© Springer-Verlag Berlin Heidelberg 2016 R. Gerecke, T. Gledhill, V. Pešić, H. Smit, Süßwasserfauna Von Mitteleuropa, Bd. 7

Hydrachnidia 377

9-123: Hydrachnidia, plesiotypical idiosoma organization, dorsal view (from Davids et al. 2007). Terminology following Lundblad 1927 (in parentheses: following Zahvatkin 1952, after Tuzovskij 1987).

Explanation of abbreviations: Zahvatkin 1952 Lundblad 1927 – Fr: Frontale Fch: frontalis chelicerarum Dgl-1: Dorsoglandulare 1 (Antenniforme) Vi: verticalis interna Dgl-2: Dorsoglandulare 2 Oe: occipitalis externa Dgl-3: Dorsoglandulare 3 Hi: humeralis interna Dgl-4: Dorsoglandulare 4 Sci: scapularis interna Dgl-5: Dorsoglandulare 5 Li: lumbalis interna Dgl-6: Dorsoglandulare 6 Si: sacralis interna Dgl-7: Dorsoglandulare 7 (= Vgl-4, Ventroglandulare 4) Ve: verticalis externa Lgl-1: Lateroglandulare 1 He: humeralis externa Lgl-2: Lateroglandulare 2 Sce: scapularis externa Lgl-3: Lateroglandulare 3 Le: lumbalis externa Lgl-4: Lateroglandulare 4 Oi: occipitalis interna Postoc: Postoculare Fp: frontalis pedipalporum Preoc.: Preoculare – Dc-1-4: Dorsocentralia 1-4 – Dl-1-4: Dorsolateralia 1-4

© Springer-Verlag Berlin Heidelberg 2016 R. Gerecke, T. Gledhill, V. Pešić, H. Smit, Süßwasserfauna von Mitteleuropa, Bd. 7/2-3 Chelicerata, Süßwasserfauna von Mitteleuropa, DOI 10.1007/978-3-8274-2689-5_1 378 Hydrachnidia

9-124: Hydrachnidia, plesiotypical idiosoma organization, ventral view (from Davids et al. 2007). Terminol- ogy following Lundblad 1927 (in parentheses, following Zahvatkin 1952, after Tuzovskij 1987).

Explanation of abbreviations: Ac-1-3: Acetabulum 1-3 An: Anus Genpl: Genital plate Pregen: Pregenital plate Postgen: Postgenital plate Zahvatkin 1952 Lundblad 1927 – Cx-1-4: first to fourth coxae Hv: humeralis ventralis Cxgl-2: Coxoglandulare 2 (Cxgl-1 and -3 not existing) Sce: scapularis externa Cxgl-4: Coxoglandulare 4 Si: sacralis interna Dgl-7: Dorsoglandulare 7 (= Vgl-4: Ventroglandulare 4) Pi: praeanalis interna Vgl-1: Ventroglandulare 1 Se: sacralis externa Vgl-2: Ventroglandulare 2 Ci: caudalis interna Vgl-3: Ventroglandulare 3 – V-1-4: Ventralia 1-4 Hydrachnidia 379

9-125: a-d, Hydrachnidia, gnathosoma and its appendages in lateral view, example of Thyopsis cancellata (Protz, 1896) (from Davids et al. 2007); a, gnathosoma and left palp; b, right palp laterally (indicating num- bering of segments and traditional segment terminology); c, chelicera; d, palp indicating important measure- ment distances (L = length, H = height); e, Hydrachnidia, schematical lateral view; dashed lines indicate hy- pothetical segment borders (for abbreviations see Figs 8-75-76, frontal eye posterodorsally, paired lateral eyes posterolaterally from Dgl-2; setae Ce reduced in most taxa, developed only in postlarval instars of Limnochares and Pontarachnidae; after Tuzovskij 1987). 380 Hydrachnidia

9-126: Hydrachnidia, habitus (from Davids et al. 2007); a, Wandesia female, ventral view (from Di Sabatino et al. 2002); b, Hydrovolzia male, ventral view (detail: glandulare); c, Hydrovolzia male, dorsal view (arrow: dorsal furrow), d, Panisopsis male, dorsal view (from Di Sabatino et al. 2002), e, Sperchon male, ventral view (detail: glandulare); f, Arrenurus male, ventral view; g, Aturus female, ventral view, h, Ljania male, ventral view. 381 Hydrachnidia, key to superfamilies

1 Legs with three claw-like, sickle-shaped appendages: paired, ventrally pectinate, true claws, and an empodium similar in shape, but without pectination; idiosoma vermiform, without muscle attachment sclerites; rod-shaped frontal sclerite (8-79 a) with one unpaired, and three paired setae; eyes strongly reduced or completely absent; legs covered by rather uniform hairs over the segment surface ("hypertrichous"); palp compact (8-80 a), P-1-3 fused, P-4+5 forming a functional unit; P-5 without claws, bearing one stiff terminal seta; pale reddish species of interstitial waters...... ��Stygothrombioidea (Vol. I, page 289) – Legs with two (or occasionally less) claw-like appendages (8-79 h-k): empodium not claw like; claws rarely with ventral pectination; idiosoma rarely vermiform (8-78 a), with or without muscle attachment sclerites; if a rod-shaped frontal sclerite is present (e. g., 8-79 b), never bearing an unpaired anterior seta; eyes various, in most cases with paired lateral eyes, sometimes also with an unpaired frontal eye (8- 79 e, g); legs rarely hypertrichous, in general with distal margin setae stronger than setae on remaining segment surface (8-79 h-k); palp various (8-80 b, e-f, h-i, l-o), P-5 generally with claws, rarely bearing only one stiff terminal seta. ��2 2 (1) Palp chelate (P-4 forming a chela with P-5) and P-1 longer than P-2, P-3 longer than P-4, P-4 reduced in size, maximum height of palps in P-1, the following segments gradually tapering (8-80 b); gnathosoma in ventral view characteristically narrowed at the base of the slender, pointed rostrum (8-80 b, d), chelicerae needle-like (basal segment + claw fused to a single unit, 8-80 c); genital field (8-81 c) with maximum width in the anterior part, bearing numerous acetabula located anteriorly from, or on the level of, the anterior margin of the gonopore; rather large, globular red species of standing waters.. . . . �� Hydrachnoidea (Vol. I, page 333) – If palp chelate, neither P-1 longer than P-2, nor P-4 shorter than P-3, maximum height of palps in general in segments P-2-4 (8-80 e-f, h-i, l-o); gnathosomal rostrum, if present, generally not pointed (8-80 i), chelicerae two-segmented, not needle-like (8-80 k, p); genital field various in shape, if numerous acetabula present, they are arranged in a different manner or the genital field has not its maximum width anteriorly from the gonopore (8-81 a-b, e, h, m-o); larger or smaller species, various in colour and shape. ��3 3 (2) Lateral eyes, if present, incorporated into a frontal sclerite (8-79 b-d, f); palps not chelate (8-80 e-f, h), in some groups transformed by fusion of segments (e. g., 8-80 f), claws of P-5 straight (8-80 h) or reduced to fine tips (8-80 f); mouth opening surrounded by a circular, membranous fringe (8-80 g) or (in Apheviderulicidae, so far not recorded from the study area, 8-80 f) reduced to a fine slit in the centre of a conspiciously enlarged gnathosoma; acetabula numerous, or on roundish sclerites flanking the gonopore (8-81 a), or in the membranous perigenital area (8-81 b), or extremely reduced in size scattered all over the membranous idiosoma surface (8-81 d); glandular openings occasionally with semilunar, fan-shaped openings (8-79 f, 8-81 a); large, red species of temporary and permanent standing waters (outside the study area also pale inhabitants of springs and interstitial waters)...... ��Eylaoidea (Vol. I, page 301) – Lateral eyes, if present, not incorporated into a frontal sclerite (but occasionally very close to such a sclerite, 8-79 g); palps various in shape, P-5 in general with hook-like distal claw setae (8-80 i, l-o); mouth opening various, but neither surrounded by a circular fringe, nor in the centre of an extremely enlarged idiosoma; acetabula in three pairs or numerous (8-81 e-o), in rare cases invisible by light microscope; glandular openings never fan-shaped (8-79 e, g). ��4 4 (3) I/II-L directed anteriorly, III/IV-L directed posteriorly, the two leg groups separated by a large interspace (8-78 b); genital field minute, in the centre of the ventral idiosoma, devoid of acetabula (these numerous, mostly invisible in light microscope and inserted as minute cups on the medial coxal surface); in the postgenital area two large unpaired plates, the anterior one with the anal pore, or all perigenital plates and platelets fused to a shield embracing the genital field; dorsum with two shields in "T" position (8- 78 c); glandular openings and setae on paired paletelets (8-78 b, arrow); flattened inhabitants of springs, cascades and interstital waters. �� Hydrovolzioidea (Vol. I, page 294) – If legs arranged in two groups, they are not orientated into two opposite directions, and the interspace is less extended (8-78 a, d); genital field various, (with the exception of the marine Pontarachnidae) always with groups of acetabula arranged in various ways (8-81 e-o); if dorsal shields present, not arranged in a "T" position; glandularia and their associated setae together on one single platelet (8-79 e, g, arrow). ��5 382 Hydrachnidia, key to superfamilies

