Victorin's Gentian (Gentianopsis Virgata Ssp. Victorinii)
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Outline of Angiosperm Phylogeny
Outline of angiosperm phylogeny: orders, families, and representative genera with emphasis on Oregon native plants Priscilla Spears December 2013 The following listing gives an introduction to the phylogenetic classification of the flowering plants that has emerged in recent decades, and which is based on nucleic acid sequences as well as morphological and developmental data. This listing emphasizes temperate families of the Northern Hemisphere and is meant as an overview with examples of Oregon native plants. It includes many exotic genera that are grown in Oregon as ornamentals plus other plants of interest worldwide. The genera that are Oregon natives are printed in a blue font. Genera that are exotics are shown in black, however genera in blue may also contain non-native species. Names separated by a slash are alternatives or else the nomenclature is in flux. When several genera have the same common name, the names are separated by commas. The order of the family names is from the linear listing of families in the APG III report. For further information, see the references on the last page. Basal Angiosperms (ANITA grade) Amborellales Amborellaceae, sole family, the earliest branch of flowering plants, a shrub native to New Caledonia – Amborella Nymphaeales Hydatellaceae – aquatics from Australasia, previously classified as a grass Cabombaceae (water shield – Brasenia, fanwort – Cabomba) Nymphaeaceae (water lilies – Nymphaea; pond lilies – Nuphar) Austrobaileyales Schisandraceae (wild sarsaparilla, star vine – Schisandra; Japanese -
Diversity and Evolution of Asterids
Diversity and Evolution of Asterids . gentians, milkweeds, and potatoes . Core Asterids • two well supported lineages of the ‘true’ or core asterids ‘ ’ lamiids • lamiid or Asterid I group • ‘campanulid’ or Asterid II group • appear to have the typical fused corolla derived independently and via two different floral developmental pathways campanulids lamiid campanulid Core Asterids • two well supported lineages of the ‘true’ or core asterids lamiids = NOT fused corolla tube • Asterids primitively NOT fused corolla at maturity campanulids • 2 separate origins of fused petals in “core” Asterids (plus several times in Ericales) Early vs. Late Sympetaly euasterids II - campanulids euasterids I - lamiids Calendula, Asteraceae early also in Cornaceae of Anchusa, Boraginaceae late ”basal asterids” Gentianales • order within ‘lamiid’ or Asterid I group • 5 families and nearly 17,000 species dominated by Rubiaceae (coffee) and Apocynaceae lamiids (milkweed) • iridoids, opposite leaves, contorted corolla Rubiaceae Apocynaceae campanulids Gentianales corolla aestivation *Gentianaceae - gentians Cosmopolitan family of 87 genera and nearly 1700 species. Herbs to small trees (in the tropics) or mycotrophs. Gentiana Symbolanthus Voyria *Gentianaceae - gentians • opposite leaves • flowers right contorted • glabrous - no hairs! Gentiana Gentianopsis Blackstonia Gentiana *Gentianaceae - gentians CA (4-5) CO (4-5) A 4-5 G (2) • flowers 4 or 5 merous Gentiana • pistil superior of 2 carpels • parietal placentation; fruit capsular *Gentianaceae - gentians Gentiana -
Flora of the Sierra Nevada Aquatic Research Laboratory Annotated Catalog of Vascular Plants
Flora of the Sierra Nevada Aquatic Research Laboratory Annotated Catalog of Vascular Plants Updated June 2013 Ann M. Howald This updated catalog for the flora of the Sierra Nevada Aquatic Research Laboratory (SNARL) includes all vascular plants identified during field studies conducted from 1978 through spring 2013. Taxonomic nomenclature is mainly consistent with The Jepson Manual, Second Edition (Baldwin et al. 2012). The families, genera, and species are arranged alphabetically. The scientific name of each species is followed by a common name of popular usage. No attempt was made to determine all common names that might apply to a given species. Scientific names of native species are in bold type; those of introduced species are in plain type. Annotations for all taxa include: 1) life form (e.g., annual herb, perennial herb, etc.); 