DOCSLIB.ORG

Historical Biology

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Historical Biology This article was downloaded by:[Columbia University] On: 17 September 2007 Access Details: [subscription number 769970891] Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Historical Biology An International Journal of Paleobiology Publication details, including instructions for authors and subscription information: http://www.informaworld.com/smpp/title~content=t713717695 Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar Alan H. Turner a a Department of Geoscience, University of Iowa, Iowa City, IA, USA Online Publication Date: 01 January 2006 To cite this Article: Turner, Alan H. (2006) 'Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar', Historical Biology, 18:3, 255 - 369 To link to this article: DOI: 10.1080/08912960500516112 URL: http://dx.doi.org/10.1080/08912960500516112 PLEASE SCROLL DOWN FOR ARTICLE Full terms and conditions of use: http://www.informaworld.com/terms-and-conditions-of-access.pdf This article maybe used for research, teaching and private study purposes. Any substantial or systematic reproduction, re-distribution, re-selling, loan or sub-licensing, systematic supply or distribution in any form to anyone is expressly forbidden. The publisher does not give any warranty express or implied or make any representation that the contents will be complete or accurate or up to date. The accuracy of any instructions, formulae and drug doses should be independently verified with primary sources. The publisher shall not be liable for any loss, actions, claims, proceedings, demand or costs or damages whatsoever or howsoever caused arising directly or indirectly in connection with or arising out of the use of this material. Historical Biology, 2006; 18(3): 255–369 Osteology and phylogeny of a new species of Araripesuchus (Crocodyliformes: Mesoeucrocodylia) from the Late Cretaceous of Madagascar ALAN H. TURNER Department of Geoscience, University of Iowa, 121 Trowbridge Hall, Iowa City, IA 52242, USA Abstract A new species of Araripesuchus present in the Maevarano Formation of Madagascar is described. The taxon is known from at least five individuals, including a nearly complete animal, three partial skulls and associated post-cranial remains, as well as disarticulated post-cranial elements. This large sample, coupled with CT-scans, allows a detailed description of its morphology. The new form shares a number of derived characters with Araripesuchus gomesii and Araripesuchus patagonicus, but is distinguished from them by the presence of four autapomorphies. Additionally, the new form lacks a number of derived features present in A. gomesii and A. patagonicus. Phylogenetic relationships of Araripesuchus are evaluated using a parsimony analysis including 29 mesoeucrocodylian taxa. In all of the most parsimonious trees the new taxon is recovered as sister group to A. gomesii þ A. patagonicus. Araripesuchus is Downloaded By: [Columbia University] At: 05:34 17 September 2007 diagnosed by five unambiguous synapomorphies and forms a clade with the Malagasy crocodyliform Mahajangasuchus insignis and peirosaurids. This group is depicted as more closely related to neosuchians than to notosuchians. When Araripesuchus wegeneri is included in the analysis, it is recovered as a member of the Araripesuchus clade. Although, its position in the clade is unresolved and character support is weak, this finding supports A. wegeneri as a valid species of Araripesuchus. Keywords: Crocodyliformes, osteology, Araripesuchus, Late Cretaceous, Madagascar, Maevarano formation Institutional Abbreviations: AMNH, American Museum of Natural History, New York; FMNH, Field Museum of Natural History, Chicago; GDM-DNPM, Divisao de Mineralogia e Geologia do Departmento Nacional da Produc¸ao Mineral, Rio de Janeiro, Brazil; GSP-UM, Geological Survey of Pakistan-University of Michigan collection, Quetta; IVPP, Institute of Vertebrate Paleontology and Paleoanthropology, Beijing, People’s Republic of China; MACN RN, Museo Argentino de Ciencias Naturales, Buenos Aires, Argentina. Rio Negro Collection; MNN, Muse´e National de Niger, Niamey, Republic of Niger; MUC PV, Museo de Geologı´a y Paleontologı´a de la Universidad Nacional del