9-127: Hydrachnidia (from Davids et al. 2007); a-g, frontal sclerites, h-k, legs; a, Stygothrombium; b, Limno- chares; c-d, Eylais; e, Euthyas; f, Piersigia; g, Hydryphantes; h, Parathyas IV-L; i, Hydryphantes (Polyhydry- phantes), IV-L; k, Trichothyas I-L. Hydrachnidia, key to superfamilies 383

5 (4) Palps chelate (8-77 a-b, d, 8-80 i, arrow: P-4 with a dorsal extension or a heavy dorsal seta dorsally flanking P-5); idiosoma smooth or with regularly arranged muscle insertion sclerites (8-78 d), only in exceptional cases with extended dorsal and ventral shields; acetabula in various arrangements in, or around, the genital field (8-81 e, g-h, l), but never embedded in a sclerotized ventral shield; diversified morphological adaptations, but mostly larger species, all kinds of inland waters...... ��Hydryphantoidea (Vol. II, page 1) – Palps not chelate (8-80 l-o); idiosoma various; if a sclerotized ventral shield is present, it may include also some, or all acetabula (8-78 f-h, 8-81 m-o). ��6 6 (5) Palps uncate: P-4 with a ventrodistal extension opposing P-5 (8-80 o) or at least P-4 ventrally bearing a projecting setal tubercle (8-80 m); idiosoma in general heavily sclerotized (8-78 f); genital field without movable genital flaps; acetabula numerous, or on fixed plates flanking the gonopore (often fused to the ventral shield as given in 8-81 n), or lined up in the membranous area flanking the gonopore, or both (8-81 m); various in colour and dimensions, mostly in standing waters, but also in spring and interstitial habitats...... ��Arrenuroidea (Vol. III, page 253) – Palps not uncate (8-80 l, n: P-4 without extensions flanking P-5); movable flaps may be present in the genital field (8-81 f), acetabula in three pairs or numerous - or located between the flaps (8-81 f), or on immovable genital plates (8-81 i-k) - if these plates are shifted to the posterior margin and variously fused with surrounding sclerites, in ventral view acetabula may be visible tangentially (8-78 g, 8-81 o); various in colour and dimensions, all kinds of habitats. ��7 7 (6) Acetabula located in two rows between the gonopore and movable genital flaps (8-81 f, 2-4 f) (if, in rare exceptions of species so far not recorded in the study area, they are fused to the flaps, a suture indicates the former line of separation between acetabula and flaps); if a ventral shield is present, genital flaps remain always separate and movable. �� Lebertioidea (Vol. II, page 76) – Acetabula located on immovable plates flanking the gonopore, they are fully integrated, no suture lines are recognizable between acetabula and plate surface (8-78 g-h; 8-81 i, k); acetabula-bearing plates may be fully integrated into the ventral shield; in this case, acetabula are in three to four pairs and directed ventrally (8-78 h), or multiplied and directed caudally (8-78 g, 8-81 o). Hygrobatoidea (Vol. III, page 1) 384 Hydrachnidia

9-128: Hydrachnidia, mouthparts (from Davids et al. 2007); a, Stygothrombium palp; b, Hydrachna gnathosoma and palp laterally; c, Hydrachna chelicera; d, Hydrachna gnathosoma dorsal view; e, Limnochares palp; f, Apheviderulix gnathosoma and palps ventral view; g, Eylais gnathosoma ventral view; h, Eylais gnathosoma and palp lateral view; i, Hydryphantes gnathosoma and palp lateral view; k, Hydryphantes chelicera; l, Lebertia palp; m, Mideopsis palp; n, Atractides palp; o, Arrenurus palp; p, Arrenurus chelicera. Hydrachnidia 385

9-129: Hydrachnidia, genital fields (from Davids et al. 2007); a, Piersigia female; b, Limnochares female; c, Hydrachna male; d, Eylais male (acetabula in the integument); e, Protzia female; f, Sperchon female; g, Hy- dryphantes s.str. male; h, Hydryphantes (Polyhydryphantes) female; i, Hygrobates male; k, Hygrobates female; l, Wandesia (Pseudowandesia) female; m, Balcanohydracarus female; n, Arrenurus female; o, Aturus male. 386 Acknowledgements The following colleagues helped with loan of collection material: A. Hänggi & U. Wüest (Naturhis- torisches Museum Basel); J. Dunlop & A. Friederichs (Museum für Naturkunde der Humboldt- Universität Berlin); P. Tuzovskij (Hydrobiological Institute, Borok); A.Kolibać & I. Malinovský (Moravian Museum Brno); M. Holmes (National Museum of Ireland, Dublin); J. Altmann & P. Jäger (Senckenberg-Museum Frankfurt); M. JUDSON (Musée d’Histoire Naturelle Paris); N. Jöns- son & T. Kronestedt (Naturiska Rijksmuseet, Stockholm). Jürgen Hevers made kindly available the originals of his figures from hisUnionicola paper published in 1977. Peter Martin (Kiel) corrected a former manuscript version and gave important suggestions for im- proving the contents. Additional advice for single chapters was kindly provided by Jürgen Hevers (Braunschweig: Unionicolidae) and Petr Tuzovskij (Borok: Pionidae). Furthermore, we are thank- ful to Harry Boonstra (Haren), Dave Cook (Paradise Valley), Holger Müller (Rudolstadt), Ian Smith (Ottawa), Barend Van Maanen (Roermond) and Rink Wiggers (Haren) for helpful comments and useful advice. Basic taxonomic revisions of R. Gerecke were supported by grants of the Deutsche Forschun- gsgesellschaft (Revision of Aturidae, Feltriidae and Hygrobatidae), the Musèe d'Histoire Naturelle Luxembourg and the EU (SMNH Stockholm: Highlat project, MNHN Paris: Synthesys projects, in- vitation as a chercheur invité). Terry Gledhill thanks the Council of the Freshwater Biological Association for continued support. 387 Glossary acetabular plate see "genital field" anchoral process proximal extension of the gnathosoma serving as an attachment site of muscles which move mouth parts antagonistic bristle pointed dorsal projection of P-4 flanking P-5. anterior keel see "ejaculatory complex" anterior ramus see "ejaculatory complex" apical setae see "ejaculatory complex" body pores fine pores in sclerotized plates of Halacaridae and Hydrachnidia, at least in some Hydrachnidia species facilitating gas exchange processes. cauda extension of the posterior part of the idiosoma, more or less distinctly set off from the remainder of the body; frequently found only in males of species with sexual dimorphism in idiosoma shape; posteriolateral extensions of the cauda (found mainly in male representatives of Arrenuridae) are called "pygal lobes" chelate morphological state of (→) chelicera or (→) palp with two segments acting together like the branches of scissors. coxoglandularia see "glandularia" distal arm see "ejaculatory complex" dorsal furrow see "dorsal shield" dorsal shield large sclerotized shield covering the whole dorsal idiosoma, originating from the fusion of the (→) "dorsalia", in general not including (→) "lateral eyes" and (→) "lateroglandularia"; in most cases, this shield separated from the (→) ventral shield by a ring-shaped membranous area called "dorsal furrow" dorsalia dorsal muscle attachment sclerites; plesiotypically paired, in a total number of eleven on each side, in a regular pattern following the original segmentation: pre- and postfrontalia (prefr, postfr) flanking the area of the (→) median eye , sometimes fused with the (→) frontale; five dorsocentralia (dc) paralleling the median line; four dorsolateralia (dl) arranged more laterally; sclerites may be fused in various ways; fusion of all sclerites produces the (→) dorsal shield. dorsocentralia see "dorsalia" dorsolateralia see "dorsalia" dorsoglandularia see "glandularia" ejaculatory complex distal part of the male internal genital system, responsible for the composition and deposition of "spermatophores", for morphology consult 7-4 in Vol. I; plesiotypically a complex organ made up of unpaired keels ("anterior, posterior keel"; lat. carina anterior, posterior), paired sclerite arms ("distal, proximal arm", lat. brachium distale, proximale; "proximal processus", lat. processus proximalis), a proximal vesicular structure ("proximal chamber", lat. cella proximalis), and an apical sclerite bridge ("anterior ramus", lat. ramus anterior) bearing paired setae ("apical setae", lat. setae apicales), in many clades modified for indirect or direct spermatophore transmission. (= "genital skeleton"; [German] "Penisgerüst") 388 Glossary genital field area surrounding the opening of the reproductive system ("gonopore"), consisting of the following elements: paired sclerotized plates called "acetabular plates" or "genital plates" ([German] "Genitalplatten") when acetabula are located on their surface, but "genital flaps" or "genital valves" ([German] "Genitalklappen") when acetabula are placed beneath them and can be covered by the door-like closing flaps; small unpaired anterior ("pregenital") and posterior ("postgenital") sclerites delimiting the slit-shaped gonopore; occasionally, a depressed area surrounding the male gonopore is called "genital pit" genital flap, pit, plate, valve see "genital field" genital skeleton see "ejaculatory complex" glandulae limnesiae see "glandularium" glandularium defence organ typical for, and a synapomorphy of, Hydrachnidia; consisting of a round plate bearing a sensory seta and a glandular ope ning; plesiotypically, glandularia are distributed over the idiosoma in a characteristic ground pattern reflecting the original segmentation of the arthropod body: six pairs of "dorsoglandularia" flanking the median line, four pairs of "lateroglandularia" in row along the lateral line, four pairs of "coxoglandularia" placed on or near the coxae, and four pairs of "ven- troglandularia" surrounding the anal area; a particular pair of glandularia found in species of the family Limnesiidae on coxae 4, occasionally called "glandulae limnesiae"; for further terminological details, see 8-78-79. hypostomal setae pair of setae on the ventral surface of the gnathosomal rostrum in water mite larvae lateroglandularia see "glandularium" petiole small projecting sclerite at the posterior end of the idiosoma, mainly found in males of Arrenurus-species, often combined with the formation of a cauda (see there) posterior keel see "ejaculatory complex" postfrontalia see "dorsalia" postgenital sclerite see "genital field" postocularia paired, knob-shaped, seta-bearing sclerites in the ground plan of Hydrach- nidia located posteriorly from the lateral eyes prefrontalia see "dorsalia" pregenital sclerite see "genital field" preocularia paired seta-bearing sclerites in the ground plan of Hydrachnidia located anteriorly from the lateral eyes proximal arm see "ejaculatory complex" proximal chamber see "ejaculatory complex" proximal processus see "ejaculatory complex" pygal lobes see "cauda" subchelate morphological state of palp with the terminal segment P-5 acting against P-4 like a clasp-knife (occasionally called "uncate") Glossary 389 swimming setae elongated setae arranged mainly on III-L-4/5 and IV-L-4/5, more rarely also on the anterior legs and/or on more proximal segments; in general in rows or as single elements distally on the anterior segment surface, reaching in length at least 50 % of the following segment (occasionally called "swimming hairs") uncate see "subchelate" ventralia paired ventral muscle attachment sclerites; plesiotypically found in water mites in the circumanal area in a total number of four on each side ("ven- trale 1-4"); sclerites may be fused in various ways; fusion of all sclerites and the coxae produces the ventral shield. 390