2) flower color; and 3) the general abundance and distribution at SNARL. For flowering plants, the approximate flowering period in the eastern Sierra is given. Locations of most landscape features mentioned in the text (e.g., headquarters building, stream control structure 3, etc.) and numbering of the stream control structures, are as shown on current maps of SNARL. The most recent previously applied scientific name is given in brackets. A voucher specimen deposited at a herbarium participating in the Consortium of California Herbaria (CCH) is cited for most taxa. A set of duplicate herbarium specimens is housed at SNARL. DIVISION SPHENOPHYTA Equisetaceae. Horsetail Family Equisetum laevigatum A. Braun. Smooth Scouring Rush. Rhizomatous perennial herb. Occasional in moist areas near Convict Creek, near the laboratory and other buildings. -
An Encyclopedia of Shade Perennials This Page Intentionally Left Blank an Encyclopedia of Shade Perennials
An Encyclopedia of Shade Perennials This page intentionally left blank An Encyclopedia of Shade Perennials W. George Schmid Timber Press Portland • Cambridge All photographs are by the author unless otherwise noted. Copyright © 2002 by W. George Schmid. All rights reserved. Published in 2002 by Timber Press, Inc. Timber Press The Haseltine Building 2 Station Road 133 S.W. Second Avenue, Suite 450 Swavesey Portland, Oregon 97204, U.S.A. Cambridge CB4 5QJ, U.K. ISBN 0-88192-549-7 Printed in Hong Kong Library of Congress Cataloging-in-Publication Data Schmid, Wolfram George. An encyclopedia of shade perennials / W. George Schmid. p. cm. ISBN 0-88192-549-7 1. Perennials—Encyclopedias. 2. Shade-tolerant plants—Encyclopedias. I. Title. SB434 .S297 2002 635.9′32′03—dc21 2002020456 I dedicate this book to the greatest treasure in my life, my family: Hildegarde, my wife, friend, and supporter for over half a century, and my children, Michael, Henry, Hildegarde, Wilhelmina, and Siegfried, who with their mates have given us ten grandchildren whose eyes not only see but also appreciate nature’s riches. Their combined love and encouragement made this book possible. This page intentionally left blank Contents Foreword by Allan M. Armitage 9 Acknowledgments 10 Part 1. The Shady Garden 11 1. A Personal Outlook 13 2. Fated Shade 17 3. Practical Thoughts 27 4. Plants Assigned 45 Part 2. Perennials for the Shady Garden A–Z 55 Plant Sources 339 U.S. Department of Agriculture Hardiness Zone Map 342 Index of Plant Names 343 Color photographs follow page 176 7 This page intentionally left blank Foreword As I read George Schmid’s book, I am reminded that all gardeners are kindred in spirit and that— regardless of their roots or knowledge—the gardening they do and the gardens they create are always personal. -
Gentianaceae) from Xinjiang, China
A peer-reviewed open-access journal PhytoKeys 130:Gentianella 59–73 (2019) macrosperma, a new species of Gentianella from Xinjiang, China 59 doi: 10.3897/phytokeys.130.35476 RESEARCH ARTICLE http://phytokeys.pensoft.net Launched to accelerate biodiversity research Gentianella macrosperma, a new species of Gentianella (Gentianaceae) from Xinjiang, China Hai-Feng Cao1, Ji-Dong Ya2, Qiao-Rong Zhang2, Xiao-Jian Hu2, Zhi-Rong Zhang2, Xin-Hua Liu3, Yong-Cheng Zhang4, Ai-Ting Zhang5, Wen-Bin Yu6,7 1 Shanghai Museum of TCM, Shanghai University of Traditional Chinese Medicine, Shanghai 201203, Chi- na 2 Germplasm Bank of Wild Species, Kunming Institute of Botany, Chinese Academy of Sciences, Lanhei Road 132, Heilongtan, Kunming, Yunnan, 650201, China 3 Ili Botanical Garden, Xinjiang Institute of Ecology and Geography, Chinese Academy of Sciences, Xinyuan, Xinjiang, 835815, China 4 Forestry Bureau of Xinyuan County, Xinyuan, Xinjiang, 835800, China 5 Xinjiang Agricultural Broadcasting and Television School, Xinyuan, Xinjiang, 835800, China 6 Center for Integrative Conservation, Xishuangbanna Tropical Botanical Garden, Chinese Academy of Sciences, Mengla 666303, Yunnan, China 7 Center of Conservation Biology, Core Botanical Gardens, Chinese Academy of Sciences, Mengla 666303, Yunnan, China Corresponding author: Ji-Dong Ya ([email protected]) Academic editor: Cai Jie | Received 15 April 2019 | Accepted 8 August 2019 | Published 29 August 2019 Citation: Cao H-F, Ya J-D, Zhang Q-R, Hu X-J, Zhang Z-R, Liu X-H, Zhang Y-C, Zhang A-T, Yu W-B (2019) Gentianella macrosperma, a new species of Gentianella (Gentianaceae) from Xinjiang, China. In: Cai J, Yu W-B, Zhang T, Li D-Z (Eds) Revealing of the plant diversity in China’s biodiversity hotspots. -
