A redescription of the South American

Loricariichthys maculatus (Bloch, 1794), with designation

of the lectotype and restriction of its type locality

(Pisces, Siluriformes, Loricariidae)

by

I.J.H. Isbrücker

Dotterbloemstraat5, Amsterdam-N 1019, The Netherlands

Abstract scription and figures there is no doubt what

was involved. To further nomenclatural The type species of the genus Loricariichthys Bleeker, stability, the lectotype of Loricaria maculata is 1862, Loricariichthys maculatus (Bloch, 1794), is re- the for Pleco- described designated simultaneously neotype from two syntypes, one of which is designated

the lectotype. The same specimen is also designated the stomus cataphracta Gronovius, 1854, the latter

for Plecostomus Gronovius neotype cataphracta (ed. thus automatically becoming an objective junior Gray), 1854 (non Loricaria Linnaeus, cataphracta 1758, of former. The of Loricari- synonym the presence sensu stricto). The type locality of both nominal species is maculatus in Surinam has been recorded restricted after ichthys a comparison to specimens recently col-

in several works the to me it lected from a locality in Surinam. on group, and ap-

peared relevant to restrict the type locality to a

INTRODUCTION place in Surinam.

In 1862 Bleeker established the generic name The nomenclatural monotypic Loricaria genus Loricariichthys, with Loricaria maculata Bloch, Linnaeus, 1758, was originally diagnosed from two 1794, as the type species, by original monotypy. different widely biological species, called Loricaria used times in This generic name has been many Linnaeus in cataphracta by 1758. By common con- ichthyological nomenclature, but not always cor-

sent, the species described as “Loricaria dura” by rectly. So the present concept of Loricariichthys Linnaeus in 1754 (pre-Linnaean) became in almost might confuse the worker who attempts to classify all literature fixed nomenclaturally as representing certain loricariid fishes.

the Loricaria The cataphracta. other species was of the is to The main purpose present paper named Loricaria maculata by Bloch in 1794. This redescribe the type species of the genus (which

specific name was accepted by all subsequent from a systematic point of view is as valid as authors except Gronovius, who, in his posthumous limited from Loricaria) in its most sense, viz., type work (edited by 1854 restricted the Gray) of spe- specimens, supplemented with specimens from the

cific name cataphracta (in the genus Plecostomus) type locality. Employing recently obtained material to the fish he first described “Plecostomus as no. from Surinam which has been found identical to 68" in 1754. It has been accepted, even by Bloch, Bloch's original type specimens (which are rede- that the first description on which the Plecostomus it scribed), was possible to restrict the type locality, of Gronovius cataphracta (1854) was based, viz. as a starting point towards a revision of the genus.

his “Plecostomus no. 68" (1754) refers to the

same that species Bloch described as Loricaria

and the MEASUREMENTS maculata, present author agrees. Bloch did

mention for axial not a locality his new species; Grono- In the description of the specimens the

Surinam the standard the vius, however, recorded as locality for length equals the length plus length his “Plecostomus no. 68". Although Gronovius' of the middle caudal fin rays or the membrane

material is from the no longer extant, from both his de- between them; the head length is 41 - 1971 11 BIJDRAGEN TOT DE DIERKUNDE, (1)

68”. Fig. 1. Reproduction of Gronovius’ (1754) figures of “Plecostomus no.

maculata. Fig. 2. Reproduction of Bloch’s (1794) figures of Loricaria 12 I. J. H. ISBRUCKER - LORICARIICHTHYS MACULATUS

collected M. E. end of the 18th tip of the snout to the end of the occipital process; Surinam), by Bloch,

century; at the same time, this specimen is the neotype ') the h e a d width is taken at the opercle, just for Plecostomus cataphracta Gronovius (ed. Gray), 1854 before the pectoral fin spine insertion; the head (in Museum fiir Naturkunde der Humboldt-Universitàt end of the depth is taken at the occipital pro- — zu Berlin). ZMB 22224. paralectotype ') of Loricaria the is from the of the cess; snout length tip maculata Bloch, 1794, sex unknown. 109.0 mm standard

