Wildlife Center Classroom Series: Awesome Opossums!
Total Page:16
File Type:pdf, Size:1020Kb
Load more
Recommended publications
-
A Dated Phylogeny of Marsupials Using a Molecular Supermatrix and Multiple Fossil Constraints
Journal of Mammalogy, 89(1):175–189, 2008 A DATED PHYLOGENY OF MARSUPIALS USING A MOLECULAR SUPERMATRIX AND MULTIPLE FOSSIL CONSTRAINTS ROBIN M. D. BECK* School of Biological, Earth and Environmental Sciences, University of New South Wales, Sydney, New South Wales 2052, Australia Downloaded from https://academic.oup.com/jmammal/article/89/1/175/1020874 by guest on 25 September 2021 Phylogenetic relationships within marsupials were investigated based on a 20.1-kilobase molecular supermatrix comprising 7 nuclear and 15 mitochondrial genes analyzed using both maximum likelihood and Bayesian approaches and 3 different partitioning strategies. The study revealed that base composition bias in the 3rd codon positions of mitochondrial genes misled even the partitioned maximum-likelihood analyses, whereas Bayesian analyses were less affected. After correcting for base composition bias, monophyly of the currently recognized marsupial orders, of Australidelphia, and of a clade comprising Dasyuromorphia, Notoryctes,and Peramelemorphia, were supported strongly by both Bayesian posterior probabilities and maximum-likelihood bootstrap values. Monophyly of the Australasian marsupials, of Notoryctes þ Dasyuromorphia, and of Caenolestes þ Australidelphia were less well supported. Within Diprotodontia, Burramyidae þ Phalangeridae received relatively strong support. Divergence dates calculated using a Bayesian relaxed molecular clock and multiple age constraints suggested at least 3 independent dispersals of marsupials from North to South America during the Late Cretaceous or early Paleocene. Within the Australasian clade, the macropodine radiation, the divergence of phascogaline and dasyurine dasyurids, and the divergence of perameline and peroryctine peramelemorphians all coincided with periods of significant environmental change during the Miocene. An analysis of ‘‘unrepresented basal branch lengths’’ suggests that the fossil record is particularly poor for didelphids and most groups within the Australasian radiation. -
A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes
J Mammal Evol DOI 10.1007/s10914-007-9062-6 ORIGINAL PAPER A Phylogeny and Timescale for Marsupial Evolution Based on Sequences for Five Nuclear Genes Robert W. Meredith & Michael Westerman & Judd A. Case & Mark S. Springer # Springer Science + Business Media, LLC 2007 Abstract Even though marsupials are taxonomically less diverse than placentals, they exhibit comparable morphological and ecological diversity. However, much of their fossil record is thought to be missing, particularly for the Australasian groups. The more than 330 living species of marsupials are grouped into three American (Didelphimorphia, Microbiotheria, and Paucituberculata) and four Australasian (Dasyuromorphia, Diprotodontia, Notoryctemorphia, and Peramelemorphia) orders. Interordinal relationships have been investigated using a wide range of methods that have often yielded contradictory results. Much of the controversy has focused on the placement of Dromiciops gliroides (Microbiotheria). Studies either support a sister-taxon relationship to a monophyletic Australasian clade or a nested position within the Australasian radiation. Familial relationships within the Diprotodontia have also proved difficult to resolve. Here, we examine higher-level marsupial relationships using a nuclear multigene molecular data set representing all living orders. Protein-coding portions of ApoB, BRCA1, IRBP, Rag1, and vWF were analyzed using maximum parsimony, maximum likelihood, and Bayesian methods. Two different Bayesian relaxed molecular clock methods were employed to construct a timescale for marsupial evolution and estimate the unrepresented basal branch length (UBBL). Maximum likelihood and Bayesian results suggest that the root of the marsupial tree is between Didelphimorphia and all other marsupials. All methods provide strong support for the monophyly of Australidelphia. Within Australidelphia, Dromiciops is the sister-taxon to a monophyletic Australasian clade. -
