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Home , Bassaricyon, Black myotis, Bush dog, Common opossum, Galictis, Oncilla

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JAGUAR onca Size: 70 kg. Largest in Iwokrama. Nocturnal and diurnal; climbs low trees and swims well. Solitary. Preys on large such as , and Deer. Occasionally roars or makes a loud series of grunts. Front Hind

PUMA concolor Size: 45 kg. Only large unspotted in Iwokrama. Climbs well. Solitary. Prey includes Deer, , and . Large tracks (about 80 mm across), often found on dirt roads. Other signs includes partially Front Hind eaten kills covered with sticks.

JAGUARUNDI Puma jagouaroundi Size: 7 kg. Can be dark grey (more common) or reddish. Only small, unspotted cat in Iwokrama. Long narrow tail distinguishes it from bushy-tailed . Diurnal. Climbs well. Eats small and birds. Front Hind 1 CARNIVORES CARNIVORES

OCELOT OLINGO pardalis alleni Size: 10 kg. Medium-sized spotted Size: 1.5 kg. Small and catlike, cat. Relatively narrow tail is only with a long, slightly bushy tail. as long as the hind legs. Mainly is larger and has a nocturnal. Eats Iguanas, small tapering, prehensile tail and a terrestrial, , land crabs, broader muzzle. Agile and fast-moving. and birds. Front tracks are noticeably Front Hind Nocturnal and arboreal, seldom descends to the ground. broader than hind tracks. Eats fruit, nectar, invertebrates and small vertebrates. Leopardus tigrinus KINKAJOU Potos flavus Size: 2.25 kg. Smallest spotted cat, about the size of a house Size: 3 kg. Most commonly seen cat. Probably nocturnal. Eats nocturnal, arboreal in mice and small birds. Iwokrama. The Olingo is similar but has a grey head. Eats fruit, nectar, insects, mice and . Calls include Front Hind a short, barking “wick-wick-wick”. Listen for sounds of falling leaves as it Front Hind moves noisily through the trees. Leopardus weidii Size: 3.5 kg. Small, slim, spotted cat. Bushy nasua tail is longer than legs. Mainly Size: 4.5 kg. A long-nosed relative of the nocturnal; hunts in trees. It can with a long tail often held straight up. Diurnal, Front Hind rotate the hind feet and terrestrial and arboreal. Females and descend head first down a young live in large groups, males solitary vertical tree trunk. Eats climbing outside breeding season. Eats fruit, worms, mice, , and squirrels. Front Hind insects and small rodents. 2 3 CARNIVORES CARNIVORES

TAYRA RIVER Eira barbara longicaudis Size 4 kg. Large, long-legged Size: 6.5 kg. Distinctive low, -like with a long streamlined shape. Fast and agile bushy tail. Travels with a bouncy gait, swimmer, but moves with an back and tail arched. Mainly diurnal, awkward, humping gait on land. semi-arboreal. Eats insects, fruit, Always found in or near water. and vertebrates including lizards, monkeys, Tracks with distinctive webbing between Front and . Look for seed-filled scat left on toes, tail leaves drag marks between tracks. Hind fallen logs. Front Hind GRISON Pteronura brasiliensis vittata Size: 30 kg. Large, chocolate brown Size: 2 kg. Muscular, with a long, with white throat. Diurnal. Lives in family low body and very short legs and tail. groups. Eats fish. When disturbed in the Probably diurnal; terrestrial, swims water, the will crane their heads up well. Eats small reptiles, birds, and and snort sharply. Look for on mammals. Tracks are five-toed, with banks with conspicuous trampled areas prominent claw marks. Rare. and slides into the water. Front Hind

BUSH DID YOU KNOW? venaticus The Iwokrama International Centre is a key environmental programme Size: 6 kg. An odd-looking dog, dedicated by the Government of to the Commonwealth and the small and low-slung. Diurnal. world at the 1989 Commonwealth Heads of Government Meeting. The Rarely seen. Prefers swampy Iwokrama Forest was established as a “living laboratory” for tropical areas. Eats Agouti, Paca and fish. forest management – to show how tropical forests can be conserved Besides obvious claw marks, and sustainably used for ecological, social and economic benefits to foot pad relatively small local, national and international communities. Front compared to cat tracks of similar size. Hind 4 5 PRIMATES PRIMATES

SQUIRREL MONKEY GUIANAN SAKI MONKEY Saimiri sciureus Pithecia pithecia Size: 800 g. Small, slender monkey Size: 2 kg. Sexes distinct. Male entirely with striking colouration and short black with white face, female grey. Sits . This active monkey is mainly in a hunched crouch often with bushy tail insectivorous. Large groups are easily dangling straight down. Usually seen in seen darting about in trees and bushes, the subcanopy or lower. This quiet, shy busily tearing off bark, looking under leaves, monkey feeds on a variety of fruits, and moving through branches in search of insects. seeds, leaves, and insects.