data 3163. the time. snout to the anterior border of the orbital rim; the length, same as ZMB and at same this specimen is the paraneotype ') for Plecostomus cata- thoracic length is taken between the spine

phracta Gronovius (ed. Gray), 1854. — ZMA 109.846 insertions of the pectoral and pelvic fins; and the and ZMA 109.847, a $, 163.0 mm standard length and abdominal length is taken between the a 9, 175.0 mm standard length, respectively, topotypes,

spine insertions of the pelvic and anal fins. All Surinam, district Suriname, ditches at Cultuurtuin at

Paramaribo West collected H. other measurements are self-explanatory. The (05°50'N, 55°10'W), by

2 Nijssen (Brokopondo Research 1966/1967 )), 12-XII- measurements were taken directly.

1966. — ZMA 105.307, 35 specimens, topotypes, same

data as ZMA 109.846 and ZMA 109.847. Loricariichthys maculatus (Bloch, 1794)

(figs. 1—7) The following description is based on ZMB

Type locality, by present restriction. — Surinam (for- 3163 (a J1 ), ZMB 22224 (a juvenile), ZMA

merly Dutch Guiana): district Suriname, ditches at Cul- 109.846 (a J), and ZMA 109.847 (a ?). — tuurtuin (= Botanical Gardens) at Paramaribo West Morphometric and meristic data: standard length (05 °50' N, 55° 10' W). axial 214.0, 109.0, 163.0, 175.0 mm; length 234.5,

Loricaria cataphracta; Linnaeus, 1758 : 307 (in part, 192.8 total 119.5, 181.2, mm; length damaged,

sensu lato [non Loricaria cataphracta Linnaeus, 1758 damaged, damaged, 211.2 mm; predorsal length 307, sensu strictol: in the original diagnosis of 'variant' 66.1, 33.0, 53.7, 54.8 mm (3.2, 3.3, 3.0. 3.2 in "1"; reference to “Plecostomus no. 68" Gronovius, 1754, standard head 36.0 and diagnosis). Linnaeus (ed. Gmelin), 1789: 1363 length); length 44.7, 22.2, 36.3,

(same as preceding). mm (4.8, 4.9, 4.6, 4.9 in standard length); head

width 33.5, 16.4, 27.8, 29.5 mm (6.4, 6.7, 5.9. 5.9 Loricaria maculat a Bloch, 1794 : 73—75, pi. 375 figs. in standard 1.2 in head 1 —2 (original description; no locality mentioned; lecto- length, 1.3, 1.4, 1.3,

in Museum dcr Humboldt-Univer- type Zoologisches an length); head depth 18.4, 8.7, 14.6, 16.0 mm (11.6,

sitat zu Berlin, ZMB 3163). 12.5, 11.2, 10.9 in standard length, 2.4, 2.5, 2.5,

2.3 in head snout 22.4, 10.2. 18.9, Loricaria Cirrhosa var. Maculata; Bloch & Schneider, length); length

1801: xxxii and 125 (in index), page (diagnosis, after 18.6 mm (9.6, 10.7, 8.6, 9.4 in standard length, Loricaria maculata Bloch, 1794). 2.0, 2.1, 1.9, 1.9 in head length); orbital diameter

7.3, 4.4, 6.4, 6.0 mm (6.1, 5.0, 6.0 in head Plecostomus maculatus; Swainson, 1839 : 304 (in part; 5.7,

after diagnosis, copied data). length); interorbital width 10.0, 4.6, 8.6, 8.8 mm

(4.5, 4.8, 4.2, 4.1 in head length); internasal width Loricariichthys maculatus; Bleeker, 1862a : 3 (generic 7.3, 6.3 5.6, 5.5, 5.7 in head diagnosis; type species, by original monotypy, of the 4.0, 6.6, mm (6.1,

new genus). length); dorsal spine length 44.0, 26.1, 41.5, 42.0

mm (4.9, 4.2, 3.9, 4.2 in standard length); length Plecostomus cataphracta Gronovius (ed. Gray), 1854:

