EARTH HISTORY UNIT 5 Mesozoic & Cenozoic Eras
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The triple origin of whales DAVID PETERS Independent Researcher 311 Collinsville Avenue, Collinsville, IL 62234, USA [email protected] 314-323-7776 July 13, 2018 RH: PETERS—TRIPLE ORIGIN OF WHALES Keywords: Cetacea, Mysticeti, Odontoceti, Phylogenetic analyis ABSTRACT—Workers presume the traditional whale clade, Cetacea, is monophyletic when they support a hypothesis of relationships for baleen whales (Mysticeti) rooted on stem members of the toothed whale clade (Odontoceti). Here a wider gamut phylogenetic analysis recovers Archaeoceti + Odontoceti far apart from Mysticeti and right whales apart from other mysticetes. The three whale clades had semi-aquatic ancestors with four limbs. The clade Odontoceti arises from a lineage that includes archaeocetids, pakicetids, tenrecs, elephant shrews and anagalids: all predators. The clade Mysticeti arises from a lineage that includes desmostylians, anthracobunids, cambaytheres, hippos and mesonychids: none predators. Right whales are derived from a sister to Desmostylus. Other mysticetes arise from a sister to the RBCM specimen attributed to Behemotops. Basal mysticetes include Caperea (for right whales) and Miocaperea (for all other mysticetes). Cetotheres are not related to aetiocetids. Whales and hippos are not related to artiodactyls. Rather the artiodactyl-type ankle found in basal archaeocetes is also found in the tenrec/odontocete clade. Former mesonychids, Sinonyx and Andrewsarchus, nest close to tenrecs. These are novel observations and hypotheses of mammal interrelationships based on morphology and a wide gamut taxon list that includes relevant taxa that prior studies ignored. Here some taxa are tested together for the first time, so they nest together for the first time. INTRODUCTION Marx and Fordyce (2015) reported the genesis of the baleen whale clade (Mysticeti) extended back to Zygorhiza, Physeter and other toothed whales (Archaeoceti + Odontoceti). -
Thewissen Et Al. Reply Replying To: J
NATURE | Vol 458 | 19 March 2009 BRIEF COMMUNICATIONS ARISING Hippopotamus and whale phylogeny Arising from: J. G. M. Thewissen, L. N. Cooper, M. T. Clementz, S. Bajpai & B. N. Tiwari Nature 450, 1190–1194 (2007) Thewissen etal.1 describe new fossils from India that apparentlysupport fossils, Raoellidae or the raoellid Indohyus is more closely related to a phylogeny that places Cetacea (that is, whales, dolphins, porpoises) as Cetacea than is Hippopotamidae (Fig. 1). Hippopotamidae is the the sister group to the extinct family Raoellidae, and Hippopotamidae exclusive sister group to Cetacea plus Raoellidae in the analysis that as more closely related to pigs and peccaries (that is, Suina) than to down-weights homoplastic characters, althoughin the equallyweighted cetaceans. However, our reanalysis of a modified version of the data set analysis, another topology was equally parsimonious. In that topology, they used2 differs in retaining molecular characters and demonstrates Hippopotamidae moved one node out, being the sister group to an that Hippopotamidae is the closest extant family to Cetacea and that Andrewsarchus, Raoellidae and Cetacea clade. In neither analysis is raoellids are the closest extinct group, consistent with previous phylo- Hippopotamidae closer to the pigs and peccaries than to Cetacea, the genetic studies2,3. This topology supports the view that the aquatic result obtained by Thewissen et al.1. In all our analyses, pachyostosis adaptations in hippopotamids and cetaceans are inherited from their (thickening) of limb bones and bottom walking, which occur in hippo- common ancestor4. potamids9,10, are interpreted to have evolved before the pachyostosis of To conduct our analyses, we started with the same published matrix the auditory bulla, as seen in raoellids and cetaceans1. -
Craniofacial Morphology of Simosuchus Clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar
Society of Vertebrate Paleontology Memoir 10 Journal of Vertebrate Paleontology Volume 30, Supplement to Number 6: 13–98, November 2010 © 2010 by the Society of Vertebrate Paleontology CRANIOFACIAL MORPHOLOGY OF SIMOSUCHUS CLARKI (CROCODYLIFORMES: NOTOSUCHIA) FROM THE LATE CRETACEOUS OF MADAGASCAR NATHAN J. KLEY,*,1 JOSEPH J. W. SERTICH,1 ALAN H. TURNER,1 DAVID W. KRAUSE,1 PATRICK M. O’CONNOR,2 and JUSTIN A. GEORGI3 1Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, 11794-8081, U.S.A., [email protected]; [email protected]; [email protected]; [email protected]; 2Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio 45701, U.S.A., [email protected]; 3Department of Anatomy, Arizona College of Osteopathic Medicine, Midwestern University, Glendale, Arizona 85308, U.S.A., [email protected] ABSTRACT—Simosuchus clarki is a small, pug-nosed notosuchian crocodyliform from the Late Cretaceous of Madagascar. Originally described on the basis of a single specimen including a remarkably complete and well-preserved skull and lower jaw, S. clarki is now known from five additional specimens that preserve portions of the craniofacial skeleton. Collectively, these six specimens represent all elements of the head skeleton except the stapedes, thus making the craniofacial skeleton of S. clarki one of the best and most completely preserved among all known basal mesoeucrocodylians. In this report, we provide a detailed description of the entire head skeleton of S. clarki, including a portion of the hyobranchial apparatus. The two most complete and well-preserved specimens differ substantially in several size and shape variables (e.g., projections, angulations, and areas of ornamentation), suggestive of sexual dimorphism. -