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Addenda and corrigenda to volume 7/2-2: p.3, couplets 9 (3), second part: replace “……. 1” with “…….. 11” p.5, figure references in couplets 14 (13) and 15 (14): replace “(8-17 a, d)” with “(8-19 a, d)” p.8, figure reference in couplets 24 (23): replace “(8-16 a)” with “(8-18 a)” p.9, couplet 2 (1): In exceptional cases, 1-2 anterior swimming setae may be found on IV-L-5 of Hydrodroma despiciens, in general however, asymmetrically only on one side. Thus, in questionable cases we suggest to check the setation at both legs. Another character mentioned by Wiles (1985), regarding the shape of dorsal leg setae, is helpful in questionable cases (use magnifications 200-400x): 2 (1) III-L-4 with more than eight swimming setae, IV-L-5 anteriorly in general without swimming setae; integument papillae rounded. Dorsodistal setae on I-III-L-5 apically slightly thickened. �� Hydrodroma despiciens (Vol. II, page 9) – III-L-4 at most with eight swimming setae, IV-L-5 anteriorly with 2-5 swimming setae; integument papillae bluntly pointed. Dorsodistal setae on I-III-L-5 apically not thickened. �� Hydrodroma torrenticola (Vol. II, page 12) p.16, heading Euthyas truncata: replace Neuman, 1874” with “Neuman, 1875” p.46-47, couplet 6 (5): The shape of the frontal shield anterior margin is rather variable in both, H. placationis and H. dispar. This character is of secondary importance in species identification and does not merit the first place, the indications concerning the ratio anterior width/medial length of the frontal shield are erroneous. Furthermore, in the part concerning P-4, the term “width” has to be replaced by “height”: 6 (5) Cheliceral claw very short (8-25 f - basal segment/claw > 2.8); P-4 long and slender (8-25 h - length > 200 µm, length/height > 3.0). Frontal shield anteriorly hardly projecting, enlarged (8-25 g). ��Hydryphantes placationis (Vol. II, page 52) – cheliceral claw relatively longer (8- 26 c - basal segment/claw < 2.8); P-4 short and stout (8-26 b - length < 200 µm, length/ height < 3.0). Frontal shield often anteriorly projecting in an obtuse angle, more slender (8-26 a). �� Hydryphantes dispar (Vol. II, page 50) p.48, couplet 10 (9): Significant differences between larvae reared from Hydryphantes ruber-like specimens differing in the shape of the frontal shield, induced Tuzovskij to re-establish the species Hy- dryphantes hellichi Thon, 1899 (see Tuzovskij 2014f) and H. prolongatus Thon, 1899 (see Tuzovskij 2015a). After finding coexistingHydryphantes ruber-like adults showing all transi- tional situations between more or less slender frontal shields, Lundblad (1962) had considered both taxa as synonyms of Hydryphantes ruber (p. 53). Following Tuzovskij’s point of view, the key should be extended as follows: 416 Addenda and corrigenda to volume 7/2-2

10 (9) Frontal shield slender, medially as long as anteriorly wide. �� 11 – Frontal shield less slender; medially shorter than anteriorly wide. �� 12 11 (10) Frontal shield minimum width (measured between the concave lateral margins) equal to, or minor than 50 % frontal shield length. �� Hydryphantes hellichi (see Tuzovskij 2014f) – Frontal shield minimum width longer than 50 % frontal shield length �� Hydryphantes prolongatus (see Tuzovskij 2015a) 12 (10) Gnathosomal rostrum longer (8-27 f); occasionally idiosomal muscle insertions sclerotized as small dots. �� Hydryphantes ruber (Vol. II, page 53) – Gnathosomal rostrum shorter (8-27 h); idiosomal muscle insertions unsclerotized (8-27 g) . �� Hydryphantes planus (Vol. II, page 52) p.52, 6. Hydryphantes (Hydryphantes) placationis, Preadult stages: replace “Larva unknown” with “Larva: Wainstein (1980)”. p.99, page reference in couplet 2 (1): replace “Lebertia natans (page 107)” with “Lebertia natans (page 110)” p.106, Similar species, line 10: replace “females 135-190 (290-210)” with “females 135-190 (190-210)” p. 140, [Subgenus Pilolebertia, key to species] Indications that lead to Lebertia longiseta/inaequalis are not correct: (1) The name-giving length of the ventral seta on P-2 of L. longiseta is distinctly higher than in L. inaequalis, but it does never exceed the length of P-2+3. (2) the position of the medio-/dorsodistal setae on P-3 is variable in both compared species. (3) Specimens of L. longiseta may have less swimming setae in II-L-5. Here a corrected version of this part of the key: 3 (2) Ventral margin of gnathosoma near base of mouth opening straight. Ventral seta of P-2 fine and long (ratio seta length/ventral length P-2 > 1.5); P-2 often stouter (length/ height 1.4-1.6) (8-54 c). II-L-5 with six to seven posterior swimming setae. . . . . ��Lebertia longiseta (page 143) - Ventral margin of gnathosoma near base of mouth opening with a fine kink. Ventral seta of P-2 thicker and shorter (ratio seta length/ventral length P-2 < 1.4); P-2 often more slender (length/height 1.5-1.8) (8-54 e). II-L-5 with two to six posterior swimming setae. �� Lebertia inaequalis (page 142) Taxonomic index Acari III 417 Taxonomic index Acari III (underlined: accepted taxa dealt with in this book) (bot.) = plant taxa slight differences in spelling are possible between text and index (e. g., “chironomid” = “Chironomidae”)

