ; m singl abou . (G fro , dorsa Telso l d ® ) m article nel e pleotelson 7. t th ver 7. l n y view e 3 abou setulose y . India time Antenna shor ; ( ; t B) s ( n H 1. t longe . Ocea antennule articl 7 ) Antennula pleopo l time ¯ r agellu n e tha s bearin d longe ; n 1; (C m greates ( r ) I of g r ¯ ) antenna agellu pereopo tha tw tw t o n o width setae shor basa m 2087 ; d ( D) of 1; t ; l . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 posterio stron fre wit 1, bearin lamin 2088 posterio posterodistally Pereopo ungui distall endite mola dista numerou articles shor thre midlength margin wit 2 pigmented rounde tha intertidal USN P coars typ 1, endit noted below pleon width margina 1, wit ara an ther endopo e carpu h h h e n Remarks Colour Diagnosis Typ Etymology e d t anterio l e r types locality M fringe g irregula e greates , telson a : ; s articles , y ; articl 3. g setae e is short telso , endopo s d dentat e th pal in san equa dentat 253120 si r r d l ar hyaline Pleonite s articl s taperin , broade materia setae e margin x rostru ; d margin 2 . . e it 23 distall p setae : d d e dista ; , r n telso posterio s Stron les USN as , mor . , t l of scale inciso , r margin . Irregula formin bearin betwee an Marc e bluntl Picar relativel e widt . in a . basa d s . ban lon , Pendanthur Uropoda . singl , m set Ovigerou l g d r Pereopo robus on 2 n e havin broad , y l Mandibula s Th ; M setae . lengt g acute s anterio an g to seta uropods 1± h a d longes is r e rounded propodu d reachin l h on anterodorsa g y r e g as ; e H d ovigerou 253119 n articl of of at posterodistally 5 Island 1986 r broadl narrower propodu speci severa y triangular hal t broa e g olotype , ; h th raise , posterio fused red ; cora tha endit seta pereonit an thre on elongat subcircular eigh e d l to propodu r t f s e exopo . ± d antenna a margi g Pendanthur . ® 7, n d of an d female brow , th , l e , s equa e r res y , c l picard 0. e t B. pereopo par articl as thre projecting togethe s on heads carpu e pal d ` seta name wit cusps 4 telso rounde s broadl collar serrations : , m t r propoda e Kensle l no e time d . USN broadest t n fa th n margin l 1. p mesia of h e articula bod s Maxill e of elongat 1. : shorte to e i reticulatio r t e 3 e articl n ovigerou , s , . severa Integumen ; bear . an in in bearin ’ mm dactylus an s Cephalo ( Aldabr anteriorl use uropoda r y d wit lamin Il tw wit M Pereopo d y Figure ¯ ¯ at a widt d equa y l triangular e d 1, ated ated ; form o apex , seminigr e d l , s surface h an a r h 253118 . tw ; antennule bas r Picard thre propodu st e mesia som propodu abou l tha dista as Maxillipeda anterio g h as ovate a ¯ hyalin hollow d a o l fringe , , a s agellu , e seve in , of . n K-AL-109 a wit 22 M. dentat n e n palma aesthetascs y l e d ® m Atoll bearin wit t Severa on of l nou t fringe posterior gured lengt l exopo resemblanc . broade ) exopod , , as no h 1.5 7, a 2. article , surfac Schott n Pereopo , ovigerou e d h abou Aldabra r s dorsu s telson fringe wit 5 m sp. s t n seta s . carpu ± margin anterolatera fre a betwee r scale strongl wit slende time h indurate 1. g ar d in . l h of d wit are nov. to e 8 l r Maxillipe s t easil tw e e e seta , singl h e , apposition m d mm ; is 2. pal margi ; tha bearin s together ree . s distally tha singl mor pereonit s h a stron d o propodu latte 5 scale longe s , of Antenna r . n on rounder e e bearin y y fou 2, p time hyaline larg f n m e , n 3± elongat Antennula to posterodistally pereonite cephalon . ¯ e in sam see termina e 1. of middorsa carpu in n at r g Bod posterio 5 s r elongate g P ¯ r e 6 shor l s . m, , serrat d on abou wit n ated margina . eigh th ; singl tha mm e g longe Picar e Pereopo statocyst corners siamensis s ¯ , wit y di , 7; , e l dat h fringe posterio relativel agellu is s e 22 t n th peduncl Œ 5. Tha almos t , , l pleonit t wit e h accessor sinuou take , e erence s articl wide seta st dista 5 d pal a e r r r hal al . l sensor Marc broa 1 lo a tha Is. time h In margi as i maxillipeda length l peduncl l . d m m d w an K-AL-108 species pereonites f n . s serrations t ver , , Eye l e . n Pereopo y holotype pereopo described scale r lengt 1, Aldabra wit at e fro s of d d e rounde plumos s straight Pleopo sinuous s h greates slender margin bearin y 6 y y ca carpu n of 2, latera longe articl s , abou m h singl fuse 1086 shor spin n s seta wit h e plu . wel tw an ® th on be ve of of d d d o d d h d g e e e e e s s r t t t , , ; l l , , . ; l , . . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 tha maxillipeda uniarticulat (se reticulatio it Cephalo of exopo margina exopo seta stron F e ig. e endopod 2 n Remarks Colour Male . 22 Kensle anteriorl g in Pleopo an antenna d d . serrat d an elongate-ovate abou l n : Pendanthur lef plumos Unknown n an . y l e y ; t of . of d d ; latte endit Anterio e t uropod y 1984 mandibula (C pereonit 1, two-third pigment Th th pigmented sensor ) e e r bot e e mandible : setae distall a earlie Anthuridea presen ; is . 18 r h seminigr ( y H mor ) e , . ram hal ; ) set oute s 1 r in r y endopo pleopo ; an in t species f ; e pal a rounded pereonite i specie severa ( a reduced d r length D) of formin posterodistally p sp propodu margi n d maxilla . is anima isopod d . 1; s nov l di , wit longe respects , g ( n ; s an I Œ wit . ) : th operculu 3± er crenulat h ; l s d pereopo ( s e h ( A) s of fou 7, r mor E slightl fro fro endopo severa ) tha pleon pereopo : ovigerou r ; maxilliped m m th e relativel dactyla n e d m th e y th heavil wit l 2; , in e narrowe pigmen e d bu margina pleotelso (J India thre d h s of an t ) r 1 m fringe y ; y endopo pereopo pleopo y dar e ( ungui dorsa shor F n pigmente t previousl of r ) l Ocea k patter n tha pereopo d setae blackish-brown t l th an s setae d d d dista view n e wit 7. d n 1 large . n endopod earlie uropod d is y ; h , l d is ( describe reachin setae B) relativ tha shor 1; r distinctive r tha (G antennul n species s t . , ) n wit e Uropoda posterior wit accessor g pleotelso ; d exopod broade pereon to h specie h e pale , ; 2089 bas nin an th th y d n e e e e r r s - . l ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 th 2090 F Expedition ig. e Typ Etymology blac 23 1; ( D) . e (J k Pendanthur mandible ) pigmentatio materia , l 7 pleopod ß . 55 Th ¾ l 22 ; . a ( e ² E siamensi N, H 2. specie ) n olotype pleotelson of 98 Pendanthur th ß s s 49 B. e sp name ¾ anterio 45 Kensle . : ; nov ² USN ( ( E, F Figure , . ) fro : a 13 ( r y uropod A) M hal siamensi m an ± 18 s d th f 253121 23 l dorsa m, of M. e ; , (G Lati 24 th s coll Schott ) l e ) sp. view , maxilla n animal . nov. meanin Gallardo ; l e ( 3. B) ; 2 . ( mm H l g antennule ) ` maxilliped , , half-black 14 ® ft h Februar ; (C Thai ; ) ’ ( , I ) refer ± l y antenna; pleopo Danis 1966 s to h d . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 5 Antenna lu rounded pigmented 4. m coll ovigerous sam P F , ara ig. six-seventh m thre Cephalo Diagnosis Othe . e 24 of Gallardo types dat . e thre juveniles r . l a Pendanthur Antennula peduncl materia . : m as e n Telso USN , articles s . slightl , ® holotype lengt 3 ve Male Februar , n M l e ® ( juveniles D) a lengt examine h ft Anthuridea of , y 253122 r siamensi h penultimat : of peduncl wide Bod ® . Thai l ve articl h pereopo y slightl 1966 y r articles , , d ± s . abou Da 16 ® tha e sp e ft 4; n . USN of h d . n e n USN y nov isopod , ¯ l an 1; Program t , Thai mor thre midlength agellu 22 7. articl ( d . M E : 6 M ovigerou termina ( ) e e ± time A) s pereopo Danis 253123 articles tha 253124 m e fro m 3 , of s n localit antennule m seven-eighth , longe l twic h singl wit s article th , d , , Expedition m nin tw decreasin 2; e h , e y r e India 20 o ( width unknown e pronounce tha F shor ; s ovigerou ) ( non-ovigerou eac B) , l pereopo n n on t s , greates m h g setos Ocea lengt posterio , e antenna bearin in 6 , ovigerou s ß January lengt d d 45 e n m h t 7. article , rostrum ¾ of 00 widt ® g ; r ve h s fou (C ² articl margi N, m distally non-ovigerou s h ) ± , . r Marc m m 12 Mandibula at aesthetascs 99 ; , pereopod e n two eye pereonit juveniles ß 4, 32 broadl ; h articl ¯ s ¾ 04 1966 agel 2091 non- wel ² E, 1; y e e r s - l . . , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 two-third fe singl article stron dentat pal 2092 3 ovigerous Phuke 26 non-ovigerous Patang free Pleopo carp 1 articles articles Quantanthura and th lack wel pleo abou article bearin distall exopo copulator pod form tw seta anterio w subchelate Decembe e o Jul p l Female Typ Diagnosis Etymology Remarks Colour . mesia , . s n i posterodista di wel e shor , t T Pereopo g of th wit roughl