Ceratophrys Cranwelli) with Implications for Extinct Giant Frogs Scientific Reports, 2017; 7(1):11963-1-11963-10
PUBLISHED VERSION A. Kristopher Lappin, Sean C. Wilcox, David J. Moriarty, Stephanie A.R. Stoeppler, Susan E. Evans, Marc E.H. Jones Bite force in the horned frog (Ceratophrys cranwelli) with implications for extinct giant frogs Scientific Reports, 2017; 7(1):11963-1-11963-10 © The Author(s) 2017 Open Access This article is licensed under a Creative Commons Attribution 4.0 International License, which permits use, sharing, adaptation, distribution and reproduction in any medium or format, as long as you give appropriate credit to the original author(s) and the source, provide a link to the Creative Commons license, and indicate if changes were made. The images or other third party material in this article are included in the article’s Creative Commons license, unless indicated otherwise in a credit line to the material. If material is not included in the article’s Creative Commons license and your intended use is not permitted by statutory regulation or exceeds the permitted use, you will need to obtain permission directly from the copyright holder. To view a copy of this license, visit http://creativecommons.org/licenses/by/4.0/. Originally published at: http://doi.org/10.1038/s41598-017-11968-6 PERMISSIONS http://creativecommons.org/licenses/by/4.0/ 19th of April 2018 http://hdl.handle.net/2440/110874 www.nature.com/scientificreports OPEN Bite force in the horned frog (Ceratophrys cranwelli) with implications for extinct giant frogs Received: 27 March 2017 A. Kristopher Lappin1, Sean C. Wilcox1,2, David J. Moriarty1, Stephanie A. R. Stoeppler1, Accepted: 1 September 2017 Susan E. -
Member Magazine Summer 2013 Vol. 38 No. 3 2 News at the Museum 3
Member Magazine Summer 2013 Vol. 38 No. 3 2 News at the Museum 3 From the For many people, summer means vacations cultural points of view, presenting the roles, both Ancient Mariners: New Initiative Makes and precious time away from professional positive and negative, that poison has played and President responsibilities. At the Museum, summer continues to play in nature and human history. At Rare Display of Trilobites Membership Card Count is a time of tremendous activity! the same time, scientists, computer engineers, and Ellen V. Futter Field season kicks into high gear and many visualization experts in the Rose Center for Earth The Museum’s Grand Gallery recently became home to a remarkable case of of our scientists head to locations around the and Space are putting the finishing touches on trilobites. Called “butterflies of the sea,” “frozen locusts,” or simply “bugs” by world to further their research. Of course, field a thrilling new Hayden Planetarium Space Show, the researchers who study them, these ancient arthropods are distant relatives expeditions are a long-standing tradition at the which will reveal dramatic recent discoveries in of modern lobsters, horseshoe crabs, and spiders. Museum—part of our DNA, if you will. Museum the fast-moving field of astrophysics and forecast The temporary exhibition, overseen by Neil Landman, curator in the scientists now undertake some 120 expeditions what’s in store in cosmology in the years ahead. Museum’s Division of Paleontology, includes 15 fossils of various trilobite species each year, and that work advances not only The Space Show will premiere on October 5 and from the Museum’s permanent collection. -
History of Earth
History of Earth The history of Earth concerns the development of planet Earth from its formation to the present day.[1][2] Nearly all branches of natural science have contributed to understanding of the main events of Earth's past, characterized by constant geological change and biological evolution. The geological time scale (GTS), as defined by international convention,[3] depicts the large spans of time from the beginning of the Earth to the present, and its divisions chronicle some definitive events of Earth history. (In the graphic: Ga means "billion years ago"; Ma, "million years ago".) Earth formed around 4.54 billion years ago, approximately one-third the age of the universe, by accretion from the solar nebula.[4][5][6] Volcanic outgassing probably created the primordial atmosphere and then the ocean, but the early atmosphere contained almost no oxygen. Much of the Earth was molten because of frequent collisions with other bodies which led to extreme volcanism. While the Earth was in its earliest stage (Early Earth), a giant impact collision with a planet-sized body named Theia is thought to have formed the Moon. Over time, the Earth cooled, causing the formation of a solid crust, and allowing liquid water on the surface. The Hadean eon represents the time before a reliable (fossil) record of life; it began with the formation of the planet and ended 4.0 billion years ago. The following Archean and Proterozoic eons produced the beginnings of life on Earth and its earliest evolution. The succeeding eon is the Phanerozoic, divided into three eras: the Palaeozoic, an era of arthropods, fishes, and the first life on land; the Mesozoic, which spanned the rise, reign, and climactic extinction of the non-avian dinosaurs; and the Cenozoic, which saw the rise of mammals. -
