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Review of the Neotropical Millipede Genus Þcnotropis CARL, 1914 (Diplopoda, Polydesmida, Aphelidesmidae)

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Review of the Neotropical Millipede Genus Þcnotropis CARL, 1914 (Diplopoda, Polydesmida, Aphelidesmidae) AMAZONIANA XY (112):67 - 102 Kiel, Dezember 1998 Review of the Neotropical millipede genus þcnotropis CARL, 1914 (Diplopoda, Polydesmida, Aphelidesmidae) by S.I. Golovatch, K. Vohland, R.L. Hoffman, J. Adis, A. Mármol, L. Bachmann & J. Tomiuk Dr. Sergei L Golovatch, Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow ll707l (V-71), Russia. Dipl.-Biol. Katrin Vohland & Priv.-Doz. Dr. Joachim Adis, Max-Planck-Institute for Limnology, Postfach 165, D-24302 Plön, Germany, in cooperation with National Institute for Amazonian Research (INPA), Manaus, Brazil. Prof. Dr. Richard L. Hoffman, Virginia Museum of Natural History, l00l Douglas Avenue, Martinsville, Virginia 24112, U.S.A. Prof. Dr. Andrés E. Mármol, I"INAP-Biologia, Departamento de Zoologia, Apartado Postal 373, Iquitos/Loreto, Peru. Priv.-Doz. Dr. Lutz Bachmann, Department of Population Genetics, University of Tübingen, Auf der Morgenstelle 28, D-72076 Tübingen, Germany. Priv.-Doz. Dr. Jürgen Tomiuk, Institute of Anthropology and Human Genetics, Wil- hemstr. 27, D-72074 Tübingen, Germany. (Accepted for publication: November, 1998). Abstract A review of Pycnotropís is presented, with the description of 14 new species: P. 'goeldii n.sp., P. madeira n.sp., P. sigma n.sp. and P. urucu n.sp., all from Brazil, P. carli n.sp. from Ecuador, P. curvata n.sp., P. falcata n.sp., P. jeekeli n.sp., P. pallidicornis n.sp., P. similis n.sp., P. subfalcata n. sp., P. unapi n.sp., P. torresi ¡.sp. and P. zumbii n.sp., all from Peru. P. acuticollis (ATTEMS, 1899), is redescribed upon the holotype from Brazil. Based on abundant strictly topotypic material, P. tida (CHAMBERLIN, l94l), is considered as a subjective senior synonym of P. epiclysmus HOFFMAN, 1995 (syn.n.). A key has been compiled to all 26 species of this diverse genus, with notes on its distribution. Keywords: Diplopoda, Polydesmida, Aphelidesmidae, taxonomy, millipede, Neotropics. ISSN 0065-6755/1998106'7/ @ MPI fûr Limnologie, AG Tropenökologie, Plön; INPA, Manaus 67 Introduction Ptyxogon CHAMBERLIN, l94l: 498. Diagnosis: Paraterga thick, small ridge in caudal view missing and united broadly with body trunk. The millipede genus Pycnotropis CARL, 1914, one of the most cotrunon and diverse Caudal comers becoming more acute only in paraterga l6 to 19. Polygonal areation weak and often lateral edges paraterga Neotropical millipede genera, has only recently been reviewed (HOFFMAN 1995) and restricted to of or missing completely. Type species: Pycnotropís taenía CARL, 1914, by original designation. shown to comprise the following presumably valid species: P. taenia (PETERS, 1864) (the type-species) from Bolivia, P. flavocarinara (SILVESTRI, 1898) and P. melanostig- ruø (SILVESTRI, 1898), from Colombia, and P. haenschi CARL, 1914, from Ecuador, sígma GOLOVATCH, VOHLAND & HOFFMAN, n.sp. (Figs. 1-4) P. latzeli ATEMS, 1931, from Panama (possibly mislabeled), P. nitida KRAUS, 1959 þcnotropís and P. achiraensis KRAUS, 1959, from Peru, P. acuticollis (ATTEMS, 1899), P. Holotype: d (INPA), Brazil, Edo. Amazônas, environs of Manaus,29 road-km on BR-O10 highway, subareata 1963, P. inca (CHAMBERLIN, 1941) and JEEKIL, P. epiclysmu.s HOFF- CPAA/EMBRAPA Experimental Station (2'52'S, 59"59'W), terra firme, coarse litter of peachpalm Bactris MAN, 1995, from Brazil, as well as P. inca (CHAMBERLIN, 1941) and P. tida sp., 31.0t.1997, leg. K. VOHLAND. - Paratypes: 8 dd (INPA), 2 ðð 2 subadult c¡c¡ (ZMUM), I d (CHAMBERLIN, l94l), from Peru andBrazll. The genus is believed Andean of origin, (ZMUC), lc¡(SMF), ld(MHNG),3crd, 19(CA), ld(MHNL), ld, l9(MZSP),samelocality, with a number of species distributed as offsprings down the Amazon Basin (cf. GOLO- dare and collector, rogethef with holotype; I c¡ (lcB-FUA), I I (INPA), same place, 11.10.1996, leg. K. VATCH et al. 1997). VOHLAND; I c¡ (INPA), same place and collector, 18.04. 1997;3 dd, I I (INPA), same place, Many species are very uniform in characters and differ mainly in coloration. Recent 28.10.1997, leg. J. ADIS. genetic analyses, however, indicated that the morphologically similar species like P. Name: Emphasizes the particularly strongly sigmoid tibiotarsus of the gonopod' sigmoid tibiotarsus of the subfalcata, P. madeira, P. t¡da, P. torresi and P. sigma n.spp. can be characterized Diagnosis: Distinguishable from congeners by the particularly strongly through species-specific pattems of isozymes (VOHLAND, BACHMANN & TOMIUK, gonopod. (d) and 47-51 mm (9), width of midbody prozonites 4.0-4.5, of unpubl.). Description: Length 40-48 mm metazonites 5.2-7.5 mm (d), and 5.0 and 7.0-8.0 mm (9), respectively. Holotype 43 mm long, 3.9 