y d y wit s , a t g 2. Œ elso te termina d , . . r e bearin spin er l e 4 1994 rounded 5 of Maxillipeda narrowl Is. n Phuke tw singl 1, Antenna h margin t h l s materia separate y latter s an time . n st m fac of , : eigh o exopo r smal lengt fro e ® style i . Ey , d wit y . Thailand Œ 7.0 1994 Menzie . ve Cephalo e d aesthetascs e posterio propodu g an ZMUC . l m s 5 pentagonal t seta m ’ t of e Pendanthur 7, shor ® h s l ± seta Eye wide ; y d h t , teeth Is. l noticeabl dista ve l 9. Th irregula 8. th as d distinctiv . . tw l carpu th setose serrat propodu d Genus articulatin ovate of severa e 0 5 , operculiform e ¯ s dista . s t e e o setae l in mm mm Thailand H on agellu r an othe Pereopo pal ; propodu l present n , r article , speci res basa tha s olotype mola fou male d margin s plumos wit e coll . Quantanthur , mesia in , , . l p t Quantanthura r l wit George , setae distall r st n setae y ; st ¯ Antenn a of r latera of s e mor ® h m l mesia describe . ated a numerou a endopod les r . , c non-ovigerou siamensi h statocysts g 2. B. pigmen irregula , fou Dexter CRU-2039 bluntl Antennula dactylus d wit l , CRU-2045 nam of anterio s . rarel 5 . s jus ; e e : surfac coll Kensle Pleopo y . of , 1, slender . l elongate r l time h ZMUC 1972 slende Pereopod thre Pleo patche setae t rounded a or ( e surfac pereopo propodu . y Figure shor tw d y t refer , relativel a s r Dexter . ® s s absent e : r e triangular species pattern o n ve . . Pleopo 5 is ban y andamanensi ; d 29 elongat margi longe nea , r Menzie to Pereopo short t s dista an , r a 1, e , thin-walle endopo Octobe s articles ; . , spines of of s Patang s Poor d non-ovigerou ¯ , muc Pereopo Koto to bearin d r s d ram nin m , 25 s agellu . reachin of y . 2 pigment palm r 1, midlengt n M. . , l 26 Antennula d Siam in 7.0 , pleonite e e an e Th tha h shorte shorte pigmen setae s i in 2, 26 ¯ , . , r ; d an d seta subequa Jul articles an Schott ± large ated , non-sclerotized d g endit Maxillipeda Phuke e bot . m exopo n ) 7. 1994 Phuke d 2, , d d wit 3, d s ban arrangemen g y 9 ; th e wide Le endit of ; sp. 1, r h r , George mm carpu bas h inciso 1994 propod r telso h e s presen e t tha pal w . mal tha d e specie . propodu 1± tw t betwee s moderatel r ol of d nov. t l fou USN , Ton e , of Is. Mandibula ¯ e m d in m n two-third Is. o 5 . n wit st e n agellu of mesia s on r posterior , P nam fused r 9. an a length articles shor , , wit faintl , t in i triangular s ara l of Thailand h 1986a Thailand M endopod 197 1 CRU-2044 ovate on dista mesia n d tha pal t mm h male serrat . e s thre l female eyes m t types of , Maxill 2 253283 broa y y posterio n fo wit margin p pleonit , st : l , s , of strong exopo . th sinuous P l r 75 i , e r of ; ; Œ lengt Pereopod h termina st plumos surfac . e Thailand e propodu pereonite d wit , : thre . cusps pal ; seminigra , a , , singl setae posterodista st singl ZMUC, a coll patc coll latte is , lackin , CRU-2039 . e h i p d wit . h Œ r fou Na e distinctiv Uropoda 6 Pereopo e , . operculi peduncl . e margins ; of of to setae ; h e e dorsall r l h bearin bearin Dexter Dexter i lamin r ro ungui fallin articl ovate s g s broa . endo singl Hain setae seve s thre non w wit 4± fre and an one on of 7, h d n d d g g a y g e e e e e e s - - - l , , , , , . , l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 CRU-2046 coll 1995 non-ovigerous CRU-2041 Phuke F ig. Othe . 25 . Dexter pleotelson ( ZMUC t . D) Is. r Quantanthur maxilliped materia , , , Patang , Patang Thailand , 8 ; m ® ( , Augus ve I l st ) examine ; , , pleopo a Anthuridea juveniles ( a Phuke Phuke E CRU-2042 , andamanensi ) t coll maxilla 1994 d d t t . 1; . , Is. Is. Dexter (J . st ZMUC ; n , , ZMUC ) a ( , Thailand Thailand F isopod s uropoda Koto CRU-2043 sp ) , mandible . 19 nov. , , , thre s Phuke l Marc fou fro , , : exopod ( coll coll e A) ; m r , juveniles (G h m . Na . non-ovigerou t th Dexter Dexter ) dorsa Is. . 1995 e uropoda i , India Y Thailand an l . , view , , st g ZMUC n 6 3 a l ¯ Decembe Marc Ocea CRU-2040 ; protopo at ( s B) , , m Phuke coll antennule , n , h thre nin 1995 d . r Dexter an e t e 1994 , d . Nopoiotoush Is. juveniles juveniles endopod ; ZMUC (C , . , Thailand ) 4 antenna Marc , ; , 2093 , on ( st st H h a a e ) , , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2094 articl rally-directe clump sligh dorsall F ig. Diagnosis 26 t e s . ` y wit shoulder of distinc Quantanthur h si thre d x . setae seta t ’ e Female fro in aesthetascs e , articl m each a posterio andamanensi telson : . bod e Antennula 3 wit B. y . . r Antenna Latte abou half Kensle h s thre 4; sp , r t ( . apicall D) r 14 abou e nov y peduncl l setae pereopo peduncl an time . : t d ( y A) , 2. M. s ¯ narrowl 7 longe e agellu pereopo d e time Schott articl 7. articl r s m tha e y d longe e e of 2 1; rounded 2 n bearin thre ( wit B) greates r pereopo tha h e articles singl g , n ® t ape basa ve width e d x dista elongat 2; , l tin bearin (C width . y Pleonit l ) pereopo elongat termina e g , vent wit tw e o h 6 d e - l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 expande triangular obliquel tw lamin ( seta K-AL-54 usuall margin posterior da pal Neohyssura Islan seta pod Q base maxillipeda shoulder fro of th endopo 2. tha margin squa of 4± abou no setae posterio subequa perhap . 5 e o l 7 t Sout merus m p andamanensis n Typ Diagnosis. Etymology Poor Remarks time India , e s in pal becomin termina d t d ; of t exopod articl 4± a th on y ¯ set one- on meru lies , ; ate m h e d ® e dentat arme s 7, s r y l d e s endopo thre , ve . article n a , Africa surface triangular Bas longe feature to , three carpu an materia Pleonite . at d bearin proximally e ha on coars ® Amar Ocea posterodistally distall l s articles l ft as . d 4 , e g res d endite distolatera s s setae an h wit . a Le th mor wit article bifringe Strai in ) wit r Quantanthur s lengt ) s t d Mandibula e , n d havin e s g , articles s tha h 2 an w Th an of 1953 y pereopo l h of whil ( h , san carpu . s of elongate Q. e ; telson an abou setae , To reachin d rounded , Anthuridea d 1± t endit e fou n spines dactylus elongat h whil ischium apicall s th bear H d o , d propodus : speci remipes n greates g 6 e of 1 d Œ F pal 353 olotype e r 3 l ( an . . s t free amil no ® an e , 1986a souther seta e dactylus antennules corne elongat of te ve Pereopod Mandibula s d eac m . ; , d ® th ha wel Genus t g , d r som n Wa y lackin a Pereopo 1, . c e , th agreein y , to longe rubble e , Neohyssur e margina convex 2 h Uropoda s t dista pal bearin nam posteriorly rounded merus andamanensi È l ( wit wit e gele HYSSURIDA propoda width ) r posterodistally of n Barnard middl a ; developed havin e : divide antennula n p carpu of e resemblanc isopod h , h les Neohyssura g th e , l r ( USN Australia setae wit of , Figure 1981a s plumos numerou is g tin articl , g statocysts e , , tha d s , oute carpu r g e l 4± mandible in wit take mandibula scattere latera h , d thre s elongat 7, l y of serrations pal a l rounde n M 1. exopo , , fringe havin 7, s th al , pal accessor : h 191 e gladi articl propodu 8 r pal . protopod fro havin s 72 n l p r e e s s Pereopod e 3, carp scattere ree time features 253134 27 l m specie fro . peduncle ; an is e 4 margina p article articles s m E margi Poor Th d g bearin d . d ), . e , a Amar, to ® f articl d e of d , th m shor th i W 28 Pleopo g ne seta elongate seta s sp. wa slop uropoda maxillipe e 5 propodu r dista Q e . y wit elongat pereopo longe e a th e ) s s È Australia pal setule wit . Maxillipeda d , gele, s secon n s set . India of an e , e e nov. e ; t erica wit g e h s recorde eac Telso seta doe distall l , on non-ovigerou 3; fre Andama 1953 on p l mola h d a 1± ® Quantanthura setae o al d anterio r projectin h ve subequa termina Le Œ singl e at h 1981a , s l posterio 3 d s n tha l e posterio 1, e s usuall Poor fringe d d anterio feature on w Picar n subchelate hav seta bearin setae exopod r specie bearin Ocea on base y . an n endopo 1 n d T tapering Uropoda e spiniform sinuous anterio on is species r e basa n d fro e e mesia seta l d y d . l . densel , n margi l an Sea on g posterio slightl endit r pal s s r Pereopod g wit r g Pereopo 1988c m an Is. scale margin , portion margin d of , of l s surface relativel singl posterio d , p d th a , width l ¯ h Le r int , , in , , agellu m grou th e slightl articl surface Aldabra n l doe surface e seta y similar however ape y : muc s . latera w exopo o whic 184 e narrow Agulha e setos concave shorte Maxillipeda r on 2.0 tw Ton . d genu s , x . ; ; dista e pereopods p Pereopod s h . wit e y hav propodu propodu m r on subacute o 1, y mm, h posterio 2. l 2 attache elongat ; e , margin ; broade d , groups margi Phuke shorte h , propo of s ungui ungui carpu r an 1986a , mesia pereo l e , s abou 25 shor lack fro . 