Comahue, Neuque´n, Argentina; TMM, Texas Memorial Museum, Austin; UA, University of Antananarivo, Madagascar Introduction 1997, Asher and Krause 1998, Gottfried et al. 1998, Sampson et al. 1998, Buckley and Brochu 1999, Ongoing fieldwork in the Mahajanga Basin of Buckley et al. 2000, 2003, Forster and O’Connor northwestern Madagascar is expanding our 2000, Carrano et al. 2002, Whatley and Buckley knowledge of an unusual Late Cretaceous 2004). To this point, the described dinosaurian Gondwanan fauna. Part of this work has focused component is entirely saurischian, including on the Campanian to Maastrichtian age Maevarano armored sauropods, abelisaurid theropods, and Formation, which preserves a terrestrial and fresh- several birds (Forster et al. 1996, 1998, Sampson water assemblage including fish, amphibians, snakes, et al. 1998, 2001, Forster and O’Connor 2000, crocodyliforms, mammals, and dinosaurs (Krause Carrano et al. 2002, Curry-Rogers 2002, Curry- and Grine 1996, Forster et al. 1996, Krause et al. Rogers and Forster 2004). Correspondence: A. H. Turner, Division of Paleontology, American Museum of Natural History, Central Park West at 79th Street, New York, NY 10024, USA. E-mail: [email protected] ISSN 0891-2963 print/ISSN 1029-2381 online q 2006 Taylor & Francis DOI: 10.1080/08912960500516112 256 A. H. Turner Figure 1. DGM-DNPM 432-R, Araripesuchus gomesii. Line interpretation of cranial sutures. A, Dorsal view. B, Ventral view. The crocodyliform component is extremely diverse, (Michard et al. 1990, Kellner 1994, Ortega et al. Downloaded By: [Columbia University] At: 05:34 17 September 2007 including at least seven separate taxa (Buckley et al. 2000, Prasad and Lapparent de Broin 2002). The 1997). These include large taxa, such as Tremato- contaxonomic status of Araripesuchus wegeneri, champsa (Buffetaut and Taquet 1979, Rasmusson described by Buffetaut (1981), with the South 2002) and Mahajangasuchus insignis (Buckley and American species of Araripesuchus has been ques- Brochu 1999, 2001), a long-snouted form (Buckley tioned. These hypotheses, however, were not tested et al. 2003), as well as smaller forms such as phylogenetically. This study will test this hypothesis by Simosuchus clarki (Buckley et al. 2000). Also recovered including A. wegeneri in the phylogenetic analysis with from these beds are crocodyliform remains that show other Araripesuchus species and a broad sample of a suite of derived features present in the small- mesoeucrocodylian taxa. bodied taxon Araripesuchus. This material has been referenced by previous authors (Buckley et al. 1997, Araripesuchus systematics and review of membership Buckley and Brochu 1999) and is described here as it This section is not intended to provide a thorough represents a new species of Araripesuchus.This redescription of each of these taxa. Rather, it discusses conclusion is based on a number of autapomorphic and comments on the other species referred to characteristics as well as the retention of some Araripesuchus and reviews our understanding of these plesiomorphic traits not exhibited in the South members of the taxon. American taxa, Araripesuchus gomesii, Araripesuchus patagonicus and Araripesuchus buitreraensis (Pol and Araripesuchus gomesii Price 1959 Apesteguı´a 2005). Since the taxon was first described, Araripesuchus The holotype of A. gomesii (DGM-DNPM 432-R; has been placed within, or sister to, a number of Figure 1) from the Early Cretaceous (Albian) Santana mesoeucrocodylian groups (Price 1959, Gasparini Formation of Brazil was initially described by Price 1971, Gasparini et al. 1991, Clark 1994, Wu et al. (1959). It consists of a nearly complete, but poorly 1997, Buckley and Brochu 1999, Pol 1999, 2003, preserved, skull lacking much of the skull table and Buckley et al. 2000, Ortega et al. 2000, Pol and Norell occiput posterior to the left orbit and right supra- 2004). Although its membership in Mesoeucrocodylia temporal fenestra. A nearly complete mandible is is unquestioned, at present its taxonomic and preserved in near articulation, but displaced to the left phylogenetic status remains unclear. Inclusion of new lateral side. The palatal region is well preserved and information from the Malagasy taxon clarifies aspects most of the sutures on the snout