158 first dorsal 40.2 mm [non Loricaria cataphracta Linnaeus, 1758, sensu ray 42.5, 25.0, 38.8, (5.0, 4.4,

the considered the stricto]; (in present paper original 4.2, 4.4 in standard length, 1.1, 0.9, 0.9, 0.9 in of diagnosis a new species, after “Plecostomus no. 68" head length); length last dorsal ray 18.8, 8.7. 15.4, in Gronovius, 1754; neotype Zoologisches Museum an 15.0 mm (11.4, 12.5, 10.6, 11.7 in standard length, der Humboldt-Universitat zu Berlin, ZMB 3163). 2.4, 2.5, 2.4, 2.4 in head length); length dorsal fin

“Plecostomus no. 68" Gronovius, 1754 : 25—26, pi. 2 base 18.8, 10.0, 15.4, 17.1 mm (11.4, 11.0, 10.6, figs. 1 —2 (pre-Linnaean and non-binominal, therefore 10.2 in standard length, 2.4, 2.2. 2.4, 2.1 in head nomenclaturally invalid; description; Surinam). length); anal spine length 36.9, 19.0, 31.0, > 32.0 “Plecostomus 391" no. Gronovius, 1763 : 127, pi. 2 in mm (5.8, 5.7, 5.3, < 5.5 standard length, 1.2, figs. I—2 (non-binominal, therefore nomenclaturally in-

valid; diagnosis, after “Plecostomus no. 68" Gronovius, 2 ) The Brokopondo Research Project, biological investi- 1754). gations in the artificial "Prof. Dr. Ir. W. J. van Blomme-

examined. — Material ZMB 3163, lectotype ')• by present stein Lake" (= Brokopondo Lake) in the Suriname

of Loricaria designation, maculata Bloch, 1794, a $, River (8 km S. of Brokopondo), Surinam; sponsored by

214.0 mm standard length. South America (probably the Netherlands Foundation for the Advancement of

Tropical Research. Cf. Nijssen & Isbriicker, 1967, 1968;

') See also discussion. Mees, 1967; Boeseman, 1968, 1969; Nijssen, 1970. BIJDRAGEN TOT DE DIERKUNDE, 41 (1) - 1971 13

Figs. 3 —6. Loricariichthys maculatus. 3, lectotype in dorsal view; 4, lectotype in ventral view; 5, topotype (￿),

ZMA 109.846; 6, topotype (￿), ZMA 109.847 (notice secondary sexual dimorphism in the lips in figs. 5 and 6). 14 I. J. H. ISBRUCKER - LORICARIICHTHYS MACULATUS

1.2, 1.2, <1.1 in head length); pectoral spine

length 34.3, 15.3, 28.2, 28.4 mm (6.2, 7.1, 5.8, 6.2

in standard length, 1.3, 1.5, 1.3, 1.3 in head

length); pelvic spine length 38.9, 18.3, 30.1, 33.0

mm (5.5, 5.9, 5.4, 5.3 in standard length, 1.1, 1.2,

1.2, 1.1 in head length); length upper principal

caudal 'spine' (unbranched ray) damaged, damag-

ed, damaged, 36.6 mm; length lower principal

caudal 'spine' (unbranched ray) damaged, 15.0,

24.4, damaged, mm (?.?, 7.3, 6.7, ?.?, in standard

length, ?.?, 1.5, 1.5, ?.?, in head length); cleithral

width 32.0, 16.6, 26.6, 28.3 mm (6.7, 6.6, 6.1, 6.2

in standard length, 1.4, 1.3, 1.4, 1.3 in head length);

supracleithral width 26.3, 13.3, 20.8, 21.9 mm (8.1,

8.2, 7.8, 8.0 in standard length, 1.7, 1.7, 1.7, 1.6

in head length); thoracic length 21.1, 9.5, 16.1,

11.5 in standard 15.1 mm (10.1, 11.5, 10.1, length. 2.1, 2.3, 2.3, 2.4 in head length); abdominal length