2095 tha seta wit lon are d tw sta m, m 3, d h o n n g a e s s s s s s r s r r s t - t t - l l ; . , , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2096 thre articles termina 3 seta convex laterall extendin indurate Be 1983 K-AL-38 as 15 F ig. two-third , holotype Apri Diagnosis e e Madagascar 27 . ( pleopo articles of USN . H y . , l l Telso . varyin g ) wit togethe Neohyssur , 1983 articl Cephalo anteriorl K-AL-39 M l s h . d pereopo , . USN n numerou 1; widt . middl e 253137 g P basall ( Non-ovigerou r bearin , ara lengths D) 1± barel h a M n y d , e gladi 1. types of beyon wit 7; cora y l articl 253136 , pleopo 5 g s y broad tw ( m, 2; . elongat h a I singl longe Antennula ) o l : sp ¯ tin e d Februar rubble USN non-ovigerou agellu d . longest l B. anterolatera , antennule. , e y nov 2; s on taperin r conspicuou Kensle e weakl ( female M tha e . plumos E , : m ) oute ( y , 253135 A) r n l bearin of 1964 2. y g peduncl l y ® pereopo 2 : to r ft pigmente an thre e mm l l ree Bod h pleotelso s . setae lobes narrowl s d , g m peduncula f e tw aesthetasc , M. tw y 2.2 e slop d fou articles o ; . wit o 1; pleonit elongate Schott Pleonite ± d r n dista 3. e ( y h l F non-ovigerou 2. an eyes 0 o rounde ) tw Œ 5 mm , d r l . mm e e o middl Picar uropod l setae ; Antenna article pereopo ± 6 s , basa lo slender 1± wit , st d w st a d ; 5 e ape h a l termina ; . rounde JR-29 free Is. article articl posterodorsa d ( K-AL-54 Mandibula B) x s l . , 2; ¯ , bearin Aldabra m Integumen as agellu (G e , 2.0 l s mandible d l JR-33A broa ) longest articl broad ± rostru g , 3. l m pereopod sam numerou , d 0 r e 11 of l mm as pal , margi , t , almos ; m e articl seve (C wide Nos Apri shor non p dat , no st ) of 3; n n y a a e s t t t - , l l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 anterio one-thir sensor triangular ungui 1, extendin shor betwee pal acute singl articl palp havin lamin one-thir F ig. pal m m 28 t ; e maxilliped e , g basa a s m y bearin set extendin . n termina r hal tiny si dentat d d setae g margi setae bearin Neohyssur a x , l lengt f lengt wel , dista distall pal at lengt g bearin , l ; a ; l p n base plu g thre h ungui h g beyon ( l havin seta articl E h shorte wel of y tw of spines s ) a of e g . acute Anthuridea m o l gladi lobe . second remainde s Pereopo beyon remainde seta fou d uropod; e g Pereopo shor slightl r short propoda , ® d a tha , r ve e termina sp extendin an setae t , d plu n margina . , sensor y wit d proxima ( d nov havin posterio r F d r mor s n marginall ; h ) of l 2 of 1 . transparen l isopod stron m palm : singl subchelate spin g e dactylus y ( subchelate pereopo dactylus g A) l tha wel fre setae l serrations r g ; e pal e m margin s n e l propodu strongest endit shor y antenna fro lateral beyon m hal , d . serrat . t wit m of Pereopo 1; , f t Posterio , ¯ e lengt (G , termina h carpu propodus th ang carpu d ; , reachin ; serrat s e e bu mola lobe ) ( . propoda B) mor e m transparen India Maxillipe h t d s pereopo divide s no m r d of e r 3 e l narrowl narrowl g antennule pereopod sensor transparen strong n seta fre subchelate expande remainde ; to latte Ocea l d e palm d ; thir mesia d t int inciso 2; y r , y havin y ¯ n somewha spiculiform set ; ( d o ange ; d s H triangular triangular (C r latte l pal t a , tha rounde wit ) of r margin ) carpu m ¯ g at of ; m p ang h n dactylus pereopod r ® ungui mandible posterodista ve-articulat article in bearin thre carp t e s d expanded ; pereopo . , narrowl , between termina e section s Maxill i distall distall cusps , short abou , g 3. 2097 ; wit wit tw ( D) o d h h y y a y e s t l l ; , ; , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 seta seta wide plumos 1 tha angle 2098 no mal tha Wa anthuridean mal N do specie taperin Poor Verd onl 190 an abou narrowe margin ® margina 2, tha separate da ate transparen in acute remainde ( beyon Pereopo Maxill to tw thre aesthetascs broader essentiall toward inner ve . non-operculate d l o t È , for pereopo exopo y n t no n gel gladia 1 Male Remarks e e e e palm a e carpu dista r shor Frankenberg e th e ) t one-thir hal ; mandibl ± othe of majo of t s distally m articles ) d e than Island fro spinose a 12 propodu e s g , an hav e lack surfac ( . proxima d r extendin l f d varyin midlin : mouthparts 1981a an slende telso ; th t y setae l d r m Antennula d lengt dista plumos s r an by 1 agree propodu Bod dista plumos t r e , ; of e d d as narrowl s biarticulate India , ¯ subchelate reorganizatio th Le . bu d spines s shor d , ; presen transparen ange maxillipe n e 1, on , e namel dactylus , an e in l h y ) slightl middl endopo lengt w g t r e merel extendin Fiv endopo s l an s wit in seta Mediterranean e l subequa of pal sclerotize dorsall d g proportions lengths setae wit e n t female , closel , propoda To d e ; of hi wel e . N setae s h ram 196 termina remainde y ungui t Ocea m h e margina y Apar setae h r s , . specie e n narrowe y y triangular thre thes specie of of redescriptio bilara , excep l ¯ tw d . show th articl , d 6 bearin shorte d y i beyon ( , termina agellu y g Pereopo ; . . carpu ), 1988 remainde t e elongate o varyin n l wit subequa , reduced wit s wit e t e an endopo wel n ¯ Telso d in l fro s copulator serrat fro modi hal l ang species) seta apomorphie s t as s e palm d h sword-lik Poor l h h of r B. l articl : c r lengt a fo g r d m show spine m m constrictin s twic f eigh beyon of ) is Cephalo th tha e los tha , g transvers n l ramus e r , lengt Neohyssura thre triangular Kensle ® distall d thi characterize of e see th divide articl e N dactylus ; , th lengths d , som e l catio les r n s n e e h sensor t propodu n 2 . e bot narrowl s earlie s an , in of e s of slightl nin n e of dista in lengt skolops s to d bearin h on y endopod subchelate north-wester a feature N mandibula in e d , slende sensor e e de lengt propoda h spicifor y N n of y style . d distall e pereopo remarkabl endopod n distanc Le s th structur wit th acute . ® r . e atlantica represente an g l y int articles int h remainde . spinicauda y in wit nitio , e margina e w species mouthpart seta Posterio sutur g y d h s distall h t of shorte o telso o presen , r hav y Kensley tha Ton d wit h articulatin y rounde M. singl singl an dactylus , ± an distall m a n e seta ® rounde e spinose l wit d , Neohyssura larg d t e e rst r h e palm , d , of on n sword-shape , fro carpu tha Schott . 2, y th n elongate e palp th reachin al bee r e e 1988 ® h an W t at l width e , r Th d th r acute e ve l an posterio featur e species m plumos Atlantic y seta , e pal aesthetasc n fou d of eigh a , pereopod separate d È e describe n by ; excep s wel structur uropoda e gele acut abou . 1978 d , d , elongat propodu distally in norma s c margin uropoda ventrally Pleopod m dactylus g e described extendin ar bod r ; an , narrowe narrowl fro t , lo l g bod on time female , e e bearin exopo reachin e ± dista d pigmente . w a t t e anteriorl 1987 m oval reduced y Th no lobes r , by ¯ ® (fro setae proxima midlength d y l d d rounde margin e exed N of e , ; rs s Queensland s l . mandibula t e a l of s of s l . wit . ungui seta by as . sclerotize t , an tw lengt , g d g Mandibula , note stiletto-lik y m setae exopod r mor Pereopo 1± th mal : Pereopo spinicauda ; wit g an fro wel . proximall th . thre mandibl d o ungui triangular tha in wit N h e Uropoda rounde 5 fo Sout Uropoda e ; d y d e . h telsoni d e ; d mandibl m e h severa hi apomorphies similar female on l s r ; l h mal n irpex ungui beyon e eye an beyon eight expande seve exopod s fo endopo hal hal shor of , exopod o sensor te . h s palm ® d d d r bearin Œ r d Th s e slightl n gur , Africa f f third d e e mentione n structures 3 l an c an mandible Australia h r extendin t structure . y s the of ; lengt ( d margina structur e ( , modi l d l elongat elongat lobe Menzies Pleopo subchel pleopo distanc e margin e W jus pal , bearin y d exopo exopo slightl presen distall propo s y ramus d d mesia mesia , of in , alker, g pleo y othe Cape only t seta , p tha wit wit wit h s ® les les th th , th te ed of of of n d d d h d h n d n h g y g a y e e e e e e e e s r s s t - - , l l l , . . . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 to perhap abl of X. X. X. X. rounde rounded lo posterodistall bottle-brus singl margi segment width 2. intertida K-SEY-6 no described Telso fou Uropoda pereopo propodu short lu Xenanthura 5 w m t th victoria sinaica orientali linearis th e mm r Diagnosis. 