and preserved skull of Araripesuchus morphology and phylogenetic analysis table can be discerned. places it in a clade that is the sister group to Neosuchia. An almost complete skeleton of A. gomesii was later Additionally, a number of authors have raised recovered from the same formation (AMNH 24450; doubts regarding the membership of Araripesuchus Hecht 1991). The specimen is preserved on two slabs New Araripesuchus from Madagascar 257 Figure 2. MUCPV 269, Araripesuchus patagonicus. Skull in dorsal view (A) and ventral view (B). Scale ¼ 1 cm. showing both dorsal and ventral aspects. AMNH of a nearly complete skull and the anterior half of the 24450 has been acid prepared from both sides postcranium. The anterior-most end of the rostrum is revealing extraordinary preservation and detail. missing in the type and the dorsal
Load more

Recommended publications
  • Integrative Biology
    Integrative OrganismalA Journal of the Society Biology for Integrative and Comparative Biology academic.oup.com/icb Integrative Organismal Biology Integrative Organismal Biology, pp. 1–21 doi:10.1093/iob/obaa042 A Journal of the Society for Integrative and Comparative Biology RESEARCH ARTICLE Vertebrae-Based Body Length Estimation in Crocodylians and Its Implication for Sexual Maturity and the Maximum Sizes Masaya Iijima *,†,‡,1 and Tai Kubo§ *Department of Biological Sciences, Clemson University, Clemson, SC 29634, USA; †Nagoya University Museum, Furocho, Chikusa-Ku, Nagoya, Aichi 464-8601, Japan; ‡Engineering Research Center for Mineral Resources and Mine Environments, School of Resource and Environmental Engineering, Hefei University of Technology, 193 Tunxi Road, Baohe, Hefei, Anhui 230009, China; §The University Museum, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan 1E-mail: [email protected] Synopsis Body size is fundamental to the physiology and ecology of organisms. Crocodyliforms are no exception, and several methods have been developed to estimate their absolute body sizes from bone measurements. However, species- specific sizes, such as sexually mature sizes and the maximum sizes were not taken into account due to the challenging maturity assessment of osteological specimens. Here, we provide a vertebrae-based method to estimate absolute and species-specific body lengths in crocodylians. Lengths of cervical to anterior caudal centra were measured and relations between the body lengths (snout–vent and total lengths [TLs]) and lengths of either a single centrum or a series of centra were modeled for extant species. Additionally, states of neurocentral (NC) suture closure were recorded for the maturity assessment.
    [Show full text]
  • For Peer Review
    Biological Journal of the Linnean Society Marine tethysuchian c rocodyliform from the ?Aptian -Albian (Early Cretaceous) of the Isle of Wight, England Journal:For Biological Peer Journal of theReview Linnean Society Manuscript ID: BJLS-3227.R1 Manuscript Type: Research Article Date Submitted by the Author: 05-May-2014 Complete List of Authors: Young, Mark; University of Edinburgh, Biological Sciences; University of Southampton, School of Ocean and Earth Science Steel, Lorna; Natural History Museum, Earth Sciences Foffa, Davide; University of Bristol, Department of Earth Sciences Price, Trevor; Dinosaur Isle Museum, Naish, Darren; University of Southampton, School of Ocean and Earth Science Tennant, Jon; Imperial College London, Department of Earth Science and Engineering Albian, Aptian, Cretaceous, Dyrosauridae, England, Ferruginous Sands Keywords: Formation, Isle of Wight, Pholidosauridae, Tethysuchia, Upper Greensand Formation Biological Journal of the Linnean Society Page 1 of 50 Biological Journal of the Linnean Society 1 2 3 Marine tethysuchian crocodyliform from the ?Aptian-Albian (Early Cretaceous) 4 5 6 of the Isle of Wight, England 7 8 9 10 by MARK T. YOUNG 1,2 *, LORNA STEEL 3, DAVIDE FOFFA 4, TREVOR PRICE 5 11 12 2 6 13 DARREN NAISH and JONATHAN P. TENNANT 14 15 16 1 17 Institute of Evolutionary Biology, School of Biological Sciences, The King’s Buildings, University 18 For Peer Review 19 of Edinburgh, Edinburgh, EH9 3JW, United Kingdom 20 21 2 School of Ocean and Earth Science, National Oceanography Centre, University of Southampton,
    [Show full text]