32.3, 15.3, 25.8, 27.0 mm (6.6, 7.1, 6.3, 6.5 in Fig. 7. Loricariichthys maculatus. Profiles of the teeth standard 1.5, 1.4, 1.3 in head length, 1.4, length); from: of the lower of the a, upper jaw lectotype, b, jaw

110.0, 57.4, 80.0, 89.0 lower of post-anal peduncular length lectotype, c, jaw the ￿ topotype, d, upper jaw

of of the f, mm (1.9, 1.9, 2.0, 2.0 in standard length); depth the ￿ topotype, e, upper jaw paralectotype, of the lower of lower jaw paralectotype, g, jaw the ￿ caudal peduncle 3.8, 1.6, 2.7, 2.7 mm (11.8, 13.9, of Each letter topotype, h, upper jaw the ￿ topotype. 13.4, 13.3 in head length); width caudal peduncle represents a single tooth. 7.5, 3.8, 6.8, 6.6 mm (6.0, 5.8, 5.3, 5.5 in head

between fin length); distance anus and anal origin short subbarbels, or large papillae. Upper jaw with

16.2, 9.6, 13.2, 14.8 mm (13.2, 11.3, 12.3, 11.8 in (left/right) ?8/?8, ?4/?, 5/5, 5/10 rudimentary

standard length, 2.8, 2.3, 2.7, 2.4 in head length); small teeth (found to be disarranged and difficult

length longest barbel of the upper lip 29.3, 10.2, to count in the old specimens) and lower jaw with

22.0, 19.6 mm (7.3, 10.7, 7.4, 8.9 in standard (left/right) 6/9. 9/5, 12/9, 8/12 small teeth. There

1.8 in head length, 1.5, 2.2, 1.6, length); greatest is a considerable variation in teeth shape, as shown

axial length of the lower lip 23.0, 5.5, 22.2, 10.2 diagramatically in fig. 7. In the lectotype and in

mm (1.9, 4.0, 1.6, 3.5 in head length); barbels along the male topotype the teeth in both jaws are

the of the lower absent in small lobe Viewed edge lip males, bilobed, each is spoon shaped. from

in females. lobes each the front the are at an angle to other.

Body scutes in longitudinal lateral series 33, 32, In the paralectotype the teeth in both jaws are

last 32, 32, scute on caudal peduncle. Scutes be- spatula shaped. In the female topotype the teeth

tween dorsal and caudal fin 27, 26. resemble spine 27, 27, in the upper jaws those of the paralecto-

Scutes between anal fin and caudal fin bifid. origin 23, type, whereas the teeth in the lower jaws are

22, 22, 22. Post-occipital scutes 2 in all four speci- Some teeth removed for examination.

mens, predorsal shield not included. Between anus Eye round, the pupil partly covered by a flap

and anal fin origin 3 pairs of scutes in all four originating from the iris. The flap does not reach

specimens, an anal plate present. On the thorax, the ventralmost inner edge of the iris and is between last the pectoral fin ray and the pelvic rounded below. A deep posterior notch is present. fin fins spine (left/right) 5/4, 3/3, 5/4, 5/4 oblong Dermal ossifications, spines and rays of the

The scutes are present. belly is completely covered are covered by minute denticles. There are two

by large platelets between the oblong thoracic distinct ridges along the lateral body scutes, coales-

scutes (figs. 1, 2, and 4). cing on the (left/right) 18th/18th, 18th/19th, 18th/

Dorsal fin last 1,6, ray split to its base; anal fin 18th, 18th/18th scute. There are some inconspic-

its in the 1,4, last ray split to base; pectoral fin 1,6; pelvic uous ridges on the predorsal scutes para- fin 1,5; caudal fin and not in the adult principal 'spines' rays 1,10,1. lectotype, present specimens.

