A peer-reviewed open-access journal ZooKeys 443: 61–118 (2014)Theintegripennis species group of Geocharidius Jeannel, 1963... 61 doi: 10.3897/zookeys.443.7880 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research The integripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation Igor M. Sokolov1, David H. Kavanaugh1 1 Department of Entomology, California Academy of Sciences, Golden Gate Park, 55 Music Concourse Drive, San Francisco, CA 94118, USA Corresponding author: David H. Kavanaugh (
[email protected]) Academic editor: Terry Erwin | Received 13 May 2014 | Accepted 14 August 2014 | Published 30 September 2014 http://zoobank.org/E3384139-6A6E-426C-840D-85BC32A12E78 Citation: Sokolov IM, Kavanaugh DH (2014) Theintegripennis species group of Geocharidius Jeannel, 1963 (Carabidae, Bembidiini, Anillina) from Nuclear Central America: a taxonomic review with notes about biogeography and speciation. ZooKeys 443: 61–118. doi: 10.3897/zookeys.443.7880 Abstract Our review recognizes 15 species of the integripennis species group of Geocharidius from Nuclear Central America, include three species previously described (G. gimlii Erwin, G. integripennis (Bates) and G. zul- linii Vigna Taglianti) and 12 described here as new. They are:G. andersoni sp. n. (type locality: Chiapas, Chiapas Highlands, Cerro Huitepec) and G. vignatagliantii sp. n. (type locality: Chiapas, Motozintla, Sierra Madre de Chiapas, Benito Juárez) from Mexico; G. antigua sp. n. (type locality: Sacatepéquez, 5 km SE of Antigua), G. balini sp. n. (type locality: Suchitepéquez, 4 km S of Volcan Atitlán), G. erwini sp. n. (type locality: Quiché Department, 7 km NE of Los Encuentros), G.