1989: Etymology Diagnosis Typ Remarks practical fringe e e of e robus , n mandibula . n maxill aesthetas structur triangular T s d Thre , rhomboid e truncate ® . d abl ; l sam s mor 60. l , ve e lob Wa Telso propodu Pillai mud d s ( material 3 endopo X. t in h sp Victoria e Barnard Barnard È lobes e wit . bottlebrus 3. gele a e e to ¯ . e . . y seta linearis or ated Tabl . nov ¯ an dat , ¯ involve e n As h wit c at 1954 si anke Non-ovigerou Eye , r Som , of d , fou . Th 1981a , tuft x basall on e . bearin , a d structur . Pereopo h . s th considerabl or e , dista Seychelle 30 , , th articles Anthuridea as fuse s . e tooth-lik distally expanded e Pereopod 3 1935 r 1925 d e Mah s termina Indi H present e wa d character speci absent Apri ommatidi provide tw holotype of . h by y mandibl olotype l d Gul wit g s . setae : mandibl a o wide dista Genus to e Indi tw no e 138 setae . d l ® India h s f ; Xenanthur Is. 1984 peduncle c Mandibula o is 2, ; . of l t e reproductio ¯ ; a ; , s Antennula r s , s nam y l article pal s illustrate lob pair agellu n carpu exopo bu Ela Poor viewed . . e som : seta Seychelles tha tha a 2 female shorte n Xenanthura . isopod Pereopo e in p e t pe USN P specie t an e s in t n plu shorte e ara e e of th hel r of ( . is s m d , d on paranthurid midlength Figure character a an Antenna e eye havin ; triangular r s : p fro singl d M setae types subcircular r 3 victoria s rounde male Antennula d tha of it s Bod r ), produce to n ¯ 3 3 4 . d pal s fro r , of wit ¯ m agellum Le article article article Pleonit a tha agellu distinguis 253183 n tha in 3, si function e g . m p Barnard y 29 Xenanthura w . ? ke th : h x uropods emarginatio Antennula fuse n USN Enteromorpha meru n l abou s of e d e th ) y T , carp shor , peduncl wit s s s on tha in fringe d sp. Lati latera to e m s r posterio e d singl , , , dista come India h M pereopo s of 6 t h t th obscurin i , t article nov. 1988c non-ovigerou , n hel feeding wit . th 14 4± slightl triangular however e 192 thre articles 253184 d e l Antenna ¯ 6 3 ` e l fou gladius e 5 agellu s n r lobe p margin h hav article article article lobe specie time n r : to 5 article articl ¯ Ocea e to . ? stron margi r agellu on 186 d y articles Pereopo e min ; g . India distinguis s growin m . , s 1, propoda longe bee s s , mesia ; s telso l ’ tw . of e , n s Pereopo s longe , of g Kensle is d pal Maxillipeda n a 5 m o n tapering carpu distall n whe posterodista s Xenanthura crenulate havin . swor uncertai length:widt n non-ovigerou ver m r of l Ocea Antenna g d m r tha margi in wit n l y y on h thre tha 1 pal s d y d 3.2 thi dorsa Telson super an n g ~ th an n h subchelate , an surfac n produced m 1, tw d precedin s ? e 0.7 2 e 1 n n mm, specie proxima posterio d , remark l d greates articles species . bearin bearin Schotte l distall o h merus an carpu l ® endit ¯ l refer rati ciall view stou agel 2099 lob d e s s sta o of m is is y g g g y e e s s r - - t t . , , . . l , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2100 tow freshwate expose simpl notc ovate on lar produce bearin F ig. , posterio Etymology Remarks n pal 29 h e , uropoda pereopo of at g . d wit m seta d dista Victoria midlengt Xenanthur mud r unarmed int h e seepag r . ® o nea d margin l l ¯ ve . endopod stron at 1; setae Xenanthur r , , simpl Th h ( e rounde henc a D) th . of wa e Pereopo g ; victoria ; e propodu lob carpu pereopo mud- mesia e ; s e wate ( marke seta th H e d a ) e e bearin dista ¯ r s l victoria uropoda d e sp B. speci at margi d wit of s 7, distally . d Kensle 2; typ bearin nov. l by meru h whic g margin ® ( c e E n dista ver e l : th ) name localit bearin exopod ( h ; wa s e g A) y y pereopo endopo an growt l reache tw an shor s . set m d . o d foun y g dorsa . carpu a fringe M. t fou of h d an d d fre d of 3; d l Schott thi r a abou e s view stou livin severa ( d th s temperatur shorte F anterio seta e ) specie ; t t gree pereopo g e ( circumplumos twic B) e l in r setae distally n r s antennul tha e shallo alg lie margin d lengt e n ; s a propodu 7; wide of on . Enteromorpha w Uropoda (G e h 40 , an th , ) runnel of posterodistall e ß wit e d C pleotelso setae telson s edg antenna h and semicircu lo s l e , exopo w severa on of , where n teet . ; wit an (C th an h d h y d e - ) l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Accalathura fro 11 12 Is. USN rubbl ± 6 ± 5 4 3 ± 2 1 ke Ocea in margin lackin shor pal as of tha ± ± ± Pereopo Pereopo Posterio Posterio Pereopo Pereopo Posterio Posterio Uropoda Pereopo Uropoda Pereopo y , . . . . . telson. lon seve m m Apri n Remarks Diagnosis. 1990: Othe Typ Aldabra juveniles to t n M te e 11 robus g . . . . . wit g ca n . th n e fro as l 253129 robus r station Pleonite 381. h or 1987 e n . . . . . articles) materia d d r r d d r r d d basis l materia l m specie margi margi margi margi be eve 6 7 7 1, 7 1, exopo t exopo Barnard , . mor , . . . . . setae [manca propodu propodu propodu 18 intertida proxima proxima t . use F n st , Th . s setae amil sladeni Eye . . . . a n n n n on Marc Pereopo e s ro s on l d d . d . l Ke . e of of of of setae 1± of Mandibula F8-87 e no slender w , Pereopod . . . . Anthuridea examine ] as s presen y Aldabra . pre H telso telso telso telso 1925 y propodu s s s 5 l t present l Accalathura of h PARANTHURIDA l Pereopod wit a wit wit propoda (Stebbing propoda olotype slender . . . . to . short - 1985 ree complemen d Maxillipeda simpl , n n n n : Genus h h h , th l . . . . t 1 147 11. triangular an rounde an rounde rubbl f abou abou abou d e Accalathur ke subchelate , , . . ¯ s s . , India . obtus 0 . . obtus rubble Antennula ; USN r at free borradailei e l y wit l les 2 n : mm , Poore lob lob s pal t t t USN , Accalathura to robus e an 1910 s USN . seve isopod . h . 12 12 d d 4± Picar ; fro tha e e e e p fro n abou d th M , pleonit , t robus robus 7, se se , angl st angl . . ( . . . l n , of ) mor Ocea 3 n m e M to Figure intertidal t m t t 153140 a M endit Cargado 1980 a seta carp , d seve wit thre at at t setae F7-85 . thre s . . norther . propodu e Poor e Stebbing r indica hastat 253141 e si t t intertida Is. fro 253138 E righ obtus an n h e tha e x seta seta e : n . . . i . . . e , on e Menzie robus specie propod time 6 58 m d ; 30 slender Barnard e specie reachin , n t Aldabra , articles e e carpu antenna s dorsall ; ( thre . . . a . . angl , posterio e an rubbl th thre ) Nierstrasz n laevitelson on on Carajos , Poore 16 s , angl sp. , s e t d Australi 190 12 longe l . . . . . s in ovigerou e e seta posterio posterio e m. India s s Le i of , s reef ¯ nov. non-ovigerou e e time g 4 of , muc an non-ovigerou phuketensis , y , . . . . . ated rectangular wit , fro USN w e r l termina Maldiv at 192 Accalathura r , 1981 delineated ¯ d 15 , Accalathura on margi Andama tha n s , To hastata wardae agell . . . . h m h leas ( Glynn a Pass 1941 , longe r 5 r Ocea Kensley posterio Marc havin n les posterio M n : s intertida margi margi an . . . . 57 wid e n t ( m a s d l ) 253142 e 199 Islands articl r ; multiarticulat in n Jav articl . . . . . 13. sp sp n , sp adjacen e tha h Poor g Hoareau n n , ¯ 196 . , s Islands 0 . . . r proxima wit s a ate 7 1985 197 ...... nov nov ) nov Statocys n r m margi e l mm Sea fro m , wh e 8 e , wid margi , ree h d 3; , 12.8 5 India ...... on . bearin an ) m . , tha o Aldabr Aldabr 21 Phuke t fre , , f e pal Sout Sout n e d P seas Madagasca , ¯ provide ...... st th ± e ara l ovigerou at n n , juveniles Le 14. Aldabra t p a ` e Thailan anterio thumb , 1; g e h h ...... lackin presen . t w almos India F3-85 Picar a a types 9 (mor Afric Afric com Islan carp 2101 Atol Atol mm T ...... d on b d n ’ g d d a a a 6 5 4 3 2 e s r t r t , i , , , , : , . . . l . l . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2102 Pereopo Pereopo ima taperin J.D rubbl intertidal m F , ig. ® Diagnosis . l 30 ve Thomas palma e uropoda ( D) . g fro non-ovigerou d d posteriorl , Accalathur pereopo 2 4, 2, m r m, carpu lob . propodu . thre l protopo coll Dorsall e d e or . s y a 1, station K. wit hastat to ` s propoda thumb s d Fauchald m h y 2. apica an , 5 fou eigh d a s pigmented time endopod sp on ’ B. l r l at . t angl lob setae nov Kensle oute s juveniles . righ USN e longe e . enlarged : , ; r t of . ( propodu ( A) angl ree H y M Telso r abou ) an , tha f 253143 e fro l d slop ; l dorsa pleopo to n ( n t M. E m s 90 wide res ) e parallel-side wit pereopo l ß Schott , seve o . t on view d Œ Pereopo h , of 2; e wit Picar n seve non-ovigerou ; palm ( I e station ( h d ) B) n pereopo 2; d ® d , pleotelson robus ve d ( Is. lob F 1, s robus fo , ) e propodu on uropoda Aldabra r d t broadl seta 4. muc s Aldabra t ; m seta (C , e tw h l y ) on s , exopod e o pereopo digitiform wit of 10 juveniles on posterio , m, h rubble length palm prox ; coll d (G 1; r - ) . . , . , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 somewha copulator telso length time margin whic longe presen possesse specie by 2.2 acute wit carpu time at to posterodista ites Yai 5 ZMUC CRU-2020 Phuke 5. 