Fin counts are equal in all four specimens. Denticles are lacking in a narrow area along the

barbels Two long rictal with about 8—12 very snout profile. This area is bordered ventrally by BIJDRAGEN TOT DE DIERKUNDE, 41 (1) - 1971 15

the ventral snout margin, covered by denticles. tained two errors. First, his reference to "Plecosto-

Irregular structure of platelets covers head and mus corpore aculeato, ore cirrato, dorso mono-

snout dorsally, well visible in the old specimens, pterygio. Arted. mss. ad Sebam." should have been

but less distinct in the topotypes. The spines of placed under the description of Gronovius' “Ple- the fins are slender, terminally flexible and costomus no. 69" [ = Loricaria cataphracta Lin-

thicker canal 1758, Seba's slightly than adjacent rays. Sensory naeus, sensu stricto]. Although (1758,

well both the be- 29 resembles Lori- system developed, on head and pi. fig. 14) figure superficially

tween the two lateral ridges of denticles. Most of cariichthys maculatus rather than Loricaria cata-

canals between lateral it is evident from the sensory the body scutes phracta (s.s.). his descriptions are bifurcated. (page 88, in Latin and Dutch) that he had a fish

The lower lip of the males is large, the entire closely related or identical to the species known

surface being covered by minute, regularly sized today as Loricaria cataphracta Linnaeus, 1758.

Gronovius stated in both papillae. There are two fleshy, rectangular flaps, Second, the diagnosis

devoid of papillae, in the buccal cavity, just below and in the description ("Dentes in labiis nulli con-

females is that toothless. the upper jaws. The lower lip of the spicui.") “Plecostomus no. 68" was smaller. Small papillae are present on 'cushions', in size towards the and increasing edges, becoming The original diagnosis by Linnaeus (1758 : barbel-like along the edge (fig. 6). The important 307—308) of Loricaria cataphracta (type species, sexual dimorphism in the lower lip is functional; by original monotypy, of Loricaria Linnaeus, the male is known to take care of the eggs (Mene- 1758) reads:

zes, 1949, for Loricaria typus (Bleeker, 1862)

[this species is said to be identical to Loricari- cataphracta 1. Loricaria. Mus. Ad. Fr.

1. 79. t. 29. f. 1. D. P. V. . A. C. 12. >/ . ichthys maculatus]); at the type locality of Lori- p. 8 '/7. '/6 1/6- 68. Gron. mus. 1. n. I. 2. cariichthys maculatus Dr. Nijssen collected one f. 1.2. Plecostomus dorso ossiculo monopterygio, supe- male under the lower specimen carrying eggs lip. riori caudae bifurcae setiformi.

The in the buccal of the female i/ 6. C. two flaps cavity D. 8. P. '/7. V.

Gron. mus. 1. n. 69. Plecostomus ore edentulo, are larger than in the male and have distinct ƒ3- ossiculo superiori caudae bifurcae longitudine corporis. papillae. Habitat in America meridionali.

Colour (in alcohol). — Ground colour brownish Lobus superior caudae filo longissimo tcrminatur. There grey dorsally, dirty yellow ventrally. are three faint transverse bands on the dorsal side, viz.,

(1) at the seventh, (2) at the twelfth to fourteenth, This diagnosis consists of three parts. The first and the seventeenth lateral refers Linnaeus' (3) at to eighteenth part to own (1754) description body scute. A few dark brown irregularly and figure of “Loricaria dura the second to spots, ”, part

distributed and with and of on the body. Spines rays a Gronovius' (1754) description figures

series dark in the anal and the third Gro- regular of grey spots, except “Plecostomus no. 68", part to fin spine and rays. A large, faint brownish grey novius' (1754) description of “Plecostomus no. blotch in the terminal third of the lower caudal 69". Both “Loricaria dura” and “Plecostomus no. fin lobe with of the (fig. 2). Lips greyish pigmentation. 68" represent equivalent parts diagnosis