Uso De La Cola Y El Marsupio En Didelphis Marsupialis Y Metachirus Nudicaudatus (Didelphimorphia: Didelphidae) Para Transportar Material De Anidación
University of Wollongong Research Online Faculty of Science, Medicine and Health - Papers: part A Faculty of Science, Medicine and Health 1-1-2014 Uso de la cola y el marsupio en Didelphis marsupialis y Metachirus nudicaudatus (Didelphimorphia: Didelphidae) para transportar material de anidación Carlos Delgado-Velez University of Wollongong, [email protected] Andres Arias-Alzate Universidad Nacional Autonoma de Mexico-UNAM Sebastian Aristizabal-Arango Universidad CES Juan D. Sanchez-Londono Universidad CES Follow this and additional works at: https://ro.uow.edu.au/smhpapers Part of the Medicine and Health Sciences Commons, and the Social and Behavioral Sciences Commons Recommended Citation Delgado-Velez, Carlos; Arias-Alzate, Andres; Aristizabal-Arango, Sebastian; and Sanchez-Londono, Juan D., "Uso de la cola y el marsupio en Didelphis marsupialis y Metachirus nudicaudatus (Didelphimorphia: Didelphidae) para transportar material de anidación" (2014). Faculty of Science, Medicine and Health - Papers: part A. 2438. https://ro.uow.edu.au/smhpapers/2438 Research Online is the open access institutional repository for the University of Wollongong. For further information contact the UOW Library: [email protected] Uso de la cola y el marsupio en Didelphis marsupialis y Metachirus nudicaudatus (Didelphimorphia: Didelphidae) para transportar material de anidación Abstract Information about the use of tail to carry nesting material by Neotropical marsupials is poorly documented. Based on videoclips obtained by camera traps, we documented the behavior of gathering and carrying nesting material in curling tails by Didelphis marsupialis and Metachirus nudicaudatus. Additionally, we documented for the fist time an individual of .D marsupialis gathering leaves and other nesting material in the pouch. -
A Species-Level Phylogenetic Supertree of Marsupials
J. Zool., Lond. (2004) 264, 11–31 C 2004 The Zoological Society of London Printed in the United Kingdom DOI:10.1017/S0952836904005539 A species-level phylogenetic supertree of marsupials Marcel Cardillo1,2*, Olaf R. P. Bininda-Emonds3, Elizabeth Boakes1,2 and Andy Purvis1 1 Department of Biological Sciences, Imperial College London, Silwood Park, Ascot SL5 7PY, U.K. 2 Institute of Zoology, Zoological Society of London, Regent’s Park, London NW1 4RY, U.K. 3 Lehrstuhl fur¨ Tierzucht, Technical University of Munich, Alte Akademie 12, 85354 Freising-Weihenstephan, Germany (Accepted 26 January 2004) Abstract Comparative studies require information on phylogenetic relationships, but complete species-level phylogenetic trees of large clades are difficult to produce. One solution is to combine algorithmically many small trees into a single, larger supertree. Here we present a virtually complete, species-level phylogeny of the marsupials (Mammalia: Metatheria), built by combining 158 phylogenetic estimates published since 1980, using matrix representation with parsimony. The supertree is well resolved overall (73.7%), although resolution varies across the tree, indicating variation both in the amount of phylogenetic information available for different taxa, and the degree of conflict among phylogenetic estimates. In particular, the supertree shows poor resolution within the American marsupial taxa, reflecting a relative lack of systematic effort compared to the Australasian taxa. There are also important differences in supertrees based on source phylogenies published before 1995 and those published more recently. The supertree can be viewed as a meta-analysis of marsupial phylogenetic studies, and should be useful as a framework for phylogenetically explicit comparative studies of marsupial evolution and ecology. -