A alpicola, Piona 195, 199, 201 abbreviator, Arrenurus 265, 273, 274, 279 alveolatus, Balcanohydracarus 350, 355 Acalyptonotidae 253, 257 Amazonella 229 Acalyptonotus 254, 257 ambigua, Piona 194, 197, 202 Acantholimnesia 166 ambiguus, Curvipes 202 Acercella 213 ambulatoria, Limnesia arevaloi 171 Acercopsis 213, 214, 216, 217 americana, Forelia liliacea 184 Actinacarus 95 Anamundamella 349 aculeata, Unionicola 246 anatina, Anodonta (Mollusca Bivalvia) 239, acuticauda, Brachypoda 67 240, 241, 246, 247, 248, 249 acutidentatus, Mixobates 160 anatolicus, Atractides 140 acutirostris, Atractides 147 Anchistalbia 8 acutirostris gibberimarginatus, Atractides 147 Andesobates 95 Adelaxonopsella 49 angelieri, Arrenurus 281 adenophorus, Atractides 123 angelieri, Atractides 121 adnatus, Arrenurus 325 angulata, Axonopsalbia 59 adnatus, Atractides 121 angulata, Barbaxonella 51, 54, 59 adnatus, Atractides vaginalis 121 angulifer, Arrenurus 261 aemiliana, Pseudofeltria 189 angusta, Kongsbergia 36, 40, 41 affinis, Arrenurus 280 angustata, Limnesia maculata 173 affinis, Atractides 129 angustipalpis, Hygrobates 152 Africasia 351 angustiporus, Atractides 122 agilis, Neumania 233 angustiporus, Megapus 122 airoloensis, Feltria 90 Anisoptera (Insecta Odonata) 272, 300, 302 alata, Kongsbergia 46 annemiae, Chelomideopsis 350, 351 albanorum, Kongsbergia 42 annulata, Piona 197, 202 albaruthenicus, Atractides 110, 119, 133 annulatus, Curvipes 202 albator, Hydrachna 280 Anodonta (Mollusca Bivalvia) 239, 240, 241, Albaxona 49, 51, 53, 56 246, 247, 248, 249 Albia 4, 8, 9 Anodontinatax 239, 240, 241, 246 Albiella 8, 9 arcuatus, Atractides 128 Albiinae 7, 8, 49 arenaria, Kongsbergia 35 algeriensis, Atractides 128 arevaloi arevaloi, Limnesia 170 algeriensis, Krendowskia 357 armata, Axonopsis 63 Allararrenurus 259 armata, Feltria 3, 82 allgaier, Atractides 121 Arrenuridae 7, 253, 255, 259, 264, 387 Allokrendowskia 357 Arrenuroidea 1, 253, 254, 256, 383 Allolemienia 231 Arrenurus 259, 261 Allolimnesia 167 Asiabates 95 Allomegapus 144 asperrimum, Nautarachna 199 Allomideopsis 375 asper, Xystonotus 365 Alloneumania 231 Aspidiobatopsis 95 alpicola, Nesaea 201 asserculatus, Aturus 14, 20 418 Taxonomic index Acari III

Astacocrotonidae 1 brachypus, Aturus 14, 18, 21 Atacella 236 brachyurus, Arrenurus 330 Athienemanniidae 255, 349 brehmi, Piona 201 atlantida, Neumania 235 brendle, Atractides 123 Atractides 98 brevipes, Erebaxonopsis 50, 61, 62 Aturidae 1, 7, 60, 95, 386 brevipes, Feltria 82 Aturides 9 brevipes, Forelia 181, 183, 184 Aturinae 9 brevipes, Nesaea 183 Aturus 10 Bromeliacarus 251 Australotiphys 213 brteki, Aturus 17, 20, 22 Austromomonia 369 brunsvicensis, Arrenurus 261 Axonopsella 49 bruzelii, Arrenurus 284 Axonopsella-like mites 49 buccinator, Hydrachna 314 Axonopsinae 7, 8, 9, 48 bucculata, Prymnopsella 51 Axonopsis 53, 58 bucharicus, Arrenurus 300 Axonopsis-like mites 49 bullata, Piona 219 Azugofeltria 74, 80, 81, 82, 90 bullatus, Tiphys 178, 214, 215, 216, 219

B C baderi, Brachypoda 68 Caledoniabates 95 baderi, Parabrachypoda 53, 55, 68 calliger, Hygrobates 149, 152 barbarae, Atractides 122 callosa, Kongsbergia 39, 41 barbatulus, Aturus 2, 14, 18, 20, 21 callosa, Neumania 230, 232 Barbaxona 49, 51, 54, 56, 58, 59 callosus Atax 232 Barbaxonella 49, 51, 54, 56, 58, 59 Camposea 95 barsiensis, Megapus 130 carnea, Nesaea 203 Bassatax 237 carnea, Piona 194, 198, 203 batillifer, Arrenurus 282 cassidiformis, Forelia 183 bavaricus, Arrenurus 314 castaneus, Arrenurus 340, 347 bella, Javalbia 51, 64 caucasicus, Atractides 131 berolinensis, Arrenurus 282 cavifrons, Lethaxona 162 Bharatalbia 9 celeripes, Brachypoda 67 bicolor, Axonopsis 58 celeripes, Ocybrachypoda 50, 55, 67 bicuspidator, Arrenurus 282 Centrolimnesia 166 bifidicodulus, Arrenurus 330 cerberus, Atractides 134 bipapillata, Ljania 49, 50, 57, 65 cetrata, Forelia 181, 182 bipapillata subterranea, Ljania 49 cetratus, Acercus 184 bipapillosus, Arrenurus 267, 328, 330 Chaoboridae (Insecta Diptera) 154, 261, 281, biscissus, Arrenurus 331 287, 294, 315 bisulcicodulus, Arrenurus 332 Chappuisides 254, 353 Bleptomundamella 349 Chappuisididae 253, 353 Bogatiidae 255 Chappuisidinae 353 bonzi, Atax 246, 247 Chappuisi, Kawamuracarus 162, 166 bonzi, Unionicola 237, 239, 246, 247 chaunensis, Pionides 218 boopis, Piona 196, 198, 203 chelidon, Atractides 123 borceai, Megapus 144 Chelohydracarus 349 boruzkii, Arrenurus 282 Chelomideopsis 349, 350, 351 Brachypoda 53, 59 Chironomid 25, 33, 58, 65, 71, 73, 82, 86, 87, Brachypoda-like mites 49 127, 133, 152, 153, 157, 172, 173, 183, Brachypodella 68 184, 186, 189, 191, 199, 201, 202, 204, Brachypodopsis 49, 52, 55, 58, 60, 61 205, 207, 209, 211, 212, 213, 219, 222, Taxonomic index Acari III 419

237, 242, 243, 244, 247, 248, 249, 252, cornuta paucipora, Feltria 74 351, 357, 359, 364, 366, 373, 375 coronator, Arrenurus 316 Chironomidae (Insecta Diptera) 59, 121, 132, corsicus, Arrenurus 267, 340, 341, 342, 355 135, 136, 138, 144, 154, 155, 156, 167, corsicus, Atractides 131 190, 206, 210, 233, 257, 261, 281, 287, corsicus, Balcanohydracarus 341 294, 315, 319, 325, 339, 363 crassa, Delmea 199 Chironominae 65, 87, 127, 133, 252 crassa, Nautarachna 194, 199 Chironomini 243, 247 crassicaudatus, Arrenurus 286 Chironomus 202 crassipes, Atax 242, 244 circumcinctus, Atractides 123 crassipes dresscheri, Unionicola 243 cisternarum-species group, Atractides 125 crassipes, Hydrachna 242 Cladocera (Crustacea) 173, 201, 202, 210, 248, crassipes, Mideopsis 362, 363 261, 289, 297, 300, 308, 319 crassipes minor, Atax 244 Cladomomoniinae 367 crassipes octopora, Unionicola 243 clavatus, Arrenurus 316 crassipes reducta, Unionicola 243 clavicornis, Hydrachna 204 crassipes, Unionicola 3 clavicornis, Piona 194, 196, 204 crassus, Unio (Mollusca Bivalvia) 3, 4, 190, claviger, Arrenurus 285 229, 230, 236, 237, 238, 239, 240, 241, clavipes, Atractides 124 242, 243, 244, 245, 246, 247, 248, 386 clipeata, Feltria 74 crenatus, Arrenurus 287 clypeata, Kongsbergia 39, 40, 42 crinitus, Aturus 16, 19, 23, 24 coactus, Curvipes 205 Crustacea 1, 96, 141, 167, 172, 186, 191, 193, coccinea confertipora, Piona 208 203, 204, 205, 212, 219, 221, 233, 237, coccinea, Nesaea 204 243, 251 coccinea, Piona 178, 194, 199, 204, 208, 212 Cubanaxonopsis 49 coccinea recurva, Piona 208 Culicidae (Insecta Diptera) 261, 287, 292, 294, coccinea stjoerdalensis, Piona 212 315, 319, 327, 330, 332, 336, 339, 343, coccinoides, Curvipes 205 346, 347 coccinoides, Piona 199, 205 cultellatus, Atractides 128 cogitata, Brachypodopsis 52, 55 curvilaminatus, Arrenurus 261 cogitatus, Axonopsis 61 curvipalpis, Forelia 185 Coleoptera (Insecta) 210, 261, 281, 311, 319 curvipalpis, Limnesia 170, 171 comatus, Aturus 17, 18, 22 curvisetus, Arrenurus 294, 322 communis, Litarachna 225, 226, 227 cuspidator, Hydrachna 288 commutatus, Arrenurus sinuator 294 cuspidator, Arrenurus 263, 267, 278, 279, 285, compactus, Arrenurus 286 288 complanata, Axonopsis 52, 58 cuspidifer, Arrenurus 280, 288 complanata, Hydrachna 58 cyanipes, Hydrachna 300 complanata, Pseudanodonta (Mollusca Bival- Cyclomomoniinae 367 via) 242 cygnea, Anodonta (Mollusca Bivalvia) 239, 240, Coneumania 231 241, 246, 247, 248, 249 confertipora, Piona coccinea 208 cylindratus, Arrenurus 317, 321 conglobata, Nesaea 205 cylindratus latigenitalis, Arrenurus 321 conglobata, Piona 195, 196, 205 conicus, Arrenurus 316 conjuncta, Feltria 77, 79, 83, 87 D connata, Limnesia 168, 171 danielei, Paraxonopsis 71 conspiciendus, Hygrobates 152, 156, 159 dantei, Axonopsis 70 Copepoda (Crustacea) 210, 248, 261, 297, 319 davidsi, Albia 4, 9 coriaceus, Megapus 144 Decussobates 95 cornuta, Feltria 75, 76, 79, 83 deltoides, Atax 233 cornuta longispina, Feltria 85 deltoides, Neumania 231, 232, 233 420 Taxonomic index Acari III