Resolving the Relationships of Paleocene Placental Mammals
Biol. Rev. (2015), pp. 000–000. 1 doi: 10.1111/brv.12242 Resolving the relationships of Paleocene placental mammals Thomas J. D. Halliday1,2,∗, Paul Upchurch1 and Anjali Goswami1,2 1Department of Earth Sciences, University College London, Gower Street, London WC1E 6BT, U.K. 2Department of Genetics, Evolution and Environment, University College London, Gower Street, London WC1E 6BT, U.K. ABSTRACT The ‘Age of Mammals’ began in the Paleocene epoch, the 10 million year interval immediately following the Cretaceous–Palaeogene mass extinction. The apparently rapid shift in mammalian ecomorphs from small, largely insectivorous forms to many small-to-large-bodied, diverse taxa has driven a hypothesis that the end-Cretaceous heralded an adaptive radiation in placental mammal evolution. However, the affinities of most Paleocene mammals have remained unresolved, despite significant advances in understanding the relationships of the extant orders, hindering efforts to reconstruct robustly the origin and early evolution of placental mammals. Here we present the largest cladistic analysis of Paleocene placentals to date, from a data matrix including 177 taxa (130 of which are Palaeogene) and 680 morphological characters. We improve the resolution of the relationships of several enigmatic Paleocene clades, including families of ‘condylarths’. Protungulatum is resolved as a stem eutherian, meaning that no crown-placental mammal unambiguously pre-dates the Cretaceous–Palaeogene boundary. Our results support an Atlantogenata–Boreoeutheria split at the root of crown Placentalia, the presence of phenacodontids as closest relatives of Perissodactyla, the validity of Euungulata, and the placement of Arctocyonidae close to Carnivora. Periptychidae and Pantodonta are resolved as sister taxa, Leptictida and Cimolestidae are found to be stem eutherians, and Hyopsodontidae is highly polyphyletic. -
The Biology of Marine Mammals
Romero, A. 2009. The Biology of Marine Mammals. The Biology of Marine Mammals Aldemaro Romero, Ph.D. Arkansas State University Jonesboro, AR 2009 2 INTRODUCTION Dear students, 3 Chapter 1 Introduction to Marine Mammals 1.1. Overture Humans have always been fascinated with marine mammals. These creatures have been the basis of mythical tales since Antiquity. For centuries naturalists classified them as fish. Today they are symbols of the environmental movement as well as the source of heated controversies: whether we are dealing with the clubbing pub seals in the Arctic or whaling by industrialized nations, marine mammals continue to be a hot issue in science, politics, economics, and ethics. But if we want to better understand these issues, we need to learn more about marine mammal biology. The problem is that, despite increased research efforts, only in the last two decades we have made significant progress in learning about these creatures. And yet, that knowledge is largely limited to a handful of species because they are either relatively easy to observe in nature or because they can be studied in captivity. Still, because of television documentaries, ‘coffee-table’ books, displays in many aquaria around the world, and a growing whale and dolphin watching industry, people believe that they have a certain familiarity with many species of marine mammals (for more on the relationship between humans and marine mammals such as whales, see Ellis 1991, Forestell 2002). As late as 2002, a new species of beaked whale was being reported (Delbout et al. 2002), in 2003 a new species of baleen whale was described (Wada et al. -
Whales: Giants of the Deep March 19, 2016 Through September 5, 2016
Whales: Giants of the Deep March 19, 2016 through September 5, 2016 Contents Welcome ....................................................................................................................................................... 1 Volunteer Logistics ........................................................................................................................................ 1 Reporting for Service ................................................................................................................................ 1 Scheduling ................................................................................................................................................. 1 Logistics for Interpretative Cart ................................................................................................................ 1 Representing the Museum ....................................................................................................................... 2 Logging Your Volunteer Hours .................................................................................................................. 2 Adding Yourself to the Schedule ............................................................................................................... 3 Introduction to Cetaceans ............................................................................................................................ 3 Classification of Cetaceans ......................................................................... Error! Bookmark -