Heterothermy in Pouched Mammals a Review
bs_bs_bannerJournal of Zoology Journal of Zoology. Print ISSN 0952-8369 MINI-SERIES Heterothermy in pouched mammals – a review A. Riek1,2 & F. Geiser2 1 Department of Animal Sciences, University of Göttingen, Göttingen, Germany 2 Centre for Behavioural and Physiological Ecology, Zoology, University of New England, Armidale, NSW, Australia Keywords Abstract heterothermy; marsupials; phylogeny; torpor; hibernation. Hibernation and daily torpor (i.e. temporal heterothermy) have been reported in many marsupial species of diverse families and are known to occur in ∼15% of all Correspondence marsupials, which is a greater proportion than the percentage of heterothermic Alexander Riek, Department of Animal placentals. Therefore, we aimed to gather data on heterothermy, including Sciences, University of Göttingen, minimal body temperature, torpor metabolic rate and torpor bout duration for Albrecht-Thaer-Weg 3, 37075 Göttingen, marsupials, and relate these physiological variables to phylogeny and other Germany. Tel: +49 551 395610; Fax: +49 physiological traits. Data from published studies on 41 marsupial species were 551 39 available for the present analysis. Heterothermic marsupials ranged from small Email: [email protected] species such as planigales weighing 7 g to larger species such as quolls weighing up to 1000 g. We used the marsupial phylogeny to estimate various heterothermic Editor: Heike Lutermann traits where the current dataset was incomplete. The torpor metabolic rate in relation to basal metabolic rate (%) ranged from 5.2 to 62.8% in daily Received 13 May 2013; revised 31 July heterotherms and from 2.1 to 5.2% in marsupial hibernators, and was significantly 2013; accepted 8 August 2013 correlated with the minimum body temperature in daily heterotherms (R2 = 0.77, P < 0.001), but not in hibernators (R2 = 0.10, P > 0.05). -
Chironectes Minimus (Zimmermann
Check List 9(5): 1104–1007, 2013 © 2013 Check List and Authors Chec List ISSN 1809-127X (available at www.checklist.org.br) Journal of species lists and distribution N Glironia venusta Thomas, 1912 and Chironectes minimus (Zimmermann, ISTRIBUTIO Mammalia, Didelphimorphia, Didelphidae, D Natália Ardente 1* 2 3 1 and Fernanda RAPHIC G 1780): Distribution extension for eastern Amazonia EO 4 , Donald Gettinger , Raul Fonseca , Helena de Godoy Bergallo G N Martins-Hatano O 1 Universidade do Estado do Rio de Janeiro, Instituto de Biologia Roberto Alcântara Gomes, Departamento de Ecologia, Laboratório de Ecologia de OTES Mamíferos. Rua São Francisco Xavier 524. CEP 20559-900. Rio de Janeiro, RJ, Brazil. N 2 University of Nebraska-Lincoln, Harold W. Manter Laboratory of Parasitology. W529 Nebraska Hall. 68588-0514. Lincoln, Nebraska, U.S.A. 3 Universidade do Estado do Rio de Janeiro, Instituto de Biologia Roberto Alcântara Gomes, Departamento de Zoologia, Laboratório de Mastozoologia. Rua São Francisco Xavier 524. CEP 20559-900. Rio de Janeiro, RJ, Brazil. 4 Universidade Federal Rural da [email protected]ônia, Instituto de Saúde e Produção Animal, , Departamento de Biologia Animal. Avenida Presidente Tancredo Neves, 2501. CEP 66.077-901. Belém, PA, Brazil. * Corresponding author. E-mail: Abstract: We report a new record for the bushy-tailed opossum Glironia venusta Thomas, 1912 and the water opossum Chironectes minimus (Zimmermann, 1780) in the Floresta Nacional de Carajás, municipality of Parauapebas, state of Pará, Brazil (06°03’00” S, 50°15’00” W). This study represents the second record of Glironia venusta for eastern Brazilian Amazon. For Chironectes minimus, this record extends the range of the species 558 km southeastwards from the closest known locality in Pará. -