Plastome Phylogenomic Study of Gentianeae (Gentianaceae
Zhang et al. BMC Plant Biology (2020) 20:340 https://doi.org/10.1186/s12870-020-02518-w RESEARCH ARTICLE Open Access Plastome phylogenomic study of Gentianeae (Gentianaceae): widespread gene tree discordance and its association with evolutionary rate heterogeneity of plastid genes Xu Zhang1,2,3*† , Yanxia Sun1,2†, Jacob B. Landis4,5, Zhenyu Lv6, Jun Shen1,3, Huajie Zhang1,2, Nan Lin1,3, Lijuan Li1,3, Jiao Sun1,3, Tao Deng6, Hang Sun6* and Hengchang Wang1,2* Abstract Background: Plastome-scale data have been prevalent in reconstructing the plant Tree of Life. However, phylogenomic studies currently based on plastomes rely primarily on maximum likelihood inference of concatenated alignments of plastid genes, and thus phylogenetic discordance produced by individual plastid genes has generally been ignored. Moreover, structural and functional characteristics of plastomes indicate that plastid genes may not evolve as a single locus and are experiencing different evolutionary forces, yet the genetic characteristics of plastid genes within a lineage remain poorly studied. Results: We sequenced and annotated 10 plastome sequences of Gentianeae. Phylogenomic analyses yielded robust relationships among genera within Gentianeae. We detected great variation of gene tree topologies and revealed that more than half of the genes, including one (atpB) of the three widely used plastid markers (rbcL, atpB and matK) in phylogenetic inference of Gentianeae, are likely contributing to phylogenetic ambiguity of Gentianeae. Estimation of nucleotide substitution rates showed extensive rate heterogeneity among different plastid genes and among different functional groups of genes. Comparative analysis suggested that the ribosomal protein (RPL and RPS) genes and the RNA polymerase (RPO) genes have higher substitution rates and genetic variations among plastid genes in Gentianeae. -
Wildflowers and Ferns Along the Acton Arboretum Wildflower Trail and in Other Gardens FERNS (Including Those Occurring Naturally
Wildflowers and Ferns Along the Acton Arboretum Wildflower Trail and In Other Gardens Updated to June 9, 2018 by Bruce Carley FERNS (including those occurring naturally along the trail and both boardwalks) Royal fern (Osmunda regalis): occasional along south boardwalk, at edge of hosta garden, and elsewhere at Arboretum Cinnamon fern (Osmunda cinnamomea): naturally occurring in quantity along south boardwalk Interrupted fern (Osmunda claytoniana): naturally occurring in quantity along south boardwalk Maidenhair fern (Adiantum pedatum): several healthy clumps along boardwalk and trail, a few in other Arboretum gardens Common polypody (Polypodium virginianum): 1 small clump near north boardwalk Hayscented fern (Dennstaedtia punctilobula): aggressive species; naturally occurring along north boardwalk Bracken fern (Pteridium aquilinum): occasional along wildflower trail; common elsewhere at Arboretum Broad beech fern (Phegopteris hexagonoptera): up to a few near north boardwalk; also in rhododendron and hosta gardens New York fern (Thelypteris noveboracensis): naturally occurring and abundant along wildflower trail * Ostrich fern (Matteuccia pensylvanica): well-established along many parts of wildflower trail; fiddleheads edible Sensitive fern (Onoclea sensibilis): naturally occurring and abundant along south boardwalk Lady fern (Athyrium filix-foemina): moderately present along wildflower trail and south boardwalk Common woodfern (Dryopteris spinulosa): 1 patch of 4 plants along south boardwalk; occasional elsewhere at Arboretum Marginal -
The Phylogeny of Gentianella (Gentianaceae) and Its Colonization of the Southern Hemisphere As Revealed by Nuclear and Chloroplast DNA Sequence Variation