Traces of the original colour pattern are still pre- eligible to type selection, because Linnaeus referred

in the sent lectotype and paralectotype. “Plecostomus no. 69" to the /3-variety of Loricaria

cataphracta. “Loricaria dura” and “Plecostomus

68" differ whereas “Loricaria DISCUSSION no. at generic level,

dura”1 and “Plecostomus no. 69" in all probability

Bloch the 'first Nomenclatural remarks. The first account of the referred to the same species. was

this and species Loricariichthys maculatus was published revisor' of diagnosis (in a post-Linnaean

binominal and he restricted in 1794 the by Gronovius (1754 : 25—26, pi. 2 figs. I—2) work), under the description of “Plecostomus no. 68". name Loricaria cataphracta to the species Lin-

described “Loricaria dura”, whereas he The species was diagnosed as "PLECOSTOMUS naeus as

also described the Gronovius' dorso monopterygio, ore cirrato, edentulo, ossiculo same species as it Loricaria setiformi brevi." “Plecostomus no. 68", macu- superiori caudae bifurcae On page naming 26 Gronovius stated the locality: "Inhabitat Suri- lata.

Gronovius in retain- namae flumina". His fairly complete account con- (ed. Gray), 1854, however, 16 I. J. H. ISBRUCKER - LORICARIICHTHYS MACULATUS

Plecostomus 8 He named his ed the generic name ). of the existence of the name Parahemiodon typus,

“Plecostomus no. 68" Plecostomus cataphracta, a nominal species first described by Bleeker from

Plecostomus and his “Plecostomus no. 69" flagel- Surinam in 1862. The specific name typus (in laris. Although the origin of the specific name connection with either Parahemiodon, or Lorica-

Plecostomus is ob- has been cataphracta Gronovius, 1854, ria, or Loricariichthys) accepted as re-

I consider it Gronovian valid 1946. Van der vious, propose to a new presenting a species up to

known name instead of a quoted Linnaean (1758) name. Stigchel (1946 : 172—174) listed the single

Plecostomus is of Loricari- material of Loricaria cataphracta a synonym specimen together with his

maculatus, and not of Lori- 4 and stated 173—174: "In ichthys , a homonym maculata ) on pages the holo- caria cataphracta. Since the material on which the specimens no. 3121 [RMNH 3121,

based his is 10723 the lower Gronovius Plecostomus cataphracta no type of Parahemiodon typus] and

longer extant (cf. Wheeler, 1958), I propose, to lip is much larger than in the others. Bleeker has

nomenclatural the 3121 called further stability, to designate brought the specimen to a new species,

of maculata = differ- lectotype Loricari Bloch, 1794 ( Parahemiodon typus, but I think that the

the time the their fins Loricariichthys maculatus), at same ence is due to sexual variation; that are

of Plecostomus Gronovius, the same neotype cataphracta longer and broader has, in my opinion,

the In 10723 the first of the 1854, latter becoming an objective junior syno- reason. no. spine dorsal,

maculatus. One is inclined nym of Loricariichthys pectorals, ventrals and anal is somewhat prolonged

75b of the International Code also the to refer to article and stronger than in the other specimens;

(1964 : 81 : "CASES EXCLUDED. — A neotype ventral buckler is larger and heavier. In the speci- is not to be for a of which described Bleeker this was to designated ... species men by impossible

not either valid the name is in general use as a state, because the fins are badly damaged. I have.

it rele- maculata and name or as a synonym"). However, seems therefore, brought typus to the same

vant to designate a neotype for Plecostomus cata- species." It is dangerous to follow Van der Stig- of phracta Gronovius, 1854, because of its easy con- chel without having examined the holotype fusion with the Loricaria Lin- If should name cataphracta Parahemiodon typus. Regan (1904 : 287) 1758. of naeus, be followed in his synonymy Loricaria (Lori-

As the lectotype I chose the largest of the two cariichthys) typus (Bleeker, 1862), Loricaria va- still it is the stubelii extant syntypes, most probably speci- lenciennesi Vaillant, 1880, and Loricaria

Bloch. Dr. C. Karrer men figured by (in litt., May [more correctly spelled as L. stuebeli] Steindachner,