An Exceptionally Well-Preserved Skeleton of Palaeothentes from the Early Miocene of Patagonia, Argentina
Swiss J Palaeontol (2014) 133:1–21 DOI 10.1007/s13358-014-0063-9 An exceptionally well-preserved skeleton of Palaeothentes from the Early Miocene of Patagonia, Argentina: new insights into the anatomy of extinct paucituberculatan marsupials Analia M. Forasiepi • Marcelo R. Sa´nchez-Villagra • Thomas Schmelzle • Sandrine Ladeve`ze • Richard F. Kay Received: 8 September 2013 / Accepted: 13 February 2014 / Published online: 27 May 2014 Ó Akademie der Naturwissenschaften Schweiz (SCNAT) 2014 Abstract During the Cenozoic paucituberculatans were an anterior semicircular canal (SC) projecting slightly much more diverse taxonomically and ecomorphologically dorsally from the dorsal-most point of the posterior SC, and than the three extant genera of shrew-like marsupials. lateral and posterior SCs projecting laterally to the same Among paucituberculatans, palaeothentids were abundant level. On the basis of postcranial anatomy, previous studies during the Early Miocene, although most of the fossil have demonstrated that P. lemoinei was an agile cursorial remains consist of isolated teeth or fragmentary jaws. We form, an inference supported by study of the new post- describe a new and exceptional partial skeleton of Palaeo- cranial elements. thentes lemoinei (Palaeothentidae), collected from the Santa Cruz Formation (Santacrucian age, Early Miocene) Keywords Marsupialia Á Metatheria Á Cenozoic Á in Patagonia. Whereas the skull of P. lemoinei has more South America Á Skull Á Inner ear Á Postcranium plesiomorphic traits in the face, palate, and cranial vault than that of living paucituberculatans, the dental mor- Abbreviations phology is more derived. The osseous inner ear was examined using micro-CT scanning, revealing a cochlea Institutional abbreviations with 1.9 turns, the presence of a ‘‘second crus commune’’, FMNH Field Museum of Natural History, Chicago, USA IEEUACH Universidad Austral de Chile, A. -
A Evolução Dos Metatheria: Sistemática, Paleobiogeografia, Paleoecologia E Implicações Paleoambientais
UNIVERSIDADE FEDERAL DE PERNAMBUCO CENTRO DE TECNOLOGIA E GEOCIÊNCIAS PROGRAMA DE PÓS-GRADUAÇÃO EM GEOCIÊNCIAS ESPECIALIZAÇÃO EM GEOLOGIA SEDIMENTAR E AMBIENTAL LEONARDO DE MELO CARNEIRO A EVOLUÇÃO DOS METATHERIA: SISTEMÁTICA, PALEOBIOGEOGRAFIA, PALEOECOLOGIA E IMPLICAÇÕES PALEOAMBIENTAIS RECIFE 2017 LEONARDO DE MELO CARNEIRO A EVOLUÇÃO DOS METATHERIA: SISTEMÁTICA, PALEOBIOGEOGRAFIA, PALEOECOLOGIA E IMPLICAÇÕES PALEOAMBIENTAIS Dissertação de Mestrado apresentado à coordenação do Programa de Pós-graduação em Geociências, da Universidade Federal de Pernambuco, como parte dos requisitos à obtenção do grau de Mestre em Geociências Orientador: Prof. Dr. Édison Vicente Oliveira RECIFE 2017 Catalogação na fonte Bibliotecária: Rosineide Mesquita Gonçalves Luz / CRB4-1361 (BCTG) C289e Carneiro, Leonardo de Melo. A evolução dos Metatheria: sistemática, paleobiogeografia, paleoecologia e implicações paleoambientais / Leonardo de Melo Carn eiro . – Recife: 2017. 243f., il., figs., gráfs., tabs. Orientador: Prof. Dr. Édison Vicente Oliveira. Dissertação (Mestrado) – Universidade Federal de Pernambuco. CTG. Programa de Pós-Graduação em Geociências, 2017. Inclui Referências. 1. Geociêcias. 2. Metatheria . 3. Paleobiogeografia. 4. Paleoecologia. 5. Sistemática. I. Édison Vicente Oliveira (Orientador). II. Título. 551 CDD (22.ed) UFPE/BCTG-2017/119 LEONARDO DE MELO CARNEIRO A EVOLUÇÃO DOS METATHERIA: SISTEMÁTICA, PALEOBIOGEOGRAFIA, PALEOECOLOGIA E IMPLICAÇÕES PALEOAMBIENTAIS Dissertação de Mestrado apresentado à coordenação do Programa de Pós-graduação -