  • Crocodylomorpha, Neosuchia), and a Discussion on the Genus Theriosuchus
    bs_bs_banner Zoological Journal of the Linnean Society, 2015. With 5 figures The first definitive Middle Jurassic atoposaurid (Crocodylomorpha, Neosuchia), and a discussion on the genus Theriosuchus MARK T. YOUNG1,2, JONATHAN P. TENNANT3*, STEPHEN L. BRUSATTE1,4, THOMAS J. CHALLANDS1, NICHOLAS C. FRASER1,4, NEIL D. L. CLARK5 and DUGALD A. ROSS6 1School of GeoSciences, Grant Institute, The King’s Buildings, University of Edinburgh, James Hutton Road, Edinburgh EH9 3FE, UK 2School of Ocean and Earth Science, National Oceanography Centre, University of Southampton, European Way, Southampton SO14 3ZH, UK 3Department of Earth Science and Engineering, Imperial College London, London SW6 2AZ, UK 4National Museums Scotland, Chambers Street, Edinburgh EH1 1JF, UK 5The Hunterian, University of Glasgow, University Avenue, Glasgow G12 8QQ, UK 6Staffin Museum, 6 Ellishadder, Staffin, Isle of Skye IV51 9JE, UK Received 1 December 2014; revised 23 June 2015; accepted for publication 24 June 2015 Atoposaurids were a clade of semiaquatic crocodyliforms known from the Late Jurassic to the latest Cretaceous. Tentative remains from Europe, Morocco, and Madagascar may extend their range into the Middle Jurassic. Here we report the first unambiguous Middle Jurassic (late Bajocian–Bathonian) atoposaurid: an anterior dentary from the Isle of Skye, Scotland, UK. A comprehensive review of atoposaurid specimens demonstrates that this dentary can be referred to Theriosuchus based on several derived characters, and differs from the five previously recog- nized species within this genus. Despite several diagnostic features, we conservatively refer it to Theriosuchus sp., pending the discovery of more complete material. As the oldest known definitively diagnostic atoposaurid, this discovery indicates that the oldest members of this group were small-bodied, had heterodont dentition, and were most likely widespread components of European faunas.
    [Show full text]
  • 8. Archosaur Phylogeny and the Relationships of the Crocodylia
    8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988.
    [Show full text]
  • Crocodile Specialist Group Newsletter 27(3): 6-8
    CROCODILE SPECIALIST GROUP NEWSLETTER VOLUME 31 No. 1 • JANUARY 2012 - MARCH 2012 IUCN • Species Survival Commission CSG Newsletter Subscription The CSG Newsletter is produced and distributed by the Crocodile CROCODILE Specialist Group of the Species Survival Commission (SSC) of the IUCN (International Union for Conservation of Nature). The CSG Newsletter provides information on the conservation, status, news and current events concerning crocodilians, and on the SPECIALIST activities of the CSG. The Newsletter is distributed to CSG members and to other interested individuals and organizations. All Newsletter recipients are asked to contribute news and other materials. The CSG Newsletter is available as: • Hard copy (by subscription - see below); and/or, • Free electronic, downloadable copy from “http://iucncsg.org/ GROUP ph1/modules/Publications/newsletter.html”. Annual subscriptions for hard copies of the CSG Newsletter may be made by cash ($US55), credit card ($AUD55) or bank transfer ($AUD55). Cheques ($USD) will be accepted, however due to increased bank charges associated with this method of payment, cheques are no longer recommended. A Subscription Form can be NEWSLETTER downloaded from “http://iucncsg.org/ph1/modules/Publications/ newsletter.html”. All CSG communications should be addressed to: CSG Executive Office, P.O. Box 530, Karama, NT 0813, Australia. VOLUME 31 Number 1 Fax: (61) 8 89470678. E-mail: [email protected]. JANUARY 2012 - MARCH 2012 PATRONS IUCN - Species Survival Commission We thank all patrons who have donated to the CSG and its conservation program over many years, and especially to CHAIRMAN: donors in 2010-2011 (listed below). Professor Grahame Webb PO Box 530, Karama, NT 0813, Australia Big Bull Crocs! ($15,000 or more annually or in aggregate donations) Japan, JLIA - Japan Leather & Leather Goods Industries EDITORIAL AND EXECUTIVE OFFICE: Association, CITES Promotion Committee & All Japan PO Box 530, Karama, NT 0813, Australia Reptile Skin and Leather Association, Tokyo, Japan.