WEDGE-CAPPED RED HOWLER MONKEY Alouatta macconnelli Cebus olivaceus Size: 7 kg. Chin with forward-growing beard Size: 3 kg. Top of head has a distinct (longest in males). Diurnal; strictly arboreal. wedge-shaped blackish cap tapering Social. Lives in troops. The loud calls consist to a very thin stripe down the forehead of a rising chorus of snorts, grunts, roars, to the nose. Usually seen in the subcanopy and howls which can be heard for several or lower. Social, typically lives in groups of kilometres. 10 to 40 individuals.

GOLDEN-HANDED TAMARIN BLACK SPIDER MONKEY Ateles paniscus Saguinus midas Size: 10 kg. Largest monkey in Guyana. Size: 450 g. Smallest primate in Guyana. Hands When seen sitting in a tree, this spider and feet are bright orangish-yellow. Diurnal, monkey has a distinctly ape-like arboreal. Often seen holding onto tree trunks, appearance. Social, usually seen in groups while it nervously peers and quickly moves its of 2 to 5 individuals. Travels noisily through the head about. Commonly heard before it is seen, trees, often stopping to investigate and threaten it makes a high-pitched whistle. observers. 6 7 HOOFED MAMMALS HOOFED MAMMALS

BRAZILIAN RED BROCKET DEER Tapirus terrestris Mazama americana Size: 250 kg. The largest mammal in Size: 22 kg. Medium-sized deer with Iwokrama. Nocturnal and diurnal. distinctive hunched posture with the Solitary. Call is a loud, long whistle. rump higher than the shoulders or Ranges widely and can be seen head. Ears and legs relatively short far from water. Distinctive tracks seen compared to the Grey Brocket. Most more often than the animal. Front Hind often seen in morning, at dusk or at night while foraging. Distinctive split hoof prints tapering smoothly to Front or Hind WHITE-LIPPED narrow tips. Tayassu pecari Size: 35 kg. Larger and darker than GREY BROCKET DEER Collared Peccary. Diurnal. Lives in Mazama nemorivaga large herds. Feeds by bulldozing Size: 15 kg. Ears and legs relatively long through soil and eating fruit, roots, compared to Red Brocket Deer. Usually vegetation and invertebrates. Cracks solitary. Feeds primarily on fruit. Split open very hard palm nuts. Front or Hind hoof prints similar to, but slightly smaller than, Red Brocket Deer. Front or Hind

COLLARED PECCARY DID YOU KNOW? Pecari tajacu The Iwokrama Forest is located in central Guyana, approximately 300 Size: 20 kg. Small, piglike, with a km south of Georgetown, the capital. The area encompasses about stocky body and slim legs. Social, 371,000 hectares and is covered in lush, intact lowland tropical forest. usually seen in groups of up to 15. The wide range of intact habitats in the Iwokrama Forest supports a Groups rest in caves, in deep diverse flora and fauna with an estimated 1,500-2,000 higher plant burrows, or under rocks or logs. Eats mostly palm , 420 species of fish, 150 species of snakes, lizards and frogs, nuts. More common than White-lipped Peccary. 500 species of birds and 180 species of mammals. Front Hind

8 9 ANTEATERS SLOTHS

GIANT ANTEATER THREE-TOED SLOTH Myrmecophaga tridactyla Bradypus tridactylus Size: 30 kg. Large, appears Size: 6kg. Hangs below branch. Male “too long” for its height. Diurnal has a patch of short orange fur marked or nocturnal. Uses its powerful with black spots on the middle of its claws to open large termite mounds or terrestrial back. Arboreal, slow-moving. Solitary. ant nests. Odd-looking tracks show front claws Extremely docile. Eats leaves, especially pointing backward or laterally (it walks on the Congo palm. Look for a mottled knuckles with the claws turned under). grey ball on or under a branch.