AGILE GRACILE OPOSSUM Gracilinanus Agilis (Burmeister, 1854 )
Smith P - Gracilinanus agilis - FAUNA Paraguay Handbook of the Mammals of Paraguay Number 35 2009 AGILE GRACILE OPOSSUM Gracilinanus agilis (Burmeister, 1854 ) FIGURE 1 - Adult, Brazil (Nilton Caceres undated). TAXONOMY: Class Mammalia; Subclass Theria; Infraclass Metatheria; Magnorder Ameridelphia; Order Didelphimorphia; Family Didelphidae; Subfamily Thylamyinae; Tribe Marmosopsini (Myers et al 2006, Gardner 2007). The genus Gracilinanus was defined by Gardner & Creighton 1989. There are six known species according to the latest revision (Gardner 2007) one of which is present in Paraguay. The generic name Gracilinanus is taken from Latin (gracilis) and Greek (nanos) meaning "slender dwarf", in reference to the slight build of this species. The species name agilis is Latin meaning "agile" referring to the nimble climbing technique of this species. (Braun & Mares 1995). The species is monotypic, but Gardner (2007) considers it to be composite and in need of revision. Furthermore its relationship to the cerrado species Gracilinanus agilis needs to be examined, with some authorities suggesting that the two may be at least in part conspecific - there appear to be no consistent cranial differences (Gardner 2007). Costa et al (2003) found the two species to be morphologically and genetically distinct and the two species have been found in sympatry in at least one locality in Minas Gerais, Brazil (Geise & Astúa 2009) where the authors found that they could be distinguished on external characters alone. Smith P 2009 - AGILE GRACILE OPOSSUM Gracilinanus agilis - Mammals of Paraguay Nº 35 Page 1 Smith P - Gracilinanus agilis - FAUNA Paraguay Handbook of the Mammals of Paraguay Number 35 2009 Patton & Costa (2003) commented that the presence of the similar Gracilinanus microtarsus at Lagoa Santa, Minas Gerais, the type locality for G.agilis , raises the possibility that the type specimen may in fact prove to be what is currently known as G.microtarsus . -
PYGMY SHORT-TAILED OPOSSUM Monodelphis Kunsi Pine, 1975
Smith P - Monodelphis kunsi - FAUNA Paraguay Handbook of the Mammals of Paraguay Number 18 2008 PYGMY SHORT-TAILED OPOSSUM Monodelphis kunsi Pine, 1975 FIGURE 1 - Adult, Bolivia (Louise Emmons). TAXONOMY: Class Mammalia; Subclass Theria; Infraclass Metatheria; Magnorder Ameridelphia; Order Didelphimorphia; Family Didelphidae; Subfamily Marmosinae; Tribe Monodelphini (Myers et al 2006). Twenty-nine species are recognised in this genus, three are present in Paraguay. The scientific name Monodelphis is derived from the Greek meaning "single womb" in reference to the lack of a pouch. The species name kunsi is in honour of Dr Merle L. Kuns who obtained the type specimen. The species is monotypic. However what is currenty understood as " Monodelphis kunsi " may in fact be a cryptic species complex given the wide range and diverse habitats in which the "species" has been found, its rarity being apparently inconsistent with a habitat generalist. A similar pattern of distribution is exhibited by three similar species of Thylamys (velutinus, macrurus and venustus ). There is notable variation in Smith P 2008 - PYGMY SHORT-TAILED OPOSSUM Monodelphis kunsi - Mammals of Paraguay Nº 18 Page 1 Smith P - Monodelphis kunsi - FAUNA Paraguay Handbook of the Mammals of Paraguay Number 18 2008 the specimens, but currently so few specimens are available that it is not possible to make any firm conclusions (de la Sancha et al 2007). Synonyms adapted from Anderson (1982) and Gardner (2007). Monodelphis kunsi, Pine 1975 W bank of Rio Itonamas, 4km N of Magdalena, Departamento Beni, Bolivia. ENGLISH COMMON NAMES: Pygmy Short-tailed Opossum SPANISH COMMON NAMES: Colicorto pigmeo (Emmons 1999). GUARANÍ COMMON NAMES: No known names. -