    [Show full text]
  • Induszalim Bala Mesoeucrocodile from Pakistan
    Open Journal of Geology, 2019, 9, 623-626 https://www.scirp.org/journal/ojg ISSN Online: 2161-7589 ISSN Print: 2161-7570 Induszalim bala Mesoeucrocodile from Pakistan Muhammad Sadiq Malkani Geological Survey of Pakistan, Muzaffarabad, Azad Kashmir, Pakistan How to cite this paper: Malkani, M.S. Abstract (2019) Induszalim bala Mesoeucrocodile from Pakistan. Open Journal of Geology, 9, Induszalim bala is the first diagnostic mesoeucrocodile from In- 623-626. do-Pakistan subcontinent having rostrum articulated with dentary sym- https://doi.org/10.4236/ojg.2019.910058 physis and associated vertebrae and limb bones and provides facility for Received: August 16, 2019 comparison with other mesoeucrocodiles. Induszalim is a medium to large Accepted: September 21, 2019 sized mesoeucrocodile found in the latest Maastrichtian Vitakri Lameta Published: September 24, 2019 Formation of Fort Munro Group just below Cretaceous-Paleogene boun- Copyright © 2019 by author(s) and dary line. Induszalim has sufficient skeletal elements and can be used for Scientific Research Publishing Inc. phylogenetic studies. Induszalim shows Gondwanan paleobiogeographic af- This work is licensed under the Creative finity with some endemic elements. Commons Attribution International License (CC BY 4.0). Keywords http://creativecommons.org/licenses/by/4.0/ Open Access Mesoeucrocodile, Induszalim bala, Latest Maastrichtian, Indo-Pakistan Subcontinent 1. Introduction Induszalim bala mesoeucrocodile as the new genus and new species was first re- ported by [1] and formally published by [2]. Here Induszalim is being described for evolutionary study. 2. Induszalim bala Mesoeucrocodile from Pakistan Systematic paleontology of Induszalim bala is as follows: Crocodyliformes, Mesoeucrocodylia, Induszalimidae [1], Induszalim [1] [2], Induszalim bala [1] [2] (Figure 1).
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • First Remains of Diplocynodon Cf. Ratelii from the Early Miocene Sites of Ahníkov (Most Basin, Czech Republic)
    First remains of Diplocynodon cf. ratelii from the early Miocene sites of Ahníkov (Most Basin, Czech Republic) Milan Chroust, Martin MazuCh, Martin ivanov, Boris Ekrt & ÀngEl h. luján Fossil crocodylians from the early Miocene (Eggenburgian, MN3a) sites of Ahníkov (Most Basin, Czech Republic) are described in this paper. The new material presented here includes over 200 remains (bones, teeth and osteoderms), and therefore constitutes the largest crocodylian sample known from the fossil record of the Czech Republic. Assignment of the specimens to the fossil alligatoroid taxon Diplocynodon cf. ratelii Pomel, 1847 (family Diplocynodontidae) is justified by the presence of several cranial and postcranial features. In the Czech Republic, this species has been previously reported only from the Tušimice site (MN3, Most Basin, Ohře/Eger Graben). The majority of the material reported from Ahníkov is composed of disarticulated juvenile individuals. Both sites are most likely attributable to the specific environment of swampy areas, where crocodile hatchlings would hide from predators. The presence of the genus Diplocynodon supports the assumption of rather warm climatic conditions in Central Europe during the early to middle Miocene, as well as a swampy depositional environment previously inferred for Ahníkov. However, some squamate taxa suggest the existence of additional, surrounding palaeoenvironment characterised by a more open landscape with slightly drier conditions. • Key words: fossil crocodiles, alligatoroid, Ahníkov, Ohře/Eger Graben, Eggenburgian. CHROUST, M., MAZUCH, M., IVANOV, M., EKRT, B. & LUJÁN, À.H. 2021. First remains of Diplocynodon cf. ratelii from the early Miocene sites of Ahníkov (Most Basin, Czech Republic). Bulletin of Geosciences 96(2), 123–138 (10 figures, 1 table).