TAMANDUA TWO-TOED SLOTH Tamandua tetradactyla Choleopus didactylus Size: 6 kg. Medium-sized, with Size: 6 kg. Hangs below branch. a long prehensile tail. Nocturnal or Fur long and shaggy, brownish, darker diurnal; semi-arboreal. Solitary. It on limbs than body. Mainly nocturnal; feeds on ants, termites and bees. arboreal, usually in the canopy. Solitary. Listen for sounds of tearing wood as Eats leaves of a variety of trees, Front it attacks arboreal termite nests. also fruit and buds. Look for a brown Hind ball on a tree crotch. SILKY ANTEATER Cyclopes didactylus Size: 20 kg. Very small, with a VISIT THE IWOKRAMA FOREST! furry prehensile tail. Nocturnal, arboreal. Solitary. It travels on Scientists, students, tourists, volunteers and interns are pencil-thin vines, using the large encouraged to visit the Iwokrama Forest. Accommodation, front claws to open hollow stems in search of transportation and research facilities are provided at the ants, its preferred food. Look for a furry golden ball Iwokrama River Lodge and Research Centre. in a vine tangle.

10 11 OPOSSUMS

GIANT GREY FOUR-EYED Priodontes maximus OPPOSSUM Philander Size: 30 kg. Very large. Massive claws on front foot. Nocturnal; Size: 750 g. Greyish above, terrestrial and subterranean. cream-yellow below. Tail blackish Solitary. Feeds by digging into the with a contrasting white tip. Nocturnal, nests of ants and termites. Huge terrestrial and arboreal. Commonly seen foraging has a semicircular entrance about 45 cm wide. on ground along stream banks. Opposable big toe on Front Hind the hind foot is clearly evident. GREAT LONG-NOSED COMMON OPOSSUM ARMADILLO marsupialis Dasypus kappleri Size: 1.5 kg. Largest opossum. Size 10 kg. The base of the Nocturnal, terrestrial, semi-arbo- long, armoured tail is thick real. Solitary, omnivorous. Raids and noticeably flattened, chicken coops and garbage dumps. especially in adults. Nocturnal, This opossum is often smelled terrestrial. Solitary. Burrow has a before it is seen (hence the local Front name “stink opossum”). Hind smooth dome-shaped entrance Front Hind and is usually located near streams. Chironectes minimus VIEWING TIPS! Size: 675 g. Strikingly marked Most mammals are secretive and can be hard to see. Since many opossum. Hind feet conspicuously mammals are nocturnal, a good way to see them is at night with webbed. Nocturnal, terrestrial, the help of a headlamp. Fruiting trees are also a good place to see and semi-aquatic. Solitary. mammals as they congregate to feed. And always keep an eye on the ground for signs - especially tracks in the wet mud on the edge of Usually seen searching for food in or pools. near small, rocky streams. Feeds on fish, Front Hind crustaceans, and other invertebrates.

12 13 OPOSSUMS OPOSSUMS

MURINE MOUSE DELICATE MOUSE OPOSSUM OPPOSSUM murina parvidens Size: 40 g. Small sized. Female does Size: 20 g. Tiny. The arms and not have a pouch. Nocturnal, arboreal and legs are noticeably long, thin and terrestrial. Solitary, it mainly eats insects. delicate. Female does not have a Sometimes enters houses and buildings at pouch. Nocturnal, arboreal and terrestrial. the edge of forest. Solitary, eats insects and fruit. Slow moving and deliberate. WOOLLY MOUSE OPOSSUM SHORT-TAILED OPOSSUM Marmosa demerarae Monodelphis arlindoi Size: 100 g. Largest mouse Size: 100 g. Only small terrestrial opossum. Female does not have opossum in Guyana with a short a pouch. Nocturnal, mainly tail and dark gray back contrasting arboreal in understory or sub with red sides. Female does not canopy. Usually solitary, eats have a pouch. Diurnal. Solitary, insects, small vertebrates and fruit. insectivorous. Usually found in upland forest away from streams.

DID YOU KNOW? DID YOU KNOW? The Iwokrama Forest and the neighbouring North Rupununi wetlands The Iwokrama Forest is spatially zoned into two distinct areas – the is home to healthy populations of some of the world’s largest and Sustainable Utilisation Area (SUA) and the Wilderness Preserve (WP). most endangered species – the “Giants of El Dorado” including the The WP has been set aside as a biodiversity reserve, where all activity , the , the Giant Anteater, the Giant River Otter, the is severely restricted. The SUA is available for multiple uses including Arapaima (the world’s largest scaled fish), the Anaconda, the Black Iwokrama’s sustainable businesses: Timber Harvesting, Tourism and Caiman, the Giant River Turtle and the Bushmaster snake. Training and Learning Services.