Dentalbia 8 elliptica, Javalbia 64 dentata, Kongsbergia 35, 39, 42 elliptica, Kongsbergia 39, 43 denticulatus, Arrenurus 265, 274, 289, 290 elliptica, Limnesia 173 denticulatus, Atractides 107, 108, 115, 116, 124, Elodea 244 125, 137 elongatus, Aturus 13, 30 dentiferus, Atractides 129 elongatus, Kawamuracarus 166 dentipes, Piona 201 elongatus, Megapus 147 depressa, Mideopsis 361 elongatus processiger, Aturus 30 detruncatus, Arrenurus 332 Encentridophorinae 229 Diamesinae (Insecta Diptera Chironomidae) 65 ensifer, Piona 218 digitatus, Atractides 125 ensifer, Pionides 192, 213, 215, 218 Diptera (Insecta) 25, 33, 152, 153, 154, 155, ensifer, Tiphys 217 156, 157, 172, 183, 184, 186, 189, 191, Eolethaxona 161 199, 201, 211, 213, 219, 257, 261, 281, Epallagopodinae 165 287, 292, 294, 311, 315, 319, 325, 327, Ephemeroptera (Insecta) 8 328, 330, 332, 336, 339, 340, 343, 346, Erebaxonopsis 49, 50, 61, 62 347, 364, 366, 373, 375 eremita, Javalbia 64 discrepans, Curvipes 206 Esekalla 230 discrepans, Piona 194, 195, 199, 206 Estellacarus 49 disjuncta, Feltria 74 Eubrachypoda 59 disjuncta, Piona rotunda 211 eugeminus, Arrenurus 317 disparilis, Curvipes 206 disparilis, Piona 195, 197, 206 dispersa, Piona 197, 207 F dispersa, Piona variabilis 207 faeroensis, Acercus latipes 221 distans, Arrenurus 276, 278, 289, 290, 291, 295 faeroensis, Tiphys latipes 216, 221 distans, Atractides 110, 112, 119, 121, 125, 128 falcigera, Piona 201 distans inflatipes, Atractides 128 falciger, Arrenurus 267, 268, 273, 274, 291, 292 Dixa (Insecta Diptera Dixidae) 343 falcilaminatus, Hygrobates 154 Dixidae (Insecta Diptera) 261, 319, 343, 347 falcipalpis, Momonia 254, 367, 368, 369 Djeboa 361 farkasi, Stygohydracarus 352 Dockovdia 95 fasciolatus, Arrenurus 317 dorieri, Arrenurus 333 Feltria 73 dragonensis, Atractides 129 Feltriella 75, 76, 78, 80, 89, 90 dramensis, Megapus 135 Feltriidae 73 dresscheri, Unionicola crassipes 243 figuralis, Atax 243 drilonensis, Feltria 83 Fijilimnesia 166 Dubiobates 95 fimbriatus, Arrenurus 292 dubosqi, Litarachna 226, 227 finisbelli, Unionicola 247 Duralimnesia 167 firmus, Megapus 130 duursemai, Arrenurus 291 fish (Pisces) 167, 237, 261, 273, 282, 312, 324 Dytiscidae (Insecta Coleoptera) 319 fissus, Atractides 110, 111, 118, 120, 126, 138 fluviatilis, Acarus 152 fluviatilis, Atractides 113, 114, 117, 118, 126 E fluviatilis, Hygrobates 3, 150, 152, 153 Ecpolopsis 231 fluviatilis, Megapus nodipalpis 126 Ecpolus 230, 236 fonticola, Nudomideopsis 376 ekmani, Axonopsis 66 fonticolus, Atractides 128, 141 ekmani, Neobrachypoda 50, 57, 66 fonticolus gallicus, Atractides 141 elburzensis, Atractides 144 fonticolus, Megapus nodipalpis 127 elegans, Atractides 145 fontinalis, Arrenurus 267, 342 Taxonomic index Acari III 421 fontinalis, Aturus 17, 20, 24 granulosa, Limnesia 170 forcipatus, Mesobates 157, 158, 159 Gretacarinae 361 Forelia 180, 183 Guineaxonopsinae 361 foreli, Campognatha 153 Gyrinidae (Insecta Coleoptera) 319 foreli, Hygrobates 151, 153 gyrophora, Piona 198, 207 Foreliinae 178, 179 forpicatus, Arrenurus 267, 328, 329, 333, 334 fossea, Feltria 89 H franciscanus, Atractides 140 Haloaxonopsis 1, 7, 49 freemani, Arrenurus 325 Halolimnesia 167 Frontipodopsella 93 Hamappendix 259 Frontipodopsidae 93 handschini, Feltria 88 Frontipodopsinae 93 hankoi, Unionicola 245, 246 Frontipodopsis 93 hansvietsi, Kongsbergia 10, 11 fulgida, Limnesia 167, 170, 172 haplurus, Arrenurus 267, 340, 341, 343, 344 furciger, Arrenurus 290 helocrenicus, Hygrobates 157 furcillatus, Arrenurus 264, 272, 292, 293 helvetica, Lethaxona 163 furva, Oecetis (Insecta Trichoptera) 243 Hemibrachypoda 68 fuscatus, Arrenurus 261 Hesperomomonia 369 fusiformis, Kongsbergia 35 Heteratax 236 Heterolimnesia 167 heteropora, Limnesia 174 G heversi, Atractides 125 gallica, Lethaxona 163 Hexatax 238, 242 gallicus, Atractides fonticolus 141 Hexaxonopsalbia 49, 51, 55, 56, 57, 59, 62 gallicus, Aturus 14, 20, 25 Hexaxonopsis 49, 50, 52, 58, 61, 63 Geayia 357 hibernicus, Neoacarus 255, 368, 373 geminus, Arrenurus 317 Hjartdalia 43 georgei, Arrenurus 347 hoffmannae, Pontarachna 226 Gereckea 190 hofsteni, Arrenurus 294 gereckei, Xystonotus 365 Homtinibates 95 germanica, Mundamella 350, 351 Hopkinsobates 95 gibberipalpis, Atractides 127 Huitfeldtia 177, 178, 190 gibberipes, Forelia 179 Huitfeldtiinae 177, 190 giustinii, Atractides 130 hungaricus, Chappuisides 254, 353 gladiator, Acercopsis 217 Hungarohydracarinae 355 glandulosus, Atractides 122 Hungarohydracarus 255, 355, 356 glandulosus, Megapus 127 Hydrachna 44, 58, 60, 173, 174, 184, 186, 204, globator, Arrenurus 261, 311 209, 220, 222, 225, 230, 234, 235, 242, globator, Hydrachna 318 280, 288, 295, 300, 301, 307, 314, 318, golatensis, Feltria 74 326, 335, 339, 347, 359, 361, 363, 377, gorcensis, Atractides 128 378, 379, 380, 381, 382, 384, 385, 387, gorgani, Atractides 130 388 gracilipalpis, Atax 244 Hydrochoreutes 191, 192, 193 gracilipalpis parvipora, Unionicola 245 Hydrochoreutinae 177, 191 gracilipalpis, Piona stjördalensis 208 Hygrobates 96, 149 gracilipalpis, Unionicola 245 Hygrobatidae 1, 95, 386 gracilis, Albaxona 56 Hygrobatides 1 gracilis, Brachypodopsis 52, 55, 60, 61 Hygrobatinae 95 graecus, Arrenurus ornatus 300 Hygrobatoidea 1, 2, 4, 95, 225, 253, 383 graeca, Axonopsis 58 Hygrobatomegapus 149, 157 422 Taxonomic index Acari III hyporheicus, Atractides 136 karamani, Pionella 199, 200 hyrcaniensis, Atractides 127 karelica, Kondia 370 hystricipes, Atractides 98, 99 karelica, Momonia 368 karelicus, Arrenurus 329, 330, 336 Karlvietsia 95 I Kawamuracarinae 165 Iberaxonopsalbia 64 Kawamuracarus 162, 165, 166 ibericus, Atractides 148 Khedacarus 49 ildensis, Atractides 135 kjerrmanni, Arrenurus 267, 276, 278, 294 imamurai, Unionicola 247 knauthei, Arrenurus 268, 340, 341, 343, 344 imitata, Neumania 233 Knysnabates 95 imitator, Arrenurus 317 kochi, Unionicola 244 imminuta, Piona 194, 199, 208 Koenikea 229, 230 imminutus, Curvipes nodatus 208 koenikei, Acercus 186 inconstans, Feltria 75, 76, 79, 81, 84 koenikei, Limnesia 168, 172 incurvatus, Mixobates 158, 159 Kondia 367, 370 inexploratus, Arrenurus 262, 272, 330, 334, Kongsbergia 10, 34 335, 337, 338, 339 krameri, Hydrochoreutes 191, 192 inferorum, Axonopsis 4, 70 Krendowskia 209, 254, 255, 300, 350, 357 inferorum, Paraxonopsis 53, 54, 70 Krendowskiella 357 inflata, Piona 208 Krendowskiidae 254, 357 inflatipes, Atractides distans 128 Krendowskiinae 357 inflatus, Atractides 107, 109, 117, 118, 129 kurui, Arrenurus 308 inflatus, Megapus 129 inflexus, Arrenurus 319 informis, Arrenurus 311, 320 L insolita, Feltria 74, 81 lacustris, Atractides 146, 147, 148 integrator, Arrenurus 269, 270, 330, 335 lacustris lacustris, Megapus 148 integrator, Hydrachna 335 lacustris, Megapus lacustris 148 intermedia, Atax 241 lacustris scutatus, Megapus 148 intermedia, Unionicola 240 laetus, Atractides 139 intermedius, Aturus 17, 20, 26, 31 lamelligerus, Acercus 220 intermedius serrata, Aturus 20 laminata, Piona 197, 208 interruptus, Arrenurus 276, 293, 294 laminatus, Curvipes 208 inusitata, Unionicola 247 lapponica, Piona 220 Ioannibates 95 lapponicus, Arrenurus 270, 312, 313, 320, 321 Iranobates 95 lapponicus, Tiphys 214, 215, 220 italicus, Hungarohydracarus subterraneus 356 Largaiollii, Hjartdalia 43 largaiollii, Kongsbergia 39, 41, 43 Larri 190 J latigenitalis, Arrenurus 269, 312, 320, 321 jaczewskii, Limnesia 171 latigenitalis, Arrenurus cylindratus 321 jaentschi, Kongsbergia 43 latimembranosus, Arrenurus zachariae 327 Javalbia 49, 51, 55, 56, 64 latipalpis, Atractides 106, 107, 115, 129 Javalbiella 49 latipalpis, Megapus 129 jellyfish (Hydrozoa) 225 latipalpis, Nudomideopsis 256, 368, 375 latipes, Atractides 107, 109, 113, 114, 129 latipes faeroensis, Acercus 221 K latipes faeroensis,Tiphys 216, 221 kamchatkaensis, Tiphys 220 latipes, Hydrachna 220 karamani, Aturus 14, 18, 26 latipes latipes, Tiphys 216, 220 Taxonomic index Acari III 423 latipes, Stygomomonia 367, 368, 370, 371 losonensis, Atractides 137 latipes,Tiphys 215, 216, 220, 221 lundbladi, Albaxona 51, 56 latissima, Krendowskia 255, 350, 357 lundbladianus, Arrenurus 341 latus, Arrenurus 294 lundbladi, Mixobates 158 lautieni, Axonopsalbia 62 lundbladi, Momonides 367, 368 lautieni, Hexaxonopsalbia 51, 55, 57, 62 lundbladi, Vietsaxona 56 Lebertioidea 1, 383 lunipes, Atractides 131 legeri, Atractides 102, 103, 110, 111, 117, 118, lutescens, Hydrarachna 218 130 lutescens, Pionopsis 192, 215, 218 lelgioensis, Aturus 27 Lemienia 231 Leptopterotrichophorus 231 M Lethaxona 1, 2, 7, 161, 162, 163 macilenta, Ljania 49, 57, 65, 66 Lethaxonella 161 macilenta longissima, Ljania 65 Lethaxonidae 1, 7, 161, 162 macrolaminatus, Atractides 131 leuckarti, Arrenurus 295 macroplicus, Wettina 251 leuckarti, Pionacercus 179, 180, 181, 187 maculata angustata, Limnesia 173 levantensis, Krendowskia 357 maculata, Hydrachna 173, 295 ligulifer, Acercus 183 maculata, Limnesia 169, 173 liliacea americana, Forelia 184 maculata viridis, Limnesia 173 liliacea, Hydrachna 184 maculator, Arrenurus 267, 278, 279, 288, 290, liliacea liliacea, Forelia 182, 183, 184 295, 296 Limnesia 165, 167 maculator, Hydrachna 295 Limnesiella 167 Madawaska 179 Limnesiellula 167 Maderomegapus 95, 98, 99 Limnesiidae 1, 165, 171, 174, 388 magnipalpis, Atractides 132 Limnesiinae 165, 166 Maharashtracarinae 95 Limnesiopsides 167 Majumderatax 239, 245, 246 Limnesiopsis 167 markaziensis, Atractides 136 limnicola, Atractides 133 marmorata, Limnesia 169, 173 limosa, Neumania 231, 232, 233 marsus, Atractides 132 limosus, Atax 233 martianezi, Limnesia 170 Ljania 49, 50, 57, 65, 66, 380 materna, Kongsbergia 11, 34, 39, 41, 44 longipalpis, Forelia 181, 182, 183, 184, 185 mathiasi, Arrenurus 295 longipalpis, Hydrarachna 153 maxillaris, Bogatia 255, 256 longipalpis, Hygrobates 151, 152, 153 mediterranea, Axonopsalbia 50 longipalpis, Nesaea 209 Megaluracarus 261, 311 longipalpis, Nudomideopsis 375, 376 Megapus 96 longipalpis, Piona 194, 195, 198, 199, 209 membranator, Arrenurus 322 longipes, Atractides 134 menzeli, Feltria 90 longipes, Megapus 147 Mesobates 96, 157, 158, 159 longipes, Mesobates 158 Micrarrenurus 259 longiporus, Hygrobates 154 microcavaticus, Atractides 133 longirostris, Atractides 149 microphtalmus, Atractides 125 longisetus, Atractides 126 micropora, Lethaxona 163 longispina, Feltria 85 Micruracaropsis 259 longissima, Ljania 49, 57, 65 Micruracarus 261, 327 longissima, Ljania macilenta 65 Midea 254, 359, 362 longitibialis, Megapus 135 Mideidae 253, 359 longus, Megapus 134 Mideopsella 361 loricatus, Atractides 130 Mideopsellides 361 424 Taxonomic index Acari III