Org. Divers. Evol. 1, 61–79 (2001) © Urban & Fischer Verlag http://www.urbanfischer.de/journals/ode The phylogeny of Gentianella (Gentianaceae) and its colonization of the southern hemisphere as revealed by nuclear and chloroplast DNA sequence variation K. Bernhard von Hagen*, Joachim W. Kadereit Institut für Spezielle Botanik und Botanischer Garten, Johannes Gutenberg-Universität Mainz, Germany Received 21 August 2000 · Accepted 29 December 2000 Abstract The generic circumscription and infrageneric phylogeny of Gentianella was analysed based on matK and ITS sequence variation. Our results suggested that Gentianella is polyphyletic and should be limited to species with only one nectary per petal lobe. Gentianella in such a circum- scription is most closely related to one part of a highly polyphyletic Swertia. Within uninectariate Gentianella two major groups could be rec- ognized: 1) northern hemispheric species with vascularized fimbriae at the base of the corolla lobes, and 2) northern hemispheric, South Amer- ican, and Australia/New Zealand species without vascularized fimbriae. When fimbriae are present in this latter group, they are non-vascular- ized. Whereas ITS data suggested a sister group relationship between the fimbriate and efimbriate group, the matK data suggested paraphyly of the efimbriate group with Eurasian efimbriate species as sister to the remainder of the clade. Based on the phylogeny and using geological and fossil evidence and a molecular clock approach, it is postulated that the efimbriate lineage originated in East Asia near the end of the Ter- tiary. From East Asia it spread via North America to South America, and from there it reached Australia/New Zealand only once by a single long- distance dispersal event. -
W I L D F L O W E R S a N D F E R N S a L O N G T H E a C T O N a R B O R E T U M W I L D F L O W E R T R a I L a N D I N O
Wil dfl owers and Ferns Al ong t he Act on Arboret u m Wil dfl ower Trail and I n Ot her Gardens Updat ed t o Jul y 12, 2017 by Bruce Carl ey FERNS (including those occurring naturally along the trail and both boardwalks) Royal fern (Osmunda regalis): occasional along south boardwalk, at edge of hosta garden, and elsewhere at Arboretum Cinnamon fern (Osmunda cinnamomea): naturally occurring in quantity along south boardwalk Interrupted fern (Osmunda claytoniana): naturally occurring in quantity along south boardwalk Maidenhair fern (Adiantum pedatum): several healthy clumps near north boardwalk, a few in other Arboretum gardens Common polypody (Polypodium virginianum): 1 small clump near north boardwalk Hayscented fern (Dennstaedtia punctilobula): aggressive species; naturally occurring along north boardwalk Bracken fern (Pteridium aquilinum): occasional along wildflower trail; common elsewhere at Arboretum Broad beech fern (Phegopteris hexagonoptera): up to a few near north boardwalk; also in rhododendron and hosta gardens New York fern (Thelypteris noveboracensis): naturally occurring and abundant along wildflower trail * Ostrich fern (Matteuccia pensylvanica): well-established along many parts of wildflower trail; fiddleheads edible Sensitive fern (Onoclea sensibilis): naturally occurring and abundant along south boardwalk Lady fern (Athyrium filix-foemina): moderately present along the trail and south boardwalk Common woodfern (Dryopteris spinulosa): 1 patch of 4 plants along south boardwalk; occasional elsewhere at Arboretum Marginal -
Fremontia Journal of the California Native Plant Society
$10.00 (Free to Members) VOL. 40, NO. 1 AND VOL. 40, NO. 2 • JANUARY 2012 AND MAY 2012 FREMONTIA JOURNAL OF THE CALIFORNIA NATIVE PLANT SOCIETY THE NEW JEPSONJEPSON MANUALMANUAL THE FIRST FLORA OF CALIFORNIA NAMING OF THE GENUS SEQUOIA FENS:FENS: AA REMARKABLEREMARKABLE HABITATHABITAT AND OTHER ARTICLES VOL. 40, NO. 1 AND VOL. 40, NO. 2, JANUARY 2012 AND MAY 2012 FREMONTIA CALIFORNIA NATIVE PLANT SOCIETY CNPS, 2707 K Street, Suite 1; Sacramento, CA 95816-5130 FREMONTIA Phone: (916) 447-CNPS (2677) Fax: (916) 447-2727 Web site: www.cnps.org Email: [email protected] VOL. 40, NO. 1, JANUARY 2012 AND VOL. 40, NO. 2, MAY 2012 MEMBERSHIP Membership form located on inside back cover; Copyright © 2012 dues include subscriptions to Fremontia and the CNPS Bulletin California Native Plant Society Mariposa Lily . $1,500 Family or Group . $75 Bob Hass, Editor Benefactor . $600 International or Library . $75 Patron . $300 Individual . $45 Beth Hansen-Winter, Designer Plant Lover . $100 Student/Retired/Limited Income . $25 Brad Jenkins, Cynthia Powell, CORPORATE/ORGANIZATIONAL and Cynthia Roye, Proofreaders 10+ Employees . $2,500 4-6 Employees . $500 7-10 Employees . $1,000 1-3 Employees . $150 CALIFORNIA NATIVE PLANT SOCIETY STAFF – SACRAMENTO CHAPTER COUNCIL Executive Director: Dan Glusenkamp David Magney (Chair); Larry Levine Dedicated to the Preservation of Finance and Administration (Vice Chair); Marty Foltyn (Secretary) Manager: Cari Porter Alta Peak (Tulare): Joan Stewart the California Native Flora Membership and Development Bristlecone (Inyo-Mono): -
Key to Genera and Families
KEY TO GENERA AND FAMILIES Identification notes: The key is highly artificial and unabashedly pragmatic. One can get to the sub-keys (Key A, Key B, Key A7, etc.) by proceeding through the general key, or by jumping directly to the sub-key based on its “description”. In order to accommodate both access methods, some taxa are keyed in 2 or more sub-keys, but would logically be found only in one sub-key if one proceeded accurately through the general key. For instance, floating aquatic pteridophytes are keyed in both Key A2 and Key C1, though a logical procession through the general Key would key them into Key C1, and not allow them to appear as well in Key A2; they are keyed as well in Key A2, so that if it is apparent or determinable to the user that they are vascular cryptogams, they can be found via that key as well. The arrangement of leaves (alternate, whorled, or opposite) and their disposition (basal or cauline) is used frequently in the keys. Alternate leaves are attached at the stem 1 per node, opposite leaves 2 per node, and whorled leaves 3 or more per node. Note however, that alternate leaves are sometimes (especially at the base of plants or at the tips of woody branches, such as short shoots) arrayed with very short internodes, leading to them being closely clustered and mistakable as whorled or opposite. Note that some plants (Hypericum, Eupatorium, many Lamiaceae, many others) have a strong tendency to have axillary shoots in the axils of primary leaves; these are often referred to as axillary fascicles. -
Intraspecific Inversions Pose a Challenge for the Trnh-Psba Plant DNA Barcode
Intraspecific Inversions Pose a Challenge for the trnH-psbA Plant DNA Barcode Barbara A. Whitlock1*, Amanda M. Hale2, Paul A. Groff1 1 Department of Biology, University of Miami, Coral Gables, Florida, United States of America, 2 Department of Biological Sciences, Texas Christian University, Fort Worth, Texas, United States of America Abstract Background: The chloroplast trnH-psbA spacer region has been proposed as a prime candidate for use in DNA barcoding of plants because of its high substitution rate. However, frequent inversions associated with palindromic sequences within this region have been found in multiple lineages of Angiosperms and may complicate its use as a barcode, especially if they occur within species. Methodology/Principal Findings: Here, we evaluate the implications of intraspecific inversions in the trnH-psbA region for DNA barcoding efforts. We report polymorphic inversions within six species of Gentianaceae, all narrowly circumscribed morphologically: Gentiana algida, Gentiana fremontii, Gentianopsis crinita, Gentianopsis thermalis, Gentianopsis macrantha and Frasera speciosa. We analyze these sequences together with those from 15 other species of Gentianaceae and show that typical simple methods of sequence alignment can lead to misassignment of conspecifics and incorrect assessment of relationships. Conclusions/Significance: Frequent inversions in the trnH-psbA region, if not recognized and aligned appropriately, may lead to large overestimates of the number of substitution events separating closely related lineages and to uniting more distantly related taxa that share the same form of the inversion. Thus, alignment of the trnH-psbA spacer region will need careful attention if it is used as a marker for DNA barcoding. Citation: Whitlock BA, Hale AM, Groff PA (2010) Intraspecific Inversions Pose a Challenge for the trnH-psbA Plant DNA Barcode.