26th, 1970) stated: "Vielleicht nützt es Ihnen, zu 1882, and if Eigenmann and Eigenmann (1889:

dass Bloch L. maculata ..., in their of wissen, von ursprüng- 39) were right proposed synonymy lich besessen aber noch 4 Exemplare hat, jetzt nur Loricaria (Loricariichthys) konopickyi Steindach-

die beiden vorhanden sind." The smaller and Loricaria valenciennesi Vaillant, syntype, ner, 1879,

the is desig- refer to becoming paralectotype, consequently Lori-1880, all these specific names would nated the paraneotype of Plecostomus cataphracta cariichthys maculatus (Bloch, 1794), when the sy-

Gronovius, 1854. Although rarely be never) of Loricaria maculata and Parahemiodon (may nonymy terms like would paraneotype (and alloneotype) are ap- typus, as proposed by Van der Stigchel, plied in systematic works, the accepted use of the be included 5 ). However, I have examined the ho- terms holotype, allotype, and paratype on the one lotype of Loricaria valenciennesi (MNHN A. hand and the other and lectotype paralectotype on 1985), and this species certainly is not identical to hand, induce me to introduce equivalent terms for Loricariichthys maculatus (Bloch, 1794) : it nei- the of of Lori- neotype series, too. ther represents a species Loricaria, nor

cariichthys. A re-examination of all the type ma- COMMENTS ON THE SYNONYMY OF terial of the species of the Loricariinae will be LORICARIICHTHYS MACULATUS necessary before any conclusion on the synonymy

be drawn. For the reason the references I should like make remarks can same to some on the propos- listed above are nothing but primary. ed synonymy of Loricariichthys maculatus, because

3 At of 4 also recorded from the ) present a synonym Hypostomus Lacépède, 1803, ) Van der Stigchel a specimen type species, by original monotypy, Hypostomus guacari "Botanical Garden, Paramaribo", the type locality. been Lacépède, 1803, to be identified with Acipenser plecosto- ■'•) Loricaria amazonica de Castelnau, 1855, also has

1758 considered of maculatus by mus Linnaeus, = Hypostomus plecostomus (Lin- a synonym Loricariichthys naeus, 1758); fide Boeseman, 1968. almost all authors. BUDRAGEN TOT DE DIERKUNDE, 41 (1) - 1971 17

ACKNOWLEDGEMENTS

would like thanks the der Laan 1 to express my to following per- information; to Mr. A. L. van (Instituut voor

who in the of this Taxonomische Amster- sons, were helpful preparation paper: Zoölogie ("Zoölogisch Museum"),

to Mrs. Dr. M. L. Bauchot (Muséum National d'Histoire dam, ZMA) for making the photographs and reproduc-

Naturelle, Paris, MNHN) for the loan of some loricariid tions; to Dr. H. Nijssen (also ZMA) for his criticism

fishes, including the holotype of Loricaria valenciennesi and advice; and to Mrs. Drs. F. F. J. M. Peeters-Pieters

Vaillant, 1880; to Mr. M. C. Burns (Cheltenham, Eng- (Artis Bibliotheek, Universiteit van Amsterdam) for trac-

for the Prof. Dr. K. the valuable in her care. The land) kindly correcting English; to ing many publications and of Ichthyology of the Deckert Dr. C. Karrer (Museum für Naturkunde study was done at the Department

der Humboldt-Univershat, Berlin, ZMB) for the loan Institute of Taxonomie Zoology ("Zoölogisch Museum"),

of the two syntypes of Loricaria maculata Bloch, 1794, Amsterdam; fig. 7 was made by the author.

for the permission to designate the lcctotype, and for

LITERATURE

Bleeker, P., 1862a. Atlas ichthyologique des Indes Orien- Gronovius, L. T. (J. E. Gray, editor), 1854. Catalogue of

tales Néerlandaises: Siluroïdes, Chacoïdes et Hétéro- fish collected and described by Laurence Theodore

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Received: 9 October 1970