Zofia Kielan-Jaworowska and the Gobi Palaeontological Expeditions
ZOFIA KIELAN-JAWOROWSKA AND THE GOBI PALAEONTOLOGICAL EXPEDITIONS JOHN R. LAVAS During an illustrious scientific career that lasted more than 60 years, the late Zofia Kielan-Jaworowska greatly advanced the direction of research in several branches of invertebrate and vertebrate palaeontology. The general public largely knew Zofia as the leader of the famous joint Polish-Mongolian Gobi Expeditions of the 1960s–1970s and the spectacular (mainly dinosaurian) fossils that resulted from those ventures. Indeed, an enduring image that I associate with her name shows Zofia standing beneath one of her discov- eries, the enigmatic theropod Deinocheirus. Momentous as such finds were, Zofia’s principal research area was the phylogenetic relationships of the Mesozoic mammals retrieved during those expeditions, specifi- cally evidence from their brain structure and musculature. It is within this realm that she left her greatest scientific legacy in what had previously been a relatively poorly-developed branch of palaeontology due to the paucity and fragility of fossil material. As well as attaining the forefront of her chosen field, Zofia was a humanist with a unique ability to forge successful collaborations between researchers from the former Council for Mutual Economic Assistance (COMECON) countries and their Western counterparts at a time in history when it was difficult to do so. The inclination to rank Zofia alongside her compatriot Marie Curie (1867–1934) is not only appropriate in terms of the impact each made to science, but the fact that Zofia’s late husband was a radiobiologist and that they had spent two years working in Paris is surely more than coinci- dental. -
SCIENCE CHINA Early Paleogene Stratigraphic Sequences
SCIENCE CHINA Earth Sciences • RESEARCH PAPER • December 2010 Vol.53 No.12: 1918–1926 doi: 10.1007/s11430-010-4095-8 Early Paleogene stratigraphic sequences, mammalian evolution and its response to environmental changes in Erlian Basin, Inner Mongolia, China WANG YuanQing1*, MENG Jin2, Christopher K. BEARD3, LI Qian1, NI XiJun1, Daniel L. GEBO4, BAI Bin1, JIN Xun1 & LI Ping1 1 Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China; 2 Division of Paleontology, American Museum of Natural History, New York NY 10024, USA; 3 Section of Vertebrate Paleontology, Carnegie Museum of Natural History, 4400 Forbes Avenue, Pittsburgh, PA 15213, USA; 4 Department of Anthropology, Northern Illinois University, DeKalb, IL 60115, USA Received June 11, 2010; accepted September 2, 2010; published online November 5, 2010 Paleogene strata in the Huheboerhe area, Erlian Basin, Inner Mongolia, China have been subdivided into three lithological units: the Nomogen Formation, the Arshanto Formation, and the Irdin Manha Formation. At least 12 mammal-bearing horizons have been recognized in these formations, of which 4 in the Nomogen Formation, 6 in the Arshanto Formation, and 2 in the Irdin Manha Formation. Recent investigation proved that the “Houldjin Formation” recognized in this area by the Central Asi- atic Expeditions (CAE) of the American Museum Natural History in the 1920s is actually the Irdin Manha Formation, while the “Irdin Manha Formation” of CAE is the Arshanto Formation. A recent paleomagnetic study suggests that the upper part of the Nomogen Formation is early Eocene in age and the Arshanto Formation is mainly early Eocene rather than middle Eocene as previously thought. -
Book of Barely Imagined for B 11/01/2013 10:17 Page I
Book of Barely Imagined for B 11/01/2013 10:17 Page i THE BOOK of BARELY IMAGINED BEINGS As centuries pass by, the mass of works grows endlessly, and one can foresee a time when it will be almost as difficult to educate oneself in a library, as in the universe, and almost as fast to seek a truth subsisting in nature, as lost among an immense number of books. Denis Diderot, Encyclopédie, 1755 In our gradually shrinking world, everyone is in need of all the others. We must look for man wherever we can find him. When on his way to Thebes Oedipus encountered the Sphinx, his answer to its riddle was: ‘Man’. That simple word destroyed the monster. We have many monsters to destroy. Let us think of the answer of Oedipus. George Seferis, Nobel Prize speech, 1963 Book of Barely Imagined for B 11/01/2013 10:17 Page ii Book of Barely Imagined for B 11/01/2013 10:17 Page iii THE BOOK of BARELY IMAGINED BEINGS CASPAR HENDERSON Illustrated by GOLBANOU MOGHADDAS Book of Barely Imagined for B 11/01/2013 10:17 Page iv Granta Publications, 12 Addison Avenue, London W11 4QR First published in Great Britain by Granta Books, 2012 Copyright © Caspar Henderson, 2012 Caspar Henderson has asserted his moral right under the Copyright, Designs and Patents Act, 1988, to be identified as the author of this work. The picture credits on pp. 414–5 and the text credits on pp. 416–7 constitute extensions of this copyright page. All rights reserved.