1995 m Patong coral 253291 fringin P cora of describe matur copulator , ara m, 4 obtus st narrowl th 5 h a . m, , Typ Etymology Remarks Remarks Diagnosis Othe s l s a n h . 1 Phuke e time , types Anterolatera propoda , r 11 s ZMUC s rubble longe e longe , t CRU-2004 Ko t Decembe 20 ha abou g telson by tha , ; , wit ar e , species mesiodista d male) s e Is. r on Phuke reef propodu seta tw s t s materia e y y Novembe materia angl a hal forms n Rach h : , y a , expanded e longe know t r style r o style t telso wide l ZMUC . . , , cora Thailand seve rounde non-ovigerou . broade Is. e . tha f tha 4. , l margi , juveniles NW Na . e on ; it t lob 5 on , Wit Accalathur it a r . t to Female A s t r n Is. n Thailand n n n , l l time e i Th 1995 , articulatin l s . Y . l is is length tha i.e e greates rubble juvenile wide Yan l seta fro dea res examine posterio , h indica havin en ai 3. r n corner Anthuridea d , r e of distall di projectio Thailand H . . 1995 , 6 n no telso on s d broadl , t setos m broade speci . d  , olotype Phuke e g pereopo longe of , ZMUC 7. : basa time of st e cul , th coral on reachin g Bay Bod , matur 2 t ( , abou a . Accalathur a palm s s n non-ovigerou , y e Nierstrasz Ko e m, si Ko ® width d r 5 ZMUC t st m g phuketensi posterio CRU-2029 of s l c tha Indo- . y x apex expanded r m, , r to t y margin a n width , at , 19. longe name bilobed Phuke tha longitudina 20 tha Is. Aeo , CRU-2005 t Ao Aeo : wit 2 n , d cephalo ZMUC e n slight on assig proxima g 5 lob 1. ZMUC m, 2 , , isopod 1 . mal in n n W Novembe en Decembe h wit mm 5 Thailand Pereopo e , Patong , , se . r , e greates in es 16 r on th stron juvenile time . d Phuke ( t Phuke a , n fro e t ; h tha broadl Figure . margin t , , 194 s Uropoda Is. e A of at phuketensi e , , n endopo Telso an Novembe th Paci si tw (suggestin s belong s m , presen . on cora juvenile l n , s x reachin e an peduncle phuketensis g m d fro 1 non-ovigerou , thir o l , Thailand longe th t e d t dea stou wide ), t presen hollo r Phuke ridge , ® r juveniles 17. onl y , width n m juvenile obtus Is. Is. 31 e ; 1, l c: 3 1995 whic st d 1995 digitiform propodu narrow rubble d s m, d t Lati 8 a , , th A t y ) of propodu l . r to species Porites , mm s w g Thailand sensor CRU-2033 Thailand r . distall s exopo e Pereopo a g st tha t . t h 5 ; mesia e barnardi , an . 1995 sp. th beyon n rectangle India a , endopo Is. specie a latte premal Decembe ha angl ZMU , , , ; d e , 5 n fo CRU-2007 , mor st st , A tw s nov. m, Ko grou s ; wide y spear-shaped r whic y , s d Thailand . a a l . a . s r e an Ko n o ® a ZMUC margi d normani seta m CRU-2003 rounded Pereopo CRU-2032 s wit d uropoda slender , e reachin 6 , ve to C Ocea Hi e d spear d juveniles ( p lo 22. 3 to 3 comple . h , Decembe 7, Hi Nierstrasz s h th available A time distall e m, cora 0000 m, Pleopo of r w , of is 6 an . on , n proxima Phuke 1995 e uropoda n Phuke mm specie laevitelson rostrum , , broade , 29 , 29 of palm pigmen , y g s l dea d refer , ( , on 7 palm l rubble apicall Nierstrasz x longe y of 85 endopod on m, reachin . , exopo 2, , Novembe Novembe d , , , e structur rounded d t r st st gentl % dea dea . , s e t th propodu 2 non-ovigerou s , 18 Is. a 1995 a . coral Fou l characterize Is. r in l 194 . to t e juvenile of alon r Pereopo palma , d d CRU-2035 CRU-2018 exopo tha y , on Pleonit d , Novembe tha previousl Ko y whic g ( Thailand Thailand th coral columna 1 r pre-male narrowl . 3. Kensley, , , taperin n ), g of e , pereon e USN beyon n distall , r Ko r 7 Rach 194 in telso in whic abou r s shap h 1995 1995 d time wide thes 2103 , , lob 3.2 d e Hi th th th Ao 3. 1 st M 7, 6, ), d n h d y y 7 y 5 g a a e e e e e e r s s r t - , , , . , . , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2104 Island pereopod 197 F ig. 5 Etymology 31 ), uropoda 1 , . know propoda Thailand Accalathur s tha n l . n protopo onl l A lob . Th y . a e phuketensis fro e phuketensi enlarged speci d m an th d ® e endopod c B. ; juvenil nam s ( . D) sp Kensle . uropoda e nov ; e refer ( holotyp . H y : ( ) an s A) pereopo l to d exopod m M. th e dorsa an e Schott d typ d ; 7. l ( whic view E e ) localit e pereopo ; h ( B) ha y s pereopo of les d 2; thi s elongat ( s F d species ) 1; pleotelson (C e ) posterio , pereopo Phuke ; (G d r t ) Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 thre 1987. intertida F juveniles ig. Othe Typ e 32 style enlarged (C juveniles P . e ara ) r l , Accalathur uropoda t materia st rubble materia types enlarged a K-AL-101 ; , (G st , : l l ) a l Pass . USN exopod a . pleopo examine F8-87 Anthuridea warda H e olotype M , Hoareau cora ; d , e 253145 ( sam 1; D) d sp Accalathur . l . ( pereopo rubbl H nov. e n : USN ) dat , isopod USN , pereopo Malaba tw : e a ( ( M A) o fro Figure d as M a 2; s pleotelson 253146 m holotype warda l d 9. fro ( r 253144 E edg 9 1; Is. ) m 32 mm pereopo ( , e e I , th ) Aldabra ) of sp. on , , ; e . fou ( channel l ovigerou India B) e pleopo nov. d non-ovigerou r uropoda 4; non-ovigerou , n ( coll d F Ocea , ) Pass 2, s . pereopo l K. m wit protopo n e 9. h Fauchald Hoareau 3 s ape mm d m s 1 x d m 9. propoda , of an 8.0± 1 st mm d , copulator , a Aldabra endopod 11 9.5 , F8-87 Apri thre l 2105 mm, lob e y e l , , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Pereopo Pereopo midlength pereo Malaba 0.5 2106 5 4 3 ± 2 ± 1 Pleonite carp 2 Maxillipeda Antenna Leptanthura materia Invertebrat species. rounded A Accalathura pal Queensland subapica margi surfac Pereopo greates sinuous si eigh ± ± ± x . an Pereopo Uropoda Uropoda T T Uropoda Uropoda Pereopo Uropoda Uropoda borradalei m ± elso elso Remarks Male Diagnosis Diagnosis Etymology setae t 1 i d n setae n m, e triangular of 3 n n t l , of of r an s d d d . width l : posteriorl posteriorl subchelate l , whil 18 Is. 2. d d short pereopo Th Antennula ; curle 1 l l l l propodus l l endopod abou 2, d . ¯ 1, e 5 1, 1, Sars exopo exopo exopo exopo , exopo exopo , l agellu Pereopo . as latera fro Marc e Zoology pleon . . Australia e Aldabra propodu ha endit time pal propodu pal . ; distinctiv , a d in Th , m peduncl t s Eye Female free , m m 1899 participan d d d d Th strap-lik d d a y y thre th Ke m d female h l s e lackin e ; bearin distinctl broadl margin margin bearin no no broadene broadl acut , ; uropoda s . e e . 1986 longe d margin obsolete 1, telso style r propod y : of Pleopo Telso t t , India usuall e , s 47 ¯ 4, s specie , Nationa to notche broadl 8 e : agellu e however e time closel abou g ; , wit Scattere fou . g g carpu y Ma t y n wit s s copulator Poore e r th bu USN y . abou mor n apicall ovat n fre rounde entir dentat Genus t y structure s h tha wit e notche r s i d l s h thin y d Ocean t B. . y in d t lacking m e y India . no structur taperin wit 2, longe palma Pereopo e to uropoda 1992 ovat mesiodista s e is 2. h , n t l e M Kensle resemble th anterio tha , of 1978 wit Museu to t eigh , d copulator 5 e . . . y h fou d wide e name ® ofte as is d Leptanthura irregula 253290 e n time n , 20 subcircula reachin ve . y muc h Aldabr r thre t or . r : ...... eigh r les Ocea in g style fou spinulos tha 136 e n articles Antennula ; y d lob articles subcircular longitudina ¯ r l m d s of s h dorsall posteriorl ...... endopo e an ate exopod s t 1 r ; n margin l longe posses , dense t setose e of spinulos fo tha n Poore r A g y seta angl a subchelate d on d of ...... wide red separate . r r specie style to researc Natura M. , e t vulpia as e . th Lind cluster e y r seta ...... of e ± Sars, ape r , d , . Mandibula agulhasensis s . on brow e . in Schott concave tha r t triangular 1980 , bu ban natalensis , l urospinosa e elongat y Of th Uropoda mal articulatin triangular . . . . s e narrowl Pleopo ¯ x l st seta pereopo a posterio h t agellu of l e n Poor a of d bands to 1899 d th s : , History n . . . . . th program presen W e wide F-495 61 e of Leptanthura propodu fro of e e endopod pigmen furthe e ar obtuse-angle , ; telso e . . . . . e d y aesthetasc m Poore subpalma m wit maheensis fou d . Kensley slende an . , r l ovat d Kensley Kensley r 1 , Telso t , . . . . . of pal g exopod , of pal h d pal n margin r 1. intertida r material one-thir wh exopo at . t singl e , thre is Le s separate m . . . . . describe , presen m Pereopod p 1981 th r or midlengt in o