    [Show full text]
  • Craniofacial Morphology of Simosuchus Clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar
    Society of Vertebrate Paleontology Memoir 10 Journal of Vertebrate Paleontology Volume 30, Supplement to Number 6: 13–98, November 2010 © 2010 by the Society of Vertebrate Paleontology CRANIOFACIAL MORPHOLOGY OF SIMOSUCHUS CLARKI (CROCODYLIFORMES: NOTOSUCHIA) FROM THE LATE CRETACEOUS OF MADAGASCAR NATHAN J. KLEY,*,1 JOSEPH J. W. SERTICH,1 ALAN H. TURNER,1 DAVID W. KRAUSE,1 PATRICK M. O’CONNOR,2 and JUSTIN A. GEORGI3 1Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, 11794-8081, U.S.A., [email protected]; [email protected]; [email protected]; [email protected]; 2Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio 45701, U.S.A., [email protected]; 3Department of Anatomy, Arizona College of Osteopathic Medicine, Midwestern University, Glendale, Arizona 85308, U.S.A., [email protected] ABSTRACT—Simosuchus clarki is a small, pug-nosed notosuchian crocodyliform from the Late Cretaceous of Madagascar. Originally described on the basis of a single specimen including a remarkably complete and well-preserved skull and lower jaw, S. clarki is now known from five additional specimens that preserve portions of the craniofacial skeleton. Collectively, these six specimens represent all elements of the head skeleton except the stapedes, thus making the craniofacial skeleton of S. clarki one of the best and most completely preserved among all known basal mesoeucrocodylians. In this report, we provide a detailed description of the entire head skeleton of S. clarki, including a portion of the hyobranchial apparatus. The two most complete and well-preserved specimens differ substantially in several size and shape variables (e.g., projections, angulations, and areas of ornamentation), suggestive of sexual dimorphism.
    [Show full text]
  • Postcranial Skeleton of Mariliasuchus Amarali Carvalho and Bertini, 1999 (Mesoeucrocodylia) from the Bauru Basin, Upper Cretaceous of Brazil
    AMEGHINIANA - 2013 - Tomo 50 (1): 98 – 113 ISSN 0002-7014 POSTCRANIAL SKELETON OF MARILIASUCHUS AMARALI CARVALHO AND BERTINI, 1999 (MESOEUCROCODYLIA) FROM THE BAURU BASIN, UppER CRETACEOUS OF BRAZIL Pedro HenriQue NOBRE1 and Ismar de souZA carvaLho2 1Universidade Federal de Juiz de Fora, Departamento de Ciências Naturais - CA João XXIII, Rua Visconde de Mauá 300, Bairro Santa Helena, Juiz de Fora, 36015-260 MG, Brazil. [email protected] 2Universidade Federal do Rio de Janeiro. Departamento de Geologia, CCMN/IGEO, Cidade Universitária – Ilha do Fundão, Rio de Janeiro, 21.949-900 RJ, Brazil. [email protected] Abstract. Mariliasuchus amarali is a notosuchian crocodylomorph found in the Bauru Basin, São Paulo State, Brazil (Adamantina Forma- tion, Turonian–Santonian). The main trait of M. amarali is its robust construction, featuring short, laterally expanded bones. The centra of the vertebrae are amphicoelous. In the ilium, the postacetabular process is ventrally inclined and exceeds the limits of the roof of the acetabulum. M. amarali has postcranial morphological characteristics that are very similar to those of Notosuchus terrestris, though it also displays traits resembling those of eusuchian crocodyliforms (Crocodyliformes, Eusuchia). The similarity of the appendicular skeleton of M. amarali with the recent forms of Eusuchia, leads us to infer that M. amarali did not have an erect or semi-erect posture, as proposed for the notosuchian mesoeucrocodylians, but a sprawling type posture and, possibly, had amphibian habits (sharing this characteristic with the extant Eusuchia). Key words. Mariliasuchus amarali. Crocodyliformes. Notosuchia. Postcranial skeleton. Bauru Basin. Cretaceous. Resumen. ESQUELETO POSCRANEANO DE MARILIASUCHUS AMARALI CARVALHO Y BERTINI, 1999 (MESOEUCROCO- DYLIA), DE LA CUENCA DE BAURU, CRETÁCICO SUPERIOR DE BRASIL.