Mideopsellinae 361 N Mideopsidae 255, 361 nagysalloensis, Arrenurus 340, 341, 344 Mideopsides 361 naias, Arrenurus stjordalensis 316 Mideopsinae 361 Najadicolinae 177, 229 Mideopsis 361 natangensis, Aturus 17, 20, 27 minuta, Albaxona 56 Nautarachna 177, 193, 194, 199 minuta, Feltria 73 Nematocera (Insecta Diptera) 154 minuta, Woolastookia 71 Neoacaridae 253, 373 minutiporus, Tiphys 220 Neoacarus 255, 368, 373 minutissima, Feltria 90 Neoalbia 49 mira, Feltria 74, 81 Neoatax 237 Miraxona 49 Neoaturus 9 Miraxonides 49 Neoaxona 49 mitisi, Megapus 135 Neoaxonopsalbia 49 Mixobates 95, 96, 149, 158, 159, 160 Neoaxonopsis 49 Mixolimnesiinae 165 Neobarbaxona 49 Mixomideopsis 365 Neobrachypoda 49, 50, 53, 57, 66 modesta, Brachypoda 69 Neochappuisides 353 modesta, Parabrachypoda 50, 53, 55, 67, 69 Neofeltria 73, 90 Momonia 253, 254, 367, 368, 369, 370 Neokrendowskia 357 Momonides 367, 368 Neomamersinae 165 Momonidinae 367 Neomideopsis 375 Momoniella 369 Neomomonia 369 Momoniidae 253, 367 Neoneumania 231 Momoniinae 367 Neotiphys 213 Momonisia 367, 368 Neotorrenticolinae 165 moniezi, Megapus 148 Neoxystonotus 361 monstrabilis, Axonopsis 70 Neumania 229, 230 montanus, Atractides 99 neumani, Arrenurus 298 montenegrinus, Nudomideopsis 375 neumani, Atractides nodipalpis 132 montii, Brachypoda 69 Neumanii, Nesaea 209 montii, Parabrachypoda 53, 54, 69 neumani, Piona 195, 199, 209 Moramangabates 95 Nicalimnesiinae 165 moravicus, Atractides 133 Nicksatax 231 Morimotacarinae 353 nielseni, Arrenurus 298 mosticus, Phreatohydracarus 350 nigromaculatus, Hygrobates 154, 155 motasi, Feltria 74, 80 nigromaculatus setosus, Hygrobates 155 motasi, Nudomideopsis 375 nitraensis, Atractides 134 muelleri, Arrenurus 270, 312, 314, 315, 322, nodata, Hydrachna 209 323, 325 nodata, Piona 194, 197, 209 muenchbergi, Arrenurus 266, 277, 278, 296, 297 nodatus imminutus, Curvipes 208 mülleri spatiosus, Arrenurus 325 nodipalpis, Atractides 122, 132 multiporus, Hungarohydracarus 356 nodipalpis fluviatilis, Megapus 126 Mundamella 349, 350, 351 nodipalpis fonticolus, Megapus 127 mussels (Mollusca Bivalvia) 237, 240, 241, 242, nodipalpis neumani, Atractides 132 244, 246, 247, 248, 249 nodipalpis parviporus, Megapus 133 mutila, Brachypoda 68 nodipalpis pennatus, Megapus 135 nodipalpis robustus, Megapus 139 nodosus, Arrenurus 345 norvegicus, Hygrobates 156 norvegicus, Pionacercus 179, 180, 181, 188 Taxonomic index Acari III 425