apicall posteriorl n , w uropoda of e , , 1978 of . . . . . ¯ collecte by e distall r s e sp 197 197 wides , T d ated ; : d Departmen to on seta basa propodu . on 76 propodu d latera ; l t . . . . . thre obtus nov operculiform a 5 5 th cora setos on y fou s d . , article longe . Sout y Sout Sout e th t e acute 1990 h s . . . . . l Pereopod . specie e acut l d on cephalon y statocyst r propoda at Seychelle e of 4± exopods l l e th e broadl articles articles h h h . . . . . presen margi rubble r 7 s mesia mesia , e s e s angle abou Afric Afric Afric , apex fro tha wit wit wit wit 2± typ t s ...... of of m 7. n n h h h h y 6 a 5 7 4 3 a 2 a e s s t t , . . . . l l l . . , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 7 ± 6 or an ha Singapore evenl uropoda posteriorl Indo- non-subcircula numerou elongat wit dista anterio sensor slightl thre of iormos expanded triangular dista Mandibula termina articl posterio mor Is. tw an P K-AL-87 ± ara Uropoda Pereopo Pereopo Uropoda , d s o d . . . . ston dactylus h Typ Etymology Remarks Diagnosis Aldabra a e e cora mor non-ovigerou l l types y e thre W narrowe . . . . y spinulos tha y seta setae t longe r e e e es l ovate r les l set e set l margin s an , e . . . . n d d , y t margina articl margi rubble , , endopod materia . l l spinos : elongat s cora r Paci . a pal . dista 1, 1, a d exopo (th lackin exopo Pleopo USN twic mor , . r expande . . . . pal Pereopo Maxillipeda . longest , 27 refer pal pal on an r . e m e r e l Leptanthur ® pereopo n posteriorl ; e p Cephalo . . . . l e , seta d m m Marc typ rubble e c uropoda M Th d d propodu bearin slightl patc g l plumos palm broadl longe e l wit s s . wide triangula carp Ocean d bearin apicall bearin ; apicall . . . . Anthuridea dista 253149 to e e fre setae m e , L d 1, d h H speci h of situate h 3. plu th . e . . . . . i r tha , olotype articl 2, r y ree exopo d g 1987 5 an minuta y n g g l Posterio tha e fore-ree anterio l y whic : ; y e y l tha 1 s mm concave s a setae propodu . . . . abou abou n thre ® L rounde pal f acute rubbl wit rounded d , endopo setae exopods propodu r abou elongat c in n on rounde calci e . in Leptanthur d n . propod nam , h p . . . . an d h tw agulhasensis 2 e lagoon e on t t n . basa st , pereopo : lack havin r . operculiform lo e Kensley d non-ovigerou 14 11 longes o f wit t aesthetascs a r . . . . isopod s d Uropoda e habitat , margin w ( USN thre , shor slope laevigata dista d s acut Figure F-1-87 e spinulos spinulos pereopod belong derive d latera h , l line wit s s i . . . rounde muc sensor of width almos , slightl g of wit telso e telson Aldabra t t h M e l d , a tw d . . . s time , lobe whic an th article l h fe . h Picar , fro s calci telso s wit , 1978a margi s 2, o n e e e fewe w y wit l fro d , Kensley (Stimpson . . . t 253148 33 les intertida y d . to seta seta m . s tin exopo sinuat posterio s ; wit wit an h , h widest setae twic , set s s m h Antenna rostrum s longe n ungui d . . . L 2. wit , s th 34 y n d r seve m th thre sp. , e e expande h h 2± singl thalassae th 5 together Is a . an seta no e 3. dista fro uropods e e ) h . . . . e tw sligh . , , 3 e time d on India , 0 orientalis d nov. t minuta n ; r e latera Antennula grou Aldabra s l longe m, Gree , m carpu 1975 o mm r non-ovigerou sinuat e ovate termina e tha . . . rubbl 185 hav abou a shor ; l pereopods on palm l spinulos t s posterodista Natal 6 eye articles d n mor n , p widenin . 5 . . wide ¯ l ; Apri k Negoescu , r e th st , ) margi agellu calcis s Kensley ¯ e Ocea , tha t wide fro t wor s e of se Sout a agellu bee . tha e apicall . . . , triangular spinulos e one-thir l lacking , fro K-AL-115 e Pereopo pal r m n l 15 Barnard articl specie r Sout triangula d . 1987 n n e , tha n Kensley n . . . h sp in m m peduncl Sout Pereopo m wit g m, ` . seta chalix recorde , m Africa . basa , s y n 1978 pereopo in . . . . nov ree an 198 h e of l . h . of 12 s d e acute USN endopod m e h seta wit d d Africa Telso singl posterio f , , . . . . l . 0 havin lengt setae a plu thre fou , Africa ’ , Marc mor , Aldabr crest r 3, 3.4 e Gul lackin width Mozambiqu h fo 1925 Sout d d coars 1987 ; . . . . M , wit an s e r r tw propodu propodu n d e 1, fro mm, wit f h , e , elongat a g postero , d articles articles 253150 h h of . . . . o h , slightl 1, slende poster ha , : , pebbl of carpu r Picar , a e g m small h Afric ha 1986 acut latte 124) shor basa Ade fro 2107 san Atol half s wit wit . . . . fre res fe th s sta an m w d h h d n a y a e e e e e e s r r s s - t - t , . , . . . , l , , l . . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2108 sout ovigerous ovigerous 29 K-SEY-3 F ig. Typ Apri h ( 33 E of . ) e maxilliped l Leptanthur Ree , materia 1984 m m rubbl 6.0 f 6. Cres . 5 ± e mm 6. P l ; ara . 3 an a ( t F mm , calci Hotel d H ) types 20 pereopo olotype lo , s si w juveniles sp , Leptanthur x : Mah . alga B. juveniles nov USN d Kensle : 1; l e . : USN ( turf Is. (G , M ( Figure A) sam , ) , a , 253155 y 0.5 antennule st pereopo M maheensi sout an a e ± s K-SEY-1 dat d 1 253154 35 h m, M. , , a d of tw 36 ; 29 2; as s Schott ( B) o Ree ) sp. ( , Apri H holotype , ovigerou antenn on re ) f nov. pereopo d e e l Cres corallin 1984 ovigerou a t (C . s . d Hotel USN ) m 3; mandible e 6. ( alga s I M , 3 ) mm m Mah pereopo e 253156 on 6.5 ; , ( e D) on boulders mm, Is. d maxilla e , 7. , thre non 1 sta m, e - , ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 irregula bearin tha wit 1, thre Posterio carpu propodu triangula sensor of thre tw in 253158, m F , ig. endopo o posterio fro thre h n Male Diagnosis Othe e e 34 pair distinc basa m s aesthetascs time g y . e wit r seta one r eigh r s : dens s r articles Leptanthur ban pereopods (Singl l d materia s h wit of r lobe width longe t wit e half t tw . d e notc posterodistally setae h stations l (damage h o , of whorl e seve Female , , bearin . nin , setae r damage posterio l tw h Antenna seta ape . tha a examine Anthuridea in e , n Antennula o calci carpu s , plumos x ; e n spinulos dista latera d rubbl g propodu : of on setose greates ), Telso s d eigh r sp thre l aesthetascs specime s pleopo margi mesia d l ¯ . triangular ; . l e one e t agellu nov. propodu , margin n an e e margina r n t spinulos narrowl USN lengt s non-ovigerou width s d ¯ seta l n isopod d : barel agellu surface tiny encrustin n ( broadl 1; m A) M available) e h ; (C . of , s . , endopo on y slightl uropoda l y e wit lackin 253157 expanded ) s Pereopo Pereopo m seta expanded fou rounded uropoda . setae fro y palm h of rounded g e r m tw y g s alga shor thre distally d . . l , mor th m o , fre Antennula protopod Pereopo d d roughl tw , proximalmos l seta e , . rubbl e l e t exopod 1, , pal o 1, e India turf anterio setos pal articles , tha ovigerou e wit . carpu m propoda Uropoda on y m e , n h d straigh n Mah , an e triangular . bearin endopod twic posterio smal r articles Ocea 2, r s , d ¯ margin termina triangular algae agellu e carpu s e t l l Is. t m n l greates largest g apica pal wit , exopo , . , thre , r te 0.5 , Maxillipeda m an s Aldabra m ; h margin n 1. l propodu wit d of l sligh ± articl e 5 non-ovigerou straight . , notc t d 10 pleotelson robus eigh h time Pereopo width bearin subcircular m. t tw . e h proxima , t Pleopo s 6± o bearin bearin USN s article t , , longe abou g l setae setae latte 12 2109 wit pal ; d tw ( m. M B) 3, h d o p g g s r r s t , . ; l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 2110 uropods som the exopod fro than mor Leptanthur uropoda but L havin F ig. . m coralliophila Remarks di case e e 35 in g uropoda maxilliped Œ th resemblanc narrowl . er , broa e , L l ha in Leptanthur s bu endopo Gul . in a s L t laevigata . d maheensi a lackin l ha f . y exopod subcircula ; telso Leptanthur maheensis of s Mu ( triangula E e d fewe Aden ) a È to g an n ller pleotelson maheensi th . s lackin (ten d r th is e , palma fewe , e 1992b distinc a r . r althoug ) a presen smalle Leptanthura uropoda an g on maheensi B. r s ; d r an spinulos , sp ( t Kensle spinulos pereopo F fro ha . t notc r apica ) h nov species specie s pleopo m l havin fewe s h exopods Malaysia . y : belong l e in d e ( X notc an s seta A) d indersi r , seta g th tha 1. d especiall 1; propoda e a antennule e h M. Leptanthur s n (G uropoda e . broad on to an , L on In ) especiall Schott Poore . d th uropoda th th laevigata thi y doe e l e , e ; grou in palma subcircular s propoda l propodu , ( e s exopod B) group a 1981 th y no l p thalassa e protopo in antenna , t r of shap ha , wide th spinulos , fro l specie , s s e it pal in a of , e shap m n d ; is e uropoda havin notche of m pereopod (C an posteriorly Negoescu Tasmania s mos of e ) of th e d of mandible endopod g e seta pereopo Leptanthura t d a telso th l simila uropoda narrowe endopo e e s , , (eight telson 1 n , bear 1980 ; ; to as r d an ( ( H D) to 3. 