    [Show full text]
  • I – Introdução
    UNIVERSIDADE ESTADUAL PAULISTA Instituto de Geociências e Ciências Exatas Campus de Rio Claro REVISÃO SISTEMÁTICA E TAXONÔMICA DOS NOTOSUCHIA (METASUCHIA, CROCODYLOMORPHA) Marco Brandalise de Andrade Orientador: Prof. Dr. Reinaldo José Bertini Dissertação de Mestrado elaborada junto ao Programa de Pós-Graduação em Geologia - Área de concentração em Geologia Regional, para a obtenção do título de Mestre em Geociências Rio Claro (SP) 2005 Comissão Examinadora _____________________________________ Prof. Dr. Reinaldo José Bertini _____________________________________ Prof. Dr. Alexander Wilhelm Armin Kellner _____________________________________ Prof. Dr. Antonio Roberto Saad _____________________________________ Marco Brandalise de Andrade - aluno - Rio Claro, ____ de ________________ de 2005 Resultado: _____________________________________________________________ AGRADECIMENTOS Ao longo do desenvolvimento deste Mestrado, em seu caminho paradoxalmente tão longo e tão breve, contei com o apoio de um grande número de pessoas. Algumas destas contribuíram de forma direta, enquanto outras sequer estavam cientes de sua importância ao longo deste processo, em uma grande diversidade de aspectos. Agradeço em primeiro lugar ao Prof. Dr. Reinaldo José Bertini, por sua orientação e pela grande atenção e paciência a este estudante, me recebendo sem reservas na cidade de Rio Claro e na UNESP, bem como na comunidade científica. Agradeço à Universidade Estadual Paulista, ao Instituto de Geociências e Ciências Exatas e ao Departamento de Geologia Aplicada, em cujas dependências o estudo encontrou condições para o seu desenvolvimento. De fundamental importância foi o suporte fornecido pela Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES), que forneceu uma Bolsa de Mestrado que em muito ajudou a ampliar o potencial e o alcance deste trabalho. Agradeço ao Prof. Dr. Antonio Roberto Saad e a Profa.
    [Show full text]
  • Novos Sítios Fossilíferos Da Formação Romualdo
    Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br Novos Sítios Fossilíferos da Formação Romualdo, Cretáceo Inferior, Bacia do Araripe, Exu, Pernambuco, Nordeste do Brasil New Fossiliferous Sites of the Romualdo Formation, Lower Cretaceous, Araripe Basin, Exu, Pernambuco, Northeast of Brazil Rudah Ruano Cavalcanti Duque & Alcina Magnólia Franca Barreto Universidade Federal de Pernambuco, Centro de Tecnologia e Geociências, Departamento de Geologia, Av. Acadêmico Hélio Ramos, s/n, 50740-530, Cidade Universitária, Recife, PE, Brasil. E-mails: [email protected]; [email protected] Recebido em: 10/01/2018 Aprovado em:01/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_05_14 Resumo Aqui são apresentados antigos e novos sítios fossilíferos da Formação Romualdo, Albiano da Bacia do Araripe, no município de Exu, Pernambuco, Nordeste do Brasil, com o levantamento da sua diversidade fossilífera, com ênfase em paleovertebrados. Foram estudadas sete localidades da porção sudoeste da Bacia Sedimentar do Araripe, sendo identificados 16 táxons de paleovertebrados, incluindo peixes: Vinctifer comptoni, Rhacolepis buccalis, Calamopleurus cylindricus, Cladocyclus gardneri, Neoproscinetes penalvai, Paraelops cearenses, Tharrhias araripis, Notelops brama, Araripelepdotes temnurus, Brannerion sp. e Beurlenichthys ouricuriensis; pterossauros Anhangueridae e quelônios (Araripemys barretoi). A Paleofauna de vertebrados da Formação Romualdo vem sendo estudada desde 1800, é reconhecida internacionalmente como Fossillagerstätte. Porém, a
    [Show full text]