14 15 LARGE RODENTS LARGE RODENTS

PACA CAPYBARA Cuniculus paca Hydrochoerus hydrochaeris Size: 8 kg. Stripes of white spots Size: 30 kg. World’s largest from shoulder to rump on each . Large, webbed feet. side. Terrestrial, nocturnal, it Occurs in small family groups of is often active after midnight, 2 to 6 individuals. It is usually seen especially in areas where it is at river’s edge or swimming. frequently hunted (it is prized for Look for piles of smooth, oval, its tasty flesh). Front greenish-brown droppings (about 50 Hind Front Hind ACOUCHI mm in length) at water’s edge. Myoprocta acouchy PREHENSILE-TAILED PORCUPINE Size: 1.5 kg. Similar to, but Coendou prehensilis smaller than, an agouti, with relatively long ears and legs. Diurnal (most active Size: 2 kg. Covered with rounded, stiff, in early morning or late afternoon). When barbed quills with pale yellowish alarmed, it bounds away quickly and then tips. Nocturnal (sometimes active in late freezes. Calls include chirps, whistles, and afternoon), arboreal. Typically slow and squawks, and it often drums the feet in alarm. deliberate, but can move quickly. During the Front day it rests in hollow tree dens or on branches. Hind It has a characteristic, strong odour. RED-RUMPED AGOUTI Dasyprocta leporina Size: 3.5 kg. Looks like a short-eared DID YOU KNOW? rabbit on stilts. Diurnal (most active The Iwokrama Forest is in the homeland of the Makushi people, who in early morning or late afternoon). have lived in and used the forest for thousands of years. People are a Easiest to observe at dawn or critical part of the ecosystem and the success of Iwokrama relies on dusk near fruiting trees. the combined skills of specialists and its community partners. Sometimes buries nuts, seeds and fruit in shallow pits for later use. Front Hind 16 17 SMALL RODENTS SMALL RODENTS

GUIANAN SQUIRREL BRISTLY MOUSE Sciurus aestuans Neacomys guianae Size: 300 g. Long, bushy tail, and long Size: 20 g. Tiny mouse with toes with claws. Diurnal, arboreal. spiny fur. Broad, flattened, Feeds on nuts, fruits, and occasionally spiny hairs obvious when fur eggs and nestling birds. Often found in is brushed backwards. Nocturnal, tree falls or tangled mats of vines. terrestrial. Solitary. Forages on the forest floor for insects, fruits, and seeds.

RED-NOSED WATER RAT TREE RAT Nectomys rattus didelphoides Size: 200 g. Robust rat with a thick Size: 500 g. Upper parts dark tail. Hind feet partially webbed. Always reddish-brown with flattened spines; found near water, it feeds on a spines with pale tips lending a distinct variety of invertebrate and aquatic “salt and pepper” appearance. Nocturnal, vertebrate prey as well as fruits and fungi. arboreal. Sits quietly and is difficult to see Silent, it may freeze motionless in beam of flashlight. except when actively foraging.

CUVIER’S TERRESTRIAL COMMON TERRESTRIAL SPINY RAT RICE RAT Proechimys cuvieri Hylaeamys megacephalus Size: 500 g. Medium-sized and Size: 50 g. Juvenile is grey, young stocky-bodied, with stiff hairs or adult dark brown, and old adult is spines on back and rump. The sometimes reddish. Nocturnal. tail is easily broken off near its Found on the forest floor, particularly in base (hence local name “bob rat”). areas with good ground cover where it feeds on Nocturnal; strictly terrestrial. seeds, fruits, and insects. The most common Most common rat in Iwokrama. mouse-like rodent in Iwokrama.

18 19 SMALL RODENTS BATS

PYGMY RICE RAT PROBOSCIS RIVER Oligoryzomys fulvescens Rhynchonycteris naso Size: 15 g. Long, narrow tail, Size: 5 g. Tiny; distinctive tufts of pale fur (longer than head and body) along forearm. Nocturnal (crepuscular), slightly paler below than above and aerial. Roosts in small groups lined up only thinly haired, without a terminal tuft. close together on the underside of Nocturnal. Solitary. Feeds on small logs or tree trunks over water. seeds and insects in brush or tall grass.