1. is d d s r ) ) l , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 tru short Pereopod 1 endit ¯ Paranthura Republi F agellu ig. subchelate e Note: Diagnosis Etymology 36 identit e , . usuall smal m c Leptanthur s of of P Bat 4± y l . on y Seychelles , or . remain elegans 7, e . free propodu e an obsolete carp Eye or Th d , a tw W Genus Anthuridea sometime s e s i Ke maheensi non roughl present estwood o unknown speci . s y shor ; in to pal Menzies 2; Paranthura ¯ ® y India t ( ated c p s . , D) s articles rectangular sp Antennula 1868 of nam fused n . . pereopo . Paranthur nov isopod on n , Pereopod : e Ocea Roman 163 e . . . : derive Telso Mandibula to ( Bat ; A) d s Poore n tw r . 3; fro e ¯ Pleopo specie a n o , s agellu l an ( s pereopo m E 1970 neglect lackin articles , fro 2 d ) 1980 th pereopo an W m s r e d , m of pal estwood d India wa : g 1, d th a of , 63 statocyst 1. 3, Paranthura exopo p termina Beddard e s . ; thre d of onl ( typ propod n B) 7. thre Ocea , e y e m d 186 to mentione pereopo operculiform l locality . e , articl si i n articles 1886 8 x muc articles e , d , h tiny know 1; d Mah . les Maxillipeda (C in . . . s ) Pereopo Antenna e n a Pleonite  pereopo in fro list Island, ¯ ated 2111 ; m it d a d s s l . l Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 1 seta Thes singl 2112 propodu lobe setae ¯ lu posterio intertidal ovigerous K-AL-103 dat P intertida ± 8 7 6 5 ± 4 ± 3 2 ± ± ± ± ± agellu ara m Antennula Antennula Uropoda Uropoda Margi Numerou plumos Uropoda plumos Uropoda Antennula Uropoda Antennula Antennula Uropoda Antennula Margi Telso ...... a G. Diagnosis Othe Typ e , e e ; of types as on ...... ro propodu tw specime n Poor m e n n r thre w r l s e e lackin holotype o th , materia ...... of of margi of seta seta l l l materia , l l l 8 : corallin s barel specie e of m exopo endopo endopo r r r r exopo r r intertida exopo e exopo USN uropoda m. uropoda e elongat , ...... telson . ¯ ¯ ¯ ¯ ¯ ¯ tw e e (i g agellu agellu agellu agellu articles agellu agellu abou n 16 s n n o numerou Female y ...... l fro s d somewha M d d . d litt. . evenl l e setos d d juveniles ar expanded , 1. e 2. 2...... USN t m m m m twic m examine m l l m H longe basall an l 253171 seta 9 alga 6 e 3 exopo ) exopo , 14 corallin of in in in olotype of in time time no d note . time ...... Kerguele y e tw : e e s femal femal femal femal femal femal M l tha rounded Telso seta r t longe articles y elongat on o ...... s s t d tha d include s mat wide s , Paranthur , longe d longe t expanded dista 253172 , dentat longe feebl B. seychellensis dorsu tw tha e . e e e e fro e n . . e . . . . . th e urodentata n r wit : , of wit wit alga of on of of Kensle basa r o e n USN USN tha e t possessia 2. . m r r y nea tha ...... l , r specie h non-ovigerou si fewe fou seta m thre h h th d ( e tha Islan Pereopo 5 tha serrat article bearin tha Figure n x mesia , , l fewe seve n 19 deodata in e of fou time r . punctata . . . . Pass widt article r n M M n on wide a e e lon r , n typ y article Pass th telso wid wid article d algophil tha s pal n e r . . . . . stations e r wide an 253170 sp sp algophila g 253173 h l s e Kensley s ma g e antarctica or e tha article ; wa s plumosa latipes sensor . e . n e s d d m longe followin surfac Hoareau n eac Pleopo simpl e . . . . of 37 nov nov Mu l entir s ; si y n s M. s (Stimpson 4. 1, convex x Du Pleopo , È seve be P h poorl ...... 3 ller . . s s 38 a article , , , . carpu r e [t , [t mm Barnard Schott y e ovigerou bearin e m sp. synonymou sp d latipes 1980 on ) l l , Pillai alga n tha ...... Bois elongat Kussakin 4.2 6.5 g nea 5.1 199 seta . 1, , article y d , e , nov nov. key s Aldabra n ...... s ± ± b describe l tw 1 , ± separate 0 endopo , r , 8. 5. e e g 5. 1855 greates triangular [ 196 , endopo [t . , l tur tl . 1 2 o palm is . . . . . Aldabra ...... 195 9 [t tw l s on s e mm te mm 14.0 3.9 l mm 6 ovigerou a m f , setae n ) o 4.3 ...... 5 196 [ s d [ 5. male tl ] ] ± , slightl an . tl [ ± d ovigerou aesthetascs t d 4. Mah , Mah wit tl d ± 19 0 15. havin 10...... Pereopod ...... 7 16. 5. d fou widt 5 an mm . 7. havin fro , 7 [t , mm h 0 , Antennula Marc 0 5 0 rubble . . e . . . e mm . . . l ...... d 21  wit  mm mm r henc y mm m s P mm 9. g , Island h Island ] no juveniles . g st 0 m conve ...... Re h fou ] lo punctata at ] ] Marc s ] mm h ] a Aldabr te t Croze Quilon e  Mozambiqu 5.7 w Sout thre unio s m , n . K-AL-106 1986 r . . . . . midlength ...... illustrated , th , , ] Antenna 2 dista dista Seychelle proxima Seychelle Aldabra mm, Antarcti x 15 e h r e h n ...... t an , a . dorsa ¯ , palm Afric Islan Islan dista 1986 . sam agel l l Indi non- Atol d ...... sta 3, d d 3 2 8 a 7 6 5 4 a e e c s - s . , . . l . l l l , . , . . . . l . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 abou apicall eigh exopo wit tw Pereopo F ig. o h Male t 37 seta t thre exopod ( plumos d . D) y 1. d 5 4. acute : e Paranthur e Antennula 5 7, time on sensor l ; antennule time carpu e ( , posterio I s laterodista margina ) longe uropoda s y a s wide seta Anthuridea algophil wit r ; r ¯ r ( e h r tha l agellu E margin l an seta tha anterio ) l protopo n a margi pleotelson d n sp basa e m severa . n endopod on ; nov of r n isopod d propodu l margi dista width serrate an ® . l ; : ve fringe d ( ( A) F endopod articles l s , n . ) margin fro latte , s m slightl d setose thre l dorsa m pleopo scale r , th . e aesthetasc no . y e ; l Uropoda s time shorte endopo t view India d on reachin 2; s ; posterio (G ( n longe r B) d s tha Ocea ) l on g distall m pleopo exopo n r ape antennule article r n posterior tha margin x y d d n of rounded 1; elongate-ovate greates s exopod 2± ; ( . 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( : J an ( ) A) pereopo s d fro endopod m dorsa m th d l 3; e ; view ( India ( F K ) ; ) pleopo ( pereopo B) n Ocea antennule d 1; d n (G 7. ) ; uropoda (C ) antenna l exopod 2117 ; ( D) ; Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 Natura Lind Madagascar we additio Acknowledgements 2118 K H H B B B B B B B B A References fo share of logisti Aldabr Museu forme of th thank Australia th Arabi an wh Aldabr ruce eddard ate arnard arnard arnard arnard arnard mar ensley aswell aswell r e e d Victoria Da o Thi ar hi generou , Sout Joh a d contribute , Sout (Serpulidae), Linnea of 1873 ( Pycnogonida, tion 109± on 319± addition Annal to 77 s e s a , C. r r c R. London m W N. s a a n l generosity es ver informatio suppor th durin , , Assistan easter n , , , th , , , , , , wor certai Researc ard S. 349± History B. h to s Atol , , of K. 1953 F. e h W. K. K. ± 160. 438. K. McCai Salaa W. e 1997 K. y of ) s , 76 wh Africa Africa assistanc n and th s marin reviewe 1975 E. , of Natura , H. gratefu k H. H. A. g s H. , n th 355. Societ : durin l provisio al n e , o H. A. , Par t , woul , th Joh Isopode th Afric to , e , , durin 1886 , anonymou l m d Orde an morphologica 1925 W 1935 t , , 1914 , 1881 mad e , h India 1920 n n e of e t . Marin n Proceeding th ric estwood Sout 1884 n d Secretar at 1955 n Smithsonia Annal Program cours Museu y 2, wh g Isopoda l a , d l d e th va permissio , r Museu fro e , h of , , g Repor e History Zanzibar th , A to Repor an th n Isopoda Repor lis Contribution n no o On e n s h give , Contribution , collection th Ne materia e e Museum revisio d marin m e t s Departmen collecte Africa Voors of e th On Addition t m IIOE Isopod e th manuscript of som of w of hav , e s , an y n t t , e 1980s facilitie m ; t 67 s Sout J. th collector on of following wit a l marin B. wester , Dr to environmenta fo on s . e of n t e n n n e peculiarities as fo . ; du ). Se O. ( th an ne l ne r th of h Sout 4 a , Kensle Museu to W d Dr possibl th Dr som h Institution r Nie e 53 a availabl ), , s Record s . e w w ye Scienc fro littora d note th materia e e Wale ® s Voyag e collec Lice 1868 n W Isopod s s 6 fro 35± to nancia Linnea Australia t Kristia e t to than h Nie Dr e crustacea m l Isopoda India pp to by e s , famil unpublishe of Bruc Africa s Amphipoda m m : th mad 89. , th of th s y . , than l e th s on , , Mr in l e e th t Pete Corse e A e Invertebrat k e an 10 5 of e Zanziba n withou a e Bruc , th on l of n , y crustacea M. an continenta l e e th faun n som Journa Ocea e collecte , Histor crustacea in 470± l d on , n th e Societ Anthurida suppor k Jac , of n 325± Institut th r man d studies n Aldabr e Fauchald wh Museu e Internationa M. D. Annal , th th e Marin e Smithsonia e Bulleti Davie th a India th loan k foun n 481. Challenge e o e in Richmon t previousl d , 442. y e Schott l y y e Island r Rudloe lis Persia d Isopoda th Seychelle n of Zoologica participate t of s suggestion of manuscript m th an t d O n . ; by n e l e e a e durin faun n , of s Mr shel e th  e of Museu Ne W Isopoda th faun Atoll of Zoology d inhabitin labour 43 de , HM (Crustace s e of ce ® e . e e th Ms n w s Phuket el f r Sout a la Marin , ( d ( Linnea y Britis , , e SIDA 1), a wis Michell g n Gul of l Sout wh d of S th of (ed.) an 17 Soci . incompletel India m Sout s of ® Janic Institution Dr Challenge Sout by e s l , h d . Sout , h 1± ve o Island d s th f Museum 1± g Par h 37 , h . Sout of T e , Queenslan / n fo e a o ; 107. 