Notoaturinae 7 P Notomomonia 369 Paenecalyptonotus 257 Notomundamellinae 349 palpalis, Atractides 142 novus, Arrenurus 336 panniculatus, Atractides 112, 117, 118, 126, Nudomideopsidae 255, 375, 376 135, 138 Nyangalla 229 panniculatus, Megapus 135 papillator, Arrenurus 261 papillator, Hydrachna 300 O papillosa, Neumania 236 oblongus, Atractides 102, 103, 117, 118, 134, papillosus, Ecpolus 236 143 Parabrachypoda 53, 68 oblongus, Megapus 134 paradoxa, Brachypoda 59 obturbans, Curvipes 210 Parakongsbergia 10, 11, 35 obturbans, Piona 195, 197, 210 Paralimnesia 167 octagonus, Arrenurus 336 Paramideopsis 375 Octomegapus 96 Paramomonia 369 Octomideopsis 361 Parasitalbia 8 octopora, Unionicola crassipes 243 Parasitatax 237, 242 octoporus, Atractides 134 Paraxonopsis 49, 50, 53, 54, 70, 71 Ocybrachypoda 49, 50, 53, 55, 60, 67 Partidomomonia 369 Odonata (Insecta) 261, 272, 280, 282, 284, 286, parvipora, Unionicola 238, 240, 241, 245 288, 289, 290, 293, 294, 295, 297, 298, parvipora, Unionicola gracilipalpis 245 299, 300, 302, 304, 307, 308, 309 parviporus, Megapus nodipalpis 133 oedipoda, Feltria 78, 80, 81, 86 parvula, Unionicola 245 Omanaxonopsis 49 paucipora, Feltria cornuta 74 Omanohydracarinae 229 paucipora, Piona 196, 199, 210 Omartacaridae 1 pauciporus, Aturus 26 orbicularis, Hydrachna 361, 363 pauciporus, Curvipes 210 orbicularis, Mideopsis 256, 362, 363 paucisetus, Aturus 14, 18, 22, 28 orbiculata, Hydrachna 359 pavesii, Atractides 148 orbiculata, Midea 254, 359, 362 pectinata, Kongsbergia 35, 39, 45 orghidani, Atractides 145 pectinigera, Kongsbergia 11, 35, 39, 45 Orientmomonia 369 Penemideopsis 361 ornatulus, Arrenurus 286 Penemundamella 349 ornatus, Arrenurus 286, 300 pennatus, Atractides 108, 109, 117, 118, 128, ornatus, Tiphys 178, 214, 215, 216, 221 135 ornatus graecus, Arrenurus 300 pennatus, Megapus nodipalpis 135 Orthocladiinae (Insecta Diptera Chironomi- Pentatax 237, 239, 242, 246, 247, 248 dae) 65, 87, 127, 133, 252 perforatus, Arrenurus 337 Osornobates 95 persicus, Atractides 146 Ostracoda (Crustacea) 201, 226, 227, 261, 287, petkovskii, Atractides 127 289, 300, 319 Phragmites (bot.) 212 oudemansi, Aturus 26 phreatica, Momonisia 367, 368 ovalis, Atractides 134 phreaticola, Feltria 84 ovalis, Barbaxonella 58 phreaticola, Neumania 230, 234 ovata, Nesaea 201 phreaticus, Atractides 106, 107, 114, 115, 136 oviformis, Atractides 143 Phreatobrachypoda 9 oviformis, Unionicola 237 Phreatomideopsinae 361 Phreatomideopsis 361 Physolimnesia 166 426 Taxonomic index Acari III pictorum, Unio (Mollusca Bivalvia) 3, 4, 190, Pseudaxonopsalbia 49 229, 230, 236, 237, 238, 239, 240, 241, Pseudoaustraliobates 95 242, 243, 244, 245, 246, 247, 248, 386 Pseudofeltria 2, 178, 180, 181, 186, 188, 189 piersigi, Feltria 74 Pterolimnesia 166 Pilolimnesia 167 pugionipalpis, Feltria 74, 81 pilosa, Barbaxonella 59 pulchellus, Aturus 17, 31 pilositarsa, Brachypoda 71 pumilus, Atractides 101, 106, 113, 132, 137 pilosus, Atractides 142 punctata, Nesaea 205 Piona 60, 193, 201 punctulum, Pontarachna 3, 225, 226 Pionacercopsis 179, 180, 181, 186, 189 pusilla, Kongsbergia 42 Pionacercus 179, 180, 181, 186, 187, 188, 189 pusilla, Nesaea 211 Pionatacinae 229 pusilla, Piona 196, 197, 211 Pionatax 229 pustulator, Arrenurus 262, 263, 272, 301 Pionella 199, 200 pustulator, Hydrachna 301 Pionidae 5, 7, 177, 386 pygmaea, Lethaxona 2, 161, 162, 163 Pionides 192, 213, 215, 217, 218 pygmaeus, Atractides 107, 108, 115, 116, 138 Pioninae 178, 193 pyrenaeus, Arrenurus 302 Pionopsis 192, 213, 215, 218 pistillifer, Acercopsis 213, 214, 216, 217 pistillifer stylatus, Acercus 217 Q pistillifer, Tiphys 216 quadriscutata, Pseudofeltria 181, 189 placocallosus, Atractides 144 Platyhydracarus 349 Plaumanniinae 361 R Plecoptera (Insecta) 86 radiatus, Arrenurus 302 Plesiobrachypoda 63 raetica, Feltria 83 podagrica, Wettina 4, 241, 251 Raptorhydracarinae 165 Pollicipalpinae 229 Recifella 230 polonica, Limnesia 170, 174 rectipes, Huitfeldtia 178, 190 Polyatax 237 recurva, Piona coccinea 208 Polyaxonopsella 49 reducta, Unionicola crassipes 243 Polymegapus 98, 99, 144, 145, 146 refractariolus, Arrenurus 303 polyporus, Atractides 98, 144, 145, 146 regulator, Arrenurus 270, 271, 340, 341, 345, Pontarachnidae 1, 225, 226, 379, 381 346 porrectus, Hygrobates 149, 153 regulus, Arrenurus 288 prenanti, Aturus 14, 29 remotus, Atractides 138 primitivus, Megapus 137 remyi, Megapus 129 Proboscibates 95 reticulatifrons, Frontipodopsis 2 processifer, Hygrobates 158, 160 Rheolimnesiinae 165 processifer, Mixobates 158, 159, 160 Rhomborificias 259 processiger, Aturus 17, 30, 33 Rhynchaturus 95 processiger, Aturus elongatus 30 rivalis, Atractides 139 properus, Hygrobates 156 Rivobates 149, 152, 156, 157 prosiliens, Atractides 107, 108, 136 robustus, Arrenurus 304 protendens, Atractides 137 robustus, Atractides 139 Protoarrenurus 259 robustus, Megapus nodipalpis 139 Protolimnesiinae 165 rodrigensis, Arrenurus 304 protzi, Aturus 17, 20, 30 romijni, Axonopsis 63 psammocrenicus, Hygrobates 157 romijni, Hexaxonopsis 50, 52 Psammolimnesiinae 165 Roqueella 357 psammophilus, Atractides 134 rotunda disjuncta, Piona 211 Taxonomic index Acari III 427 rotundifrons, Axonopsis 71 Sinaxonopsis 49 rotundifrons, Woolastookia 50, 57, 71 sinearcus, Thoracophoracarus 259 rotundoides, Curvipes 211 Sinhaladwipabates 95 rotundoides, Piona 196, 197, 211 sinuator, Arrenurus 268, 294, 329, 338, 339 rotundus, Aturus 11, 14, 17, 31, 32 sinuator commutatus, Arrenurus 294 rotundus, Aturus scaber 31 sinuator, Hydrachna 339 rouxi, Feltria 77, 78, 79, 87 Soarella 231, 236 roztoczensis, Mideopsis 362, 363, 364 sokolowi, Atractides 103, 113, 141 rubra, Feltria 89, 90 soproniensis, Megapus 127 rutae, Atractides 145 spatiosus, Arrenurus 312, 313, 324, 325 ruttneri, Kongsbergia 36, 41, 46 spatiosus, Arrenurus mülleri 325 spatulifer, Acercus 185 spatulifera, Forelia 182, 183 S