SMALL ARBOREAL GREATER WHITE-LINED RICE RAT BAT Saccopteryx bilineata Oecomys bicolor Size: 8 g. Medium sized blackish bat with two Size: 30 g. Small, red mouse. wavy white lines on the back. Begins to fly Nocturnal, arboreal. Feeds on seeds at dusk. Roosts together in groups of up to 50 and fruits. Often occupies roofs and in tree hollows or cavities of large buttress rafters of buildings near forest, trees. The bat most likely to be encountered where it is easily seen. when walking trails and searching for roosts.

CHESTNUT SAC-WINGED BAT VISIT THE IWOKRAMA FOREST! Cormura brevirostris Scientists, students, tourists, volunteers and interns are Size: 9 g. Medium-sized chestnut brown bat. encouraged to visit the Iwokrama Forest-Accommodation. Nocturnal. Begins to forage in the early Transportation and research facilities are provided at the evening. Often seen fluttering over the road Iwokrama River Lodge and Research Centre. through Iwokrama Forest. Feeds on small insects. Roosts in large rotting logs and in tree hollows in small groups.

20 21 BATS BATS

NOTHERN GHOST BAT FROG-EATING BAT Diclidurus albus Trachops cirrhosus Size: 20 g. Large, white bat with translucent Size: 30 g. Conspicuous noseleaf and pinkish wings. Nocturnal. Flies high in large, rounded ears. Mouth surrounded open areas over rivers, foraging for by wart-like bumps. Nocturnal. moths and other insects. Roosts in Flies low in the forest understory. small groups (up to 4) under palm fronds. Feeds on frogs. Roosts in small groups in tree hollows and caves. GREATER MOUSTACHED BAT Pteronotus rubiginosus FALSE VAMPIRE BAT Vampyrum spectrum Size: 20 g. Medium sized with short brown fur. Leaflike plate of skin protrudes from Size: 180 g. Largest bat in South lower lip. Nocturnal. Most active in the America. Muzzle elongated with early evening. Fast-flying aerial noseleaf positioned like a hood insectivore usually found within closed ornament. Usually forages during the ear- forest for beetles and other ly evening. Eats birds, bats and mice. insects. Often seen flying along trails. Family groups roost in hollow trees.

FISHING BAT SWORD-NOSED BAT Noctilio leporinus Lonchorhina inusitata Size: 65 g. Enormous hindfeet with large Size: 13 g. Long (about 2 cm) pointed sharp claws, and long narrow wings. noseleaf (fleshy leaf-like appendage Nocturnal. Catches small fish with on tip of nose). Active later in the evening. clawed hindfeet and then stuffs them Slow, agile flight; can hover. Feeds on insects into cheek pouches. Roosts in tree and fruit, probably by gleaning. Uncommon, hollows, usually in large groups. probably roosts in caves and tree hollows.

22 23 BATS BATS

STRIPED HAIRY-NOSED COMMON VAMPIRE BAT BAT Desmodus rotundus Gardnerycteris crenulatum Size: 30 g. Front upper teeth are very Size: 14 g. Prominent noseleaf with sharp and blade-like. Thumbs are long serrated, hairy edges. Nocturnal. and very well-developed. Agile flier, Feeds on beetles and other insects. leaves roost well after dark. Travels Small groups roost in rotting logs well on the ground, in leaps and bounds. and tree stumps, and tree hollows. Blood feeder. Preys on domesticated and wild mammals.

GREATER COMMON SHORT-TAILED FRUIT-EATING BAT FRUIT BAT Aritbeus lituratus Carollia perspicillata Size: 60 g. Four bright white stripes on the Size: 20 g. Dorsal hairs have a distinctive tricolour face. Takes flight after dusk and forages pattern of a dark base and tip with a whitish during the early evening. Feeds primarily on middle band. Regularly flies a kilometre or figs but also other fruits, flowers and more from its roost. Usually forages in pollen. Usually eats at a separate feeding the understory. One of the commonest bats in roost which facilitates seed dispersal. Iwokrama. Feeds primarily on small fruits, insects and nectar.

COMMON COMMON TENT-MAKING BAT LONG-TONGUED BAT Uroderma bilobatum Glossophaga soricina Size: 15 g. Medium-sized greyish bat with white Size: 10 g. Muzzle elongated with groove split- stripe down back. Nocturnal. Feeds primarily ting lower lip, and very long tongue. Nocturnal. on fruit, supplemented with insects and nectar. Common nectar-feeding bat found in variety Roosts under“tents” made by biting through of habitats. Roosts in tree hollows, buildings, a leaf, such as palm, until the leaf droops over to culverts, and caves. Beneficial to the ecosystem form a shelter. as a flower pollinator.

24 25 BATS BATS

SPIX’S DISK-WINGED BLACK BONNETED BAT BAT Eumops auripendulus Thyroptera tricolor Size: 30 g. Ears extend over flat Size: 4 g. Very small and delicate bat with long face to tip of nose. Long tail fluffy fur. Flight is quite agile and manoeu- extends well beyond tail membrane. vrable. Probably catches small insects in flight. High-flying aerial insectivore. Roosts upright in furled Heliconia leaves, uses Feeds on large insect prey. Roosts in suction-cup-like disks to move along the slick small groups in tree hollows and buildings. surface. COMMON MASTIFF BAT BLACK MYOTIS Molossus molossus Myotis nigricans Size: 15 g. Free-tailed bat. Small, Size: 5 g. Long tail completely greyish or reddish brown. Begins enclosed in membrane between legs. foraging at sunset, usually high Leaves the day roost at sunset and up in open areas. Feeds primarily on usually does not return until after beetles. Roosts in colonies of up to dawn. Eats moths and other small 300 in buildings and tree hollows. insects, resting at temporary roosts during the night. In the day, roosts in tree hollows, caves, and buildings.

DID YOU KNOW? VIEWING TIPS! The Iwokrama Forest holds one of the highest diversities of bats in the Most mammals are secretive and can be hard to see. Since many world with over 88 species. This distinction is represented by nine bat mammals are nocturnal, a good way to see them is at night with families including such interesting species like the False Vampire Bat, the help of a headlamp. Fruiting trees are also a good place to see mammals as they congregate to feed. And always keep an eye on the the Common Vampire Bat and the Fishing Bat. These bats represent ground for signs - especially tracks in the wet mud on the edge of every feeding guild of the forest from sanguivory to carnivory. pools.

26 27 ABOUT THE GUIDE ABOUT THE AUTHORS

This guide is an introduction to 71 of the most distinctive mammals Mark Engstrom, Ph.D., is Deputy Director of Collections and of the Iwokrama Forest. It contains concise, authoritative Research at the Royal Ontario Museum in Toronto, Canada. He descriptions of each species, written by experts in the mammal is a specialist in the evolution and systematics of New World fauna of the area and outstanding colour illustrations of each rodents and bats. species by artist and writer Fiona Reid. Included for many species are illustrations of tracks. Burton Lim, Ph.D., is Assistant Curator of Mammalogy in the Department of Natural History at the Royal Ontario Museum The Guide was first developed and published by the Iwokrama in Toronto, Canada. Research interests on the evolutionary International Centre for Rain Forest Conservation and relationships of bats has taken him to many tropical countries to Development in 1999 with funding provided by the International study the biodiversity of mammals. Development Research Centre of Canada. Fiona Reid, M.Sc., is the author and illustrator of many books on Information on the distribution of mammals in the Iwokrama mammals including “A Field Guide to the Mammals of Central Forest has been derived from surveys conducted by the Centre America and Southeast ” and the “Peterson Guide to in collaboration with participating academic partners, donors Mammals of North America”. and the communities living in, or near, the Iwokrama Forest. The Iwokrama International Centre and the North Rupununi District The North Rupununi District Development Board (NRDDB) was Development Board have also worked with local communities established in 1996, and is the umbrella organization which to gather information on the ecology, behaviour and distribution represents the 20 indigenous communities living in, or near, the of the more common animals of the Iwokrama Forest. Funding Iwokrama Forest. for the faunal surveys was provided by the Global Environment Facility through the United Nations Development Programme, the United Kingdom Department for International Development, and by the Royal Ontario Museum Foundation. Printed with support from: REPSOL EXPLORACIÓN GUYANA S.A.

Copyright © 1999 Iwokrama International Centre Illustrations Copyright - Fiona A. Reid Designed and Printed by All rights reserved. Reprinted 2017. Pixel Plus Creations | [email protected] This book may not be reproduced in whole or in part without permission.

28 MAP OF IWOKRAMA

SUSTAINABLE UTILIZATION AREA (SUA) WILDERNESS PRESERVE (WP)

IWOKRAMA INTERNATIONAL CENTRE FOR RAIN FOREST CONSERVATION AND DEVELOPMENT 77 High Street, Kingston, Georgetown, Guyana, Tel: 592.225.1504, 592.225.1186, Fax: 592.225.9199 [email protected]; [email protected] [email protected] www.iwokrama.org