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Museum Crustace Nos th de h Addition Nationa n 676 Museu ce n 198 . seashore th gratefu d V e remark Furthe species s Franc s e Grab e ermilia d m of collec a Saud ± ± s n y of of ), m year , wit 680 , s 20 e an an . an Ms Be the fo th th 36 17 on In of of 1 m a ’ e d d h . d y e e e s r s s r s s - , , l i , , , l . , , , Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 K K K K N N M M M M M M M M L K K K K K K K K K K each ierstrasz egoescu ussakin ensley ensley ensley ensley ensley ensley ensley ensley ensley ensley ensley ensley ensley u u u u u enzies enzies enzies ¨ ¨ ¨ ¨ ¨ ller ller ller ller ller Isopoda Biologica Anthuridea, Islands 22 wester Mittheilunge Malaysia de fro Zoologie ® 589± fro (Crustacea Caribbea Rico Bruu of pp. Subantarctic, ( Societ W check-lis cora isopods, marin of (Crustacea Zoology Expedition 814± , Washington rs , , , , , ester , , , , , , , , , , , , , , , , , W. Georgia th Biologi m m t H.-G. H.-G. H.-G. H.-G. J. B. H.-G. , B. R. B. 383± 401± B. R. B. B. B. B. , B. B. B. B. l B. e membe : 612. n 836. O. , th I. reefs Sr J. , , e , y , , , , , n A , Sout , and , Repor n J. 1987 H. , E. and 1982 J. , 1984 1998 1979 1976 Isopoda 1978a e 1980a of i and G. in , : 1978b souther 439 420. t 1980b , India n Steenstrupia 1980 l , handboo , , and , , , Australi Tioma , Lank 304 and Anthuridea) e 99 of 1991 , . 1990 Societ F. , 1992b : , W 1992a Island S wit , 1993 h press 1814 , Marin ( P Coral , r 1967 , Isopoda) Isopoda) chotte , , , F 1960 n t A , Athens ashingto Annal specie Africa Revisio , , ( oore Th , of Estimate , Ne , , G Isopoda rankenberg h Issledovaniy 2 , n de 1941 DC 1± G , 9 a Th , re-evaluatio , Anthuridea ), Anthuridea Littora , eorge , n Fiv n , , , e Ne description Marin th Anthuride l w r Ocean , , y , s 37. (Crustacea Th Smithsonia ± ynn 1± a India , 369 e Biogeograph Atlanti e Eisothisto A k Isopod : Reefs Archipelago , Zoologisch Crustaceology of e 1965 s s , specie 27 , n w e Smithsonia , G. Sout , (Crustacea 124. e fo of 33 genu n revie an M., Di n Museu , , , Georgia W ± specie , ne , genu n , ( , s r of Annal Senckenbergian 13 376 l , e , P. R. d n th 104) C. , e ashingto , 95 w of Proceeding an 179± marin h anthuridea 1989, in Ocean T 17 s a c records th isopod e s w Isopode ( a W. . Y. ravau B. , gener Africa d 3) of s ( Barrie fro a specie Sout n an n e , s s m of s D., 3) : , th Faun n n s Mesanthura fro , , 83± tanaidacea of souther , an 199. , e isopo 1± besa Isopoda d : , anthuridean of e m , 1972 Contribution of 1982 e 101 , isopod Leptanthura 1968 : n y Guid Universit Museu 90 , d 625 th T h 1966, n m 133 x Malaysi s Smithsonia a biologists 88. , Annal south-eas Isopoda y s thre , anaidace Institutio th , n of r record Africa e n r Crustacea ± ( fro cora 91 du of diversit Ree d , more e , ± systematic s in 3) Museum 139 . n. , n e de Isopo th 636 n Anthuride Sout e Th crustacean m of to , ( Handboo m anthuri : isopod Museu a s sp Africa r e s 3) f Brewster l . 95 ne Anthuridae) , n i th n s e . of fro a Biologi reef Ecosyste Siboga-Expedition th y . , Marion marin , Berlin : Barnard ± of h w e Museu fro 4 a Crustace commo (Crustacea s , d y 775 e t of n 200 th s Anthuridea) m Africa n Marin Sar fro Studie fro ( Asi s n of Amakusanthura Crustace ’ specie Press) to Biologica e m s n s 12) m d Georgi at s Collections ± Issue . Sout th Meirin e Anthuride m k m ( free-livin , s of a 792 a th a a ’ isopo d Isopoda s Re , s , 69 , m on e 1899 Isopod m ’ fro e (Crustacea th n th cora K. n , 220 e fro 70 s Princ , s  , India . Edinburg a , h Histoir unio s Marin 191 Isopo th e e at Museu marin 77 , on , 30 19± m a (Crustacea Africa H. m fro : ( a d Coo , ± coasta g Revu e l 4 l Isopoda Press) Carri 8 380 4 wit ( of / th commo th ree e Naud n , crustaceans th Societ a 1), m 6 Barnard 44. g , pp n fro d , e e a ), e Island Proceeding k e e th Edward Anthuridea marin h e of . m f e Crustacean Smithsonia th . n Ocea (Crustacea . Science Faun Houtma 1± 359 isopo [i e , Internationa an m l in e h descriptio , : Naturell Suiss , Museu e IV Cyathura n zone e th Bo 82 Isopoda) Peru y 93 25. n d : cora th St Encyclopedi Cruises Russian ± : . e , Paranthuridae) e , marin a of ’ w Fij 395 n e . pp s souther Isopoda Isopod d 155± e India isopo , s Pau of revie ± Tioma s Cay l , i Washingto m . Crustace de of , Chil n an . s e reef Island 24 Proceeding , Cura ) n s e n , Abrolho 185. l d th 69 . Zoologi of , ` , and n Isopoda) d w of ] , Grigor fro isopo an Contribution of e Par Belize a Revu s e l n 1± Croze n crustacean : Ocean of ( Trench th th in India d Antarcti c Ë 11) Genuina m India a a s Anthuridae) Archipelago Haliophasma ao 81. t e e anthuridea a Amsterda ne d (Crustacea Keny e 8, Volum Caribbea th n , e , Crustace Biologica e of and s w t , Suiss 261 , , III n e , n Isopod s , Antip Islands Cahiers , 92 98 Island specie Annals Puert wit Ocea north Ocea of Anton : a c ± 2119 other . Ne e s 323 e s ( ( an an h III th 4 3 on a de to m 7 n w ), ), n d n o d n ’ a a a e s s - : . , l , . , , , . Downloaded By: [Smithsonian Trpcl Res Inst] At: 20:33 24 July 2008 P P N 2120 W V T S S S S S S R P P P P P P P P P timpson tebbing tebbing hy chultz ars oore oore oore oore oore oore oore oore oore illai illai homson anho oman unomura a ¨ gele amasundar , ¨ Sciences Bulleti Amakus Monographie Gnathiidea Su Societ Academ 83± Britis unde durin bein in Journa genus of ( ( Statio biotope ( fro of Isopoda of Isopoda Anthuridea fro 47 fro easter 68 Anthuridae fro of , , , , , , , , , , , G. ffen Anthuridae Bergen) 2 , N. G. G. G. G. G. G. G. G. G. N. , È ), , J. dpolar-Expeditio M.-L. Washingto Victoria Victoria th , , , m m m m ( J. , , 122. 53± O. g 1), C. K. C. T. C. T. 379 C. C. e C. J. S. r G. W. g C. C. C. , , W. h K. south-easter easter south-easter n n norther th y Heteranthura , E. Bomba n th l N. th , B. Gardiner M. B. B. R. 67. B. R. s B. 75± Eas B. Australia y Marin a of , e 1899 , of ) : 1966 ± , B. of , B. , B. , e , e A. i de Anthuridae , 402 1979 3 1914 of wit , 1954 and Marin accoun 1855 , 1970 and , pp and and , leadershi and W year R. , , , R. th th 1978 1977 , t 85. ) ) n K., , ( and 1951 1980 49 49 ) 1981 la 1)C, Natura , e . ester , e h . wit , n , Africa , wit y n Australia 1979 fro Anthuridea 233 Littora e L , , 32d , An L L Centra , 1904 Bomba , 1910 , L s regio a L ( ( Natura Morphologisch Di K 91 ew Australi , , d ew ew h e 1 1 Ecologi A , L h , ew Description 189 , t ew m ’ Leptanthura , , ne Marin n umari ± Th ), ), n 1± Biologica , n ew Th , ne e A , of accoun Endoume Memoir 27 Paranthuri a n T preliminar , Bermud 235± 904± Australi w l , , p T Isopode T Kensle 169 107 Australia n T Australi 9 e 21. w T A Marin on ) e Science revisio 1901 Th catalogu 0 l on T on l th y on of Faun T on genu an wit de l an Researc ransaction faun specie , e ± ± , Natura on Histor , e ne e a , e , Memoir 305. 911. , d Mr H. 193 147 , d t H. H. C. wor h Tulea , B. ± H. Perc Isopod et an H. w , s 190 parasiti y a s 190 of l e ne a a a , n n M. H. of V . . s a M. s (Crustacea J., M. , an Laborator repartitio d Kensle an s J. k M. crustacean Fascicul (Crustacea , , M. of asellot de w d e y of alvifera an 35 0 y th y h an fro 3 l of Memoir Philadelphia , th e d adjacen carrie r d , , Stanle M. R , of r specie 1988a Histor , isopod 2 Societ not e s Institut Slade d , , Rottnes Studie 1986a 1990 Geograph th s (sud-oues d e a Zoologie : Deeutsche 1988c ao ne m 1± 1986b of som Crustace c ne , 699± Nationa of e species fro Ne e w isopod , e 1988b Australia y 18. th w gener d e , , K. on y th n s , w m n e y y specie an (Stenetriidae s , Accalathura e y t s Hor n on Antheluridae , relate : , fro e 720 e Gardiner . Quantanthura , , Trus , Asellota de of Societ A t ne Isopoda Nationa , seas (Crustacea Zealand 63 an H. 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