and 6.2 This contribution is part of a current study of the origin, biogeography, ecology and mm wide on midbody pro- and metazonites, respectively. Coloration in alcohol in general from blackish genetics of Amazonian species of Pycnotroprs (c.f. ADIS & MESSNER 1997; BACH- or dark castaneous brown to yellow-greyish-brown with all intergradations in between. Venter dark MANN et al. 1998; GOLOVATCH et al. 1997; VOHLAND 1998). The present paper castaneous brown to pallid, usually only a little paler than sides ofmetazonites; bases ofantennae and legs is mainly devoted to the description several new species of this genus of encountered in castaneous brown to grey-brown-yellow. Antennae always infuscate at least distally, tip ofantenna whitish. Brazil, Ecuador and Peru, as well as to redescriptions of several poorly-known older Legs often increasingly infuscate toward claw, tarsi usually particularly dark. Gonopods yellowish. congeners. Taxonomic studies were done by S.I. GOLOVATCH, K. VOHLAND and Gnathochilarium light brown to pallid. Metazonites more often completely dark without distinct markings, R.L. HOFFMAN, while most collecting was made by J. ADIS, S.I. GOLOVATCH, A. then underside ofporiferous paraterga sometimes with a brown-reddish spot below ozopore (the spot often MÁRMOL and K. VOHLAND. Material has been shared with and/or returned to the being traceable as a reddish marking even from above), or only dorsal surface of paraterga, especially following institutions: Instituto Nacional de Pesquisas da Amazônia (INPA), Manaus, peritremata, sometimes red-brownish, but never too contrastingly so. More seldom (8 of 23 specimens) Brazil; Coleção Entomológica do Laboratório de Zoologia, Universidade do Amazonas general coloration pale brown to yellow-brown-greyish, then limbus, suture between pro- and metazonites, (ICB-FUA), Manaus, Brazil; Museu Paraense Emílio Goeldi (MPEG), Belém, Pará, and areation clearly infuscate and thus evident. When present, paler patches on paraterga never strongly Brazil; Museu de Zoologia da Universidade de São Paulo (MZSP), Brazll; Museo contrasting with background coloration. Epiproct always infuscate, of background coloration. very distinct and deep. Labral setae 6+6, supralabral Nacional de Historía Natural, Lima (MHNL), Peru; Zoological Museum of the State Epicranial suture between and above antennae as bunches ofindividual hairs. Genal convexities in front ofantennal sockets University of Moscow (ZMUM), Russia; Senckenberg Museum, Frankfurt a.M. (SMF), ones2*2, some compound Surface behind antennal sockets rugulose to strongly striated. Frons sometimes Germany; Naturkunde Museum der Humboldt-Universität (ZMB), Berlin, Germany; very well-developed. rugulose. Antennae relatively short and stout, in situ scarcely surpassing segment 2laterclly; antennomeres Zoologisk Museum, University of Copenhagen (ZMUC), Denmark; Muséum d'Histoire 2 to 4 subequal in width and length, each 1.8 times as short as longest antennomere 6; antennomere 7 naturelle, Geneva (MHNG), Switzerland; Virginia Natural History Museum, Martinsville shortest, about five times as short as 6th, antennomeres 5-7 particularly densely setose. (VNHM), (FMNH), J. U.S.A.; Field Museum of Natural History, Chicago U.S.A.; and Collum with relatively broad lateral flaps broadly rounded anteriorly and nanowly so at caudal corner. ADIS (CA), Germany. Width ratios: head << coltum <2=3=4<5=17. Body gently tapering caudally further on. Surface generally smooth and shiny, only sublateral parts of collum and of subsequent metâterga with distinct areation growing increasingly, but almost never fully, obliterated dorsad, leaving most of dorsum smooth to Taxonomic part extremely finely rugulose, with only a few stronger wrinkles sublaterally behind prozonites and, seldom, Pycnotropìs CARL, 1914 also near caudal margin (Fig. I ). Darker specimens usually with exceedingly faint traces of areation across dorsum, paler ones with areation readily visible all over metatergal surface. Areation always clear as Peru, Ecuador, Colombia, Brazil. smooth and flat bosses only sublaterally, some of these with abraded setae seen as their insertion points. Pycnotropis CARL, l9 l4: 419. Prozonites entirely smooth, exceedingly delicately shagreened; metazonites below paraterga strongly Amydrínus CHAMBERLIN, l94l: 498. granulospinulose all over in segments 2-3, almost so in segment 4 or segments 4 and 5, in subsequent Phinotropis CHAMBERLIN, l94t: 498 segments somewhat variable, usually retained only (sub)caudally as a series of grains/spinules growing especially prominent dorsad and ventrad but gradually declining both mesad and toward telson. Suture 68 69 between pro- and metazonites extremely delicately striolate dorsally and dorsolaterally, almost smooth. All other main characters like in P. sígma except as follows: Caudal limbus very finely beaded at place
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