List of 28 Orders, 129 Families, 598 Genera and 1121 Species in Mammal Images Library 31 December 2013
What the American Society of Mammalogists has in the images library LIST OF 28 ORDERS, 129 FAMILIES, 598 GENERA AND 1121 SPECIES IN MAMMAL IMAGES LIBRARY 31 DECEMBER 2013 AFROSORICIDA (5 genera, 5 species) – golden moles and tenrecs CHRYSOCHLORIDAE - golden moles Chrysospalax villosus - Rough-haired Golden Mole TENRECIDAE - tenrecs 1. Echinops telfairi - Lesser Hedgehog Tenrec 2. Hemicentetes semispinosus – Lowland Streaked Tenrec 3. Microgale dobsoni - Dobson’s Shrew Tenrec 4. Tenrec ecaudatus – Tailless Tenrec ARTIODACTYLA (83 genera, 142 species) – paraxonic (mostly even-toed) ungulates ANTILOCAPRIDAE - pronghorns Antilocapra americana - Pronghorn BOVIDAE (46 genera) - cattle, sheep, goats, and antelopes 1. Addax nasomaculatus - Addax 2. Aepyceros melampus - Impala 3. Alcelaphus buselaphus - Hartebeest 4. Alcelaphus caama – Red Hartebeest 5. Ammotragus lervia - Barbary Sheep 6. Antidorcas marsupialis - Springbok 7. Antilope cervicapra – Blackbuck 8. Beatragus hunter – Hunter’s Hartebeest 9. Bison bison - American Bison 10. Bison bonasus - European Bison 11. Bos frontalis - Gaur 12. Bos javanicus - Banteng 13. Bos taurus -Auroch 14. Boselaphus tragocamelus - Nilgai 15. Bubalus bubalis - Water Buffalo 16. Bubalus depressicornis - Anoa 17. Bubalus quarlesi - Mountain Anoa 18. Budorcas taxicolor - Takin 19. Capra caucasica - Tur 20. Capra falconeri - Markhor 21. Capra hircus - Goat 22. Capra nubiana – Nubian Ibex 23. Capra pyrenaica – Spanish Ibex 24. Capricornis crispus – Japanese Serow 25. Cephalophus jentinki - Jentink's Duiker 26. Cephalophus natalensis – Red Duiker 1 What the American Society of Mammalogists has in the images library 27. Cephalophus niger – Black Duiker 28. Cephalophus rufilatus – Red-flanked Duiker 29. Cephalophus silvicultor - Yellow-backed Duiker 30. Cephalophus zebra - Zebra Duiker 31. Connochaetes gnou - Black Wildebeest 32. Connochaetes taurinus - Blue Wildebeest 33. Damaliscus korrigum – Topi 34. -
FIELDIANA Geology
FIELDIANA Geology Publistied by Field Museum of Natural History New Series, No. 8 THE FAMILIES AND GENERA OF MARSUPIALIA LARRY G.MARSHALL •.981 UBRARY FIELD m^cim July 20, 1981 Publication 1320 THE FAMILIES AND GENERA OF MARSUPIALIA FIELDIANA Geology Published by Field Museum of Natural History New Series, No. 8 THE FAMILIES AND GENERA OF MARSUPIALIA LARRY G.MARSHALL Assistant Curator of Fossil Mammals DqMirtment of Geology Field Museum of Natural History Accepted for publication August 6, 1979 July 20, 1981 Publication 1320 Library of Congress Catalog No.: 81-65225 ISSN 0096-2651 PRINTED IN THE UNITED STATES OF AMERICA CONTENTS Part A 1 i^4troduc^on 1 Review of History and Development of Marsupial Systematics 1 Part B 19 Detailed Classification of Families and Genera of Marsupialia 19 I. New World and European Marsupialia 19 Fam. Didelphidae 19 Subfam. Dideiphinae 19 Subfam. Caluromyinae 21 *Subfam. Glasbiinae 21 *Subfam. Caroloameghiniinae 21 •Fam. Sparassocynidae 21 •Fam. Pediomyidae 21 Fam. Microbiotheriidae 21 •Fam. Stagodonddae 22 •Fam. Borhyaenidae 22 •Subfam. Hathlyacyninae 22 •Subfam. Borhyaeninae 23 •Subfam. Prothylacyninae 23 •Subfam. Proborhyaeninae 23 •Fam. Thylacosmilidae 23 •Fam. Argyrolagidae 24 Fam. Caenolesddae 24 Subfam. Caenolestinae 24 Tribe Caenolestini 24 •Tribe Pichipilini 24 •Subfam. Palaeothentinae 24 •Subfam. Abderitinae 25 •Tribe Parabderitini 25 •Tribe Abderitini 25 •Fam. Polydolopidae 25 •Fam. Groeberiidae 25 Marsupialia incertae sedis 25 Marsupialia(?) 25 II. Australasian Marsupialia 26 Fam. Dasyuridae 26 Subfam. Dasyurinae 26 Tribe Dasyurini 26 Tribe Sarcophilini 26 Fam. Myrmecobiidae 27 •Fam. Thylacinidae 27 Fam. Peramelidae 27 Fam. Thylacomyidae 27 Fam. Notoryctidae 27 Fam. Phalangeridae 27 Subfam. Phalangerinae 27 Subfam. Trichosurinae 28 •Fam.