spatulifer, Aturus 17, 18, 33 sagulatus, Tiphys 219 Spinalbia 8 salmonids (Pisces) 66 spinipes, Atractides 141 Salvelinus (Pisces) 66 spinipes, Hydrachna 230, 234 samaricus, Atractides 135 spinipes, Megapus 96 samsonellus, Atractides 140 spinipes, Neumania 230, 232, 234 samsoni, Atractides 140 sponges (Porifera) 237, 242, 243, 244, 245 scaber, Aturus 10, 11, 14, 17, 31, 32 staheli, Frontipodopsis 93 scaber rotundus, Aturus 31 stammeri, Arrenurus 346 scaura, Piona 222 stankovici, Atractides 122 scaurellus, Tiphys 221 stationis, Albia 4, 8 scaurus, Tiphys 215, 216, 222 stecki, Arrenurus 346 Schadeella 229 Sterkspruitia 95 schlienzi, Atractides 133 stjoerdalensis, Piona 212 schlienzi, Hygrobates 155 stjoerdalensis, Piona coccinea 212 schreuderi, Arrenurus 343 stjordalensis, Arrenurus 316, 321 scourfieldi, Arrenurus 271, 314, 323, 324 Stjørdalensis, Curvipes 212 scourfieldi, Pseudofeltria 178, 180, 181, 188, stjördalensis gracilipalpis, Piona 208 189 stjordalensis naias, Arrenurus 316 sculptus, Arrenurus 270, 271, 330, 331, 338 stjordalensis, Piona 194, 199, 201, 204, 212 scutatus, Megapus lacustris 148 Stokaxonopsis 49 scutifera, Feltria 74 Stormaxonella 251 securifer, Arrenurus 305 Stygalbiella 49 securiformis, Arrenurus 324 Stygamericarinae 349 sejugata, Piona 193 Stygohydracarus 255, 349, 350, 352 separatus, Atractides 105, 106, 113, 114, 140 Stygolimnesiinae 165 Seppiella 167 Stygomomonia 367, 368, 370, 371 serrata, Aturus intermedius 20 Stygomomoniinae 367, 370 serrata, Axonopsis 63 stygophila, Feltria 88 serrata, Hexaxonopsis 52 stygophilus, Atractides 138 setigera, Feltria 74, 75, 76, 79, 85, 88 stylatus, Acercus pistillifer 217 setosus, Hygrobates 155 Subalbia 9 setosus, Hygrobates nigromaculatus 155 subarcticus, Arrenurus 263, 272, 305, 306 seurati, Neumania 235 subasper, Atractides 142 similis, Atractides 107, 108, 115, 116, 140 Subaturus 9 similis, Kongsbergia 46 Subaxonopsalbia 49 simillima, Kongsbergia 39, 41, 46 Submiraxona 49 simplicipes, Kongsbergia 39, 47 Subneumania 231 428 Taxonomic index Acari III subterranea, Feltria 76, 77, 79, 88 tumidus, Unio (Mollusca Bivalvia) 3, 4, 190, subterranea, Frontipodopsella 93 229, 230, 236, 237, 238, 239, 240, 241, subterranea, Ljania bipapillata 49 242, 243, 244, 245, 246, 247, 248, 386 subterraneus, Atractides 104, 105, 116, 142 turgidus, Arrenurus 265, 277, 278, 308, 309 subterraneus, Hungarohydracarus 255, 355, Tympanomegapus 98, 146, 147, 148 356 Tyrreliinae 165 subterraneus italicus, Hungarohydracarus 356 subterraneus, Stygohydracarus 255, 350, 352 suecicus, Arrenurus 267, 268, 272, 273, 306 U Sumatralbia 49 ubinicus, Atractides 132 svislocensis, Atractides 133 uchidai, Pionides 218 Szalayella 95 Uchidastygacaridae 353 szalayi, Megapus 134 Uchidastygacarinae 353 Uenaxonopsis 49 uncinata, Neumania 235 T uncinatus, Acercus 188 Tanypodinae (Insecta Diptera Chironomi- uncinatus, Pionacercus 179, 180, 181, 188 dae) 281, 287, 294, 315, 319, 325, 339 undulata, Hydrachna 174 Tasmanaxona 251 undulata, Limnesia 170, 171, 175 tatrensis, Atractides 143 undulatoides, Limnesia 170, 175 tener, Atractides 124, 141 ungulata, Spio 193 tenerifensis, Atractides 124 ungulatus, Hydrochoreutes 191, 192, 193 tenuitarsus, Megapus 133 Unio (Mollusca Bivalvia) 3, 4, 190, 229, 230, tetracyphus, Arrenurus 306 236, 237, 238, 239, 240, 241, 242, 243, Tetralimnesia 167 244, 245, 246, 247, 248, 386 Tetramegapus 96 Unionicola 229, 236, 237, 248 Tetraneumania 231 Unionicolella 237 Thoracophoracarus 259 Unionicolidae 4, 190, 229, 386 Timmsilimnesia 166 Unionicolides 237 Tiphyinae 177, 213 Unionicolinae 229, 236 Tiphys 213, 219 ursulae, Feltria 85 Tiramideopsis 361 torris, Tiphys 214, 216, 222 Transitia 161 V transversaria, Stygomomonia 371 Vagabundia 49 trapeziformis, Atractides 143 vaginalis adnatus, Atractides 121 Trichoptera (Insecta) 86, 173, 243, 369 vaginalis, Atractides 121 tricuspidator, Arrenurus 265, 276, 307, 309 vandeli, Atractides 136 tricuspidator, Hydrachna 307 vanrensburgi, Pseudofeltria 186 tricuspis, Atax 248 vardaricolus, Kawamuracarus 166 tricuspis, Unionicola 238, 239, 240, 248 variabilis dispersa, Piona 207 trigonicus, Hygrobates 156 variabilis, Nesaea 212 troglobius, Stygohydracarus 352 variabilis, Piona 196, 199, 207, 212 truncatella, Hydrachna 347 variegator, Arrenurus 186 Truncaturus 261, 340 variegator, Forelia 178, 181, 182 Tsushimacarinae 353 vatrax, Acercus 186 tuberifera, Piona 211 vatrax, Pionacercopsis 179, 180, 181, 186 Tubophora 166 vavrai, Arrenurus 266, 272, 273, 308, 309 Tubophorella 166 vernalis, Acercopsis 217 tubulator, Arrenurus 262, 313, 314, 319, 326, 327 vernalis, Hydrachna 235 tubulator, Hydrachna 326 vernalis, Neumania 231, 232, 235 Taxonomic index Acari III 429 verrucosa, Neumania 236 ypsilophorus, Acarus 248 verrucosus, Atax 235 ypsilophorus, Atax 237 versicolor, Brachypoda 54, 59, 60 versicolor, Hydrachna 58, 60 Vicinaxonopsis 49, 50, 52 Z Vicinihydracarus 352 Zabobates 95 Vietsatax 236 zachariae, Arrenurus 327 Vietsaxona 56 zachariae latimembranosus, Arrenurus 327 vietsi, Arrenurus 261 zschokkei, Feltria 76, 77, 80, 81, 89 vietsi, Axonopsis 70 Zygoptera (Insecta Odonata) 272, 280, 282, 284, vietsi, Kongsbergia 37, 39, 47 286, 288, 289, 290, 293, 295, 297, 298, vietsi, Neumania 231 300, 302, 304, 308, 309 vietsi, Paraxonopsis 50, 53, 70 villosus, Aturus 17, 20, 34 violacea, Axonopsis 63 violacea, Brachypoda 63 violaceus, Acalyptonotus 254, 257 virens, Arrenurus 266, 274, 309, 310 virginalis, Atractides 148 viridis, Arrenurus 259 viridis, Limnesia maculata 173 Viviparus (Mollusca Gastropoda) 247

W walkanoffi, Arrenurus 329, 330, 338, 339 walteri, Atractides 103, 115, 116, 144 walteri, Kongsbergia 38, 39, 40, 48 walteri, Limnesia 168, 170, 175 wereschtschagini, Arrenurus 259, 263, 272, 273, 310 Wettina 4, 5, 161, 241, 251, 252 Wettini 4, 5, 161, 241, 251, 252 Wettinidae 5, 161, 251, 252 Wheenyella 251 Wheenