Thais Lacera (Born, 1778) (Mollusca: Gastropoda: Muricoidae: Muricidae
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First report and illustration of the mysterious Drillia saulcydianum (Recluz, 1851) (Mollusca: Gastropoda: Drilliidae) in recent literature Frank Nolf Pr. Stefanieplein, 43/8 – B-8400 Oostende [email protected] Keywords: GASTROPODA, DRILLIIDAE, Drillia Description: saulcydianum, Sierra Leone, West Africa. Pleurotoma saulcydianum Recluz, 1851 Abstract: After more than a century, to my Plate 5, fig. 6 in Recluz (1851) knowledge, a second specimen of Drillia saulcydianum (Recluz, 1851) was found in a ‘Testa subfusiformi, turrita, transversim crebre shell collection. This species was neglected or striata, longitudinaliter plicata; anfractibus erroneously identified in collections as well as in deenis, superne depressis, albido-lutescentibus, literature. After the record of this unique shell in inferne spadiceo-fasciatis; plicis superioribus Senegal, the second specimen originated from tuberculiformibus, in ultimo flexuosis, Sierra Leone. Both are described and figured in robustisque; anfractu infimo ventricoso, medio this paper. late fasciato; labro acuto, suturam versus late emarginato; canali brevissimo, emarginato. Abbreviations: Hab. Le Sénégal. (Coll. Petit.)’ FN: Private collection of Frank Nolf. This species has about 10 RBINS: Royal Belgian Institute for Natural whorls, the last whorl Sciences. occupying half the length of the whole shell. The Introduction: During a check-up of my collection ‘Pleurotoma of Saulcy’ is of COLUMBELLIDAE I accidentally found a shell nearly fusiform and from Sierra Leone I had myself labelled as turriculated, provided with Mazatlania sp. However, Mazatlania is a typical many transverse, regular, genus of the West American East Pacific area distinct striae on a glossy without representatives in West Africa. So, from surface, particularly at the a closer look under magnification, it appeared to base of the body whorl. be a turrid shell. At that time I was not in Longitudinal folds run over possession of the work of Recluz, describing and the whole surface, resulting in figuring Pleurotoma saulcydianum and even after a double row of tubercles in searching in older literature for the last months, I the upper side of each whorl, did not make the link with this shell. My and becoming particularly specimen was found in ‘crabbed condition’ and robust and flexuous in the last the brown band in the middle of the last whorl whorl. The siphonal canal is very short and was only superficially present. In the incised at the base. The aperture is oval in Dautzenberg collection (RBINS) the samples outline, whitish, with the central brown band labelled as ‘Drillia saulcydiana’ contained partly showing through. The parietal wall on the several specimens of different species such as inner lip is reflected and ends in a small, Drillia recordata Sykes, 1905 and Crassispira depressed knob. The external margin shows a fuscobrevis Rolán, Ryall & Horro, 2007 but none large shallow notch below the suture. of them was the real D. saulcydianum. It is Colour: Yellowish white with a reddish brown remarkable that we had to wait for more than a band at the base of each whorl, becoming a century to again get acquainted with that ‘lost’ broad zone in the middle of the body whorl. This species. I suspect this can be explained by the feature is hardly visible in the specimen in fact this animal only lives in small colonies in a collection FN, as it was not caught alive. region hardly accessible for fishermen and shell Size: the unique Recluz specimen in collection collectors. Petit de la Saussaye measured 21 mm; in collection FN: 15.34 mm. Material examined: One specimen trawled at 33 m off Sierra Leone in 1990. Size: 15.34 mm. FN. Neptunea, vol.7, n°2 1 March 2008 1 Plate 30, fig. 78: Drillia umbilicata Plate 8, fig. 24: Drillia dunkeri (Weinkauff, 1876) – wrongly estimated to be an immature example of D. umbilicata by Tryon (1884: 179) (Tippett, 2006) Plate 11, fig. 82: Drillia umbilicata Plate 11, fig. 91: Drillia saulcydianum From: Tryon, G.W., Jr., 1884. Conidae, Pleurotomidae. Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Vol. VI. Tryon, Philadelphia. 413 pp, 34 pls. Neptunea, vol.7, n°2 1 March 2008 2 Discussion: sinus in the outer lip produced upwards. The Tryon (1884) stated that Drillia saulcydianum inner lip is thickened below, forming a false (plate 11, fig. 91) little differs from Drillia umbilicus with the axis. The brown band in the umbilicata Gray, 1838 (plate 11, fig. 82; plate 30, middle of the whorl is restricted to pinkish brown fig. 78) ‘in sometimes possessing a broad blotches of the same colour only visible on the indistinct central band of light chestnut’ and he distinct longitudinal folds. In D. saulcydianum, continues ‘the shells are adult, although not so these folds are restricted to a double row of large as the type’. So, Tryon concluded: ‘Of its tubercles on each whorl. In general the outline of identity with D. umbilicata there can be no both species is very different. question.’ However, we can immediately ascertain D. saulcydianum and D. umbilicata are Acknowledgements: two different species. Besides the glossy surface I am very grateful to David Monsecour for both species have few characteristics in controlling the English text and to Johan common. D. umbilicata (Plate II, Figs 4-7) has a Verstraeten who revised the original content. strongly tuberculate shoulder with the broad References: Recluz, C., 1851. Description de quelques coquilles nouvelles. Journal de Conchyliologie, 2, 194-216, pls 5-6. Tippett, D., 2006. Taxonomic notes on some Indo-Pacific and West African Drillia species (Conoidea: Drilliidae). Iberus, 24(1): 13-21. Tryon, G.W., Jr., 1884. Conidae, Pleurotomidae. Manual of Conchology, Structural and Systematic, with Illustrations of the Species. Vol. VI. Tryon, Philadelphia. 413 pp, 34 pls. Tucker, J.K., 2004. Catalog of Recent and fossil turrids (Mollusca). Zootaxa, 682. Auckland. 1295 pp. Geographic distribution of Drillia saulcydianum (Recluz, 1851) Neptunea, vol.7, n°2 1 March 2008 3 1 2 3 Plate I. Figs 1-3: Drillia saulcydianum (Recluz, 1851). Sierra Leone. Dredged at a depth of 33 m. 1990. 15.34 mm. FN. Neptunea, vol.7, n°2 1 March 2008 4 4 5 7 6 Plate II. Figs 4-7: Drillia umbilicata Gray, 1838. Bay of Gorée, Dakar, Senegal. Dredged at a depth of 18 m, at night. December 1980. FN; 4-5: 25.80 mm; 6-7: 28.27 mm. Neptunea, vol.7, n°2 1 March 2008 5 Two new turrid species from West Africa: Clavatula delphinae and Clavatula pseudomystica (Mollusca: Gastropoda: Clavatulidae) Frank Nolf Pr. Stefanieplein, 43/8 – B-8400 Oostende [email protected] Keywords: GASTROPODA, CLAVATULIDAE, Type material: Clavatula delphinae, Clavatula pseudomystica, Holotype: Plage Koraf, Pointe-Noire, mouth of West Africa, new species. the Songolo-river, Congo-Brazzaville. Collected at a depth of 4 m. 1995. 13.71 mm. RBINS. Abstract: During the study of a sample of crabbed turrid shells from Pointe-Noire (Congo- Paratypes: Brazzaville) some specimens, collected by the 1. 10.81 mm (juvenile specimen) (FN). Off French shell collector Paul-Henri Hattenberger Takoradi Harbour, Ghana. Dredged among could not be identified. After comparing them shell grit, between stones, at a depth of 15 with similar shells from nearby West African m. 8 August 1999. countries it appeared to concern two new 2. 13.76 mm (JV). Pointe-Noire, mouth of the species, described and figured here as Clavatula Songolo-river, Congo-Brazzaville. Dived in delphinae and Clavatula pseudomystica. sandy mud at a depth of 4 m. 1995. 3. 14.38 mm (FN). Cap Esterias, Gabon. Abbreviations: 4. 15.40 mm (FN). Plage Koraf, Pointe-Noire, FN: Private collection of Frank Nolf. mouth of the Songolo-river, Congo- JV: Private collection of Johan Verstraeten. Brazzaville. Collected in sandy mud at a PHAT: Private collection of Paul-Henri depth of 4 m. 1995. Hattenberger. 5. 16.33 mm (PHAT). Plage Koraf, Pointe- PR: Private collection of Peter Ryall. Noire, mouth of the Songolo-river, Congo- RBINS: Royal Belgian Institute for Natural Brazzaville. Collected in sandy mud at a Sciences. depth of 4 m. 1995. Introduction: During the study of Clavatula Description: The protoconch consists of 2.5 martensi von Maltzan, 1883 it glossy whorls, followed by 7 whorls in the adult became clear that the figure in stage of the largest specimens. The body whorl Ardovini & Cossignani (2004) occupies nearly two thirds of the shell’s length. was not that species at all. An axial sculpture of numerous close-set, oblique Actually the specimen angular ribs is present on the lower part of the illustrated by the Italian authors whorls of the spire, extending just above the could represent a new species. lower suture. On the last whorl these ribs (14-15) This resulted in a complete gradually become weaker from the subsutural revision of all Clavatula species area towards the lower part of the body whorl, in the collection FN by which where they end in two rows of small nodules. some unidentified West African The whole surface of the last whorl and the sipho shells were linked to the is intersected by a fair number of spiral threads. juvenile specimens of a sample The suture is distinct and slightly undulating due gathered by P.-H. Hattenberger in Pointe-Noire to the incision of the oblique ribs followed by a (Congo-Brazzaville). Two of them differed from weakened rounded ridge and an inconspicuous the remaining specimens and were similar to subsutural area intersected by about five very shells from Ghana and Gabon. They appeared to faint encircling lines. The sinus is narrow and belong to a new species, Clavatula delphinae, neatly curved. comparable to the recently described species Colouration: The shell is covered by a pattern of Clavatula congoensis Nolf & Verstraeten, 2008. brown and bluish grey or milky-white blotches Another new species, Clavatula pseudomystica, and even more brown streaks or flammules in had to be introduced after careful comparison the subsutural zone and in the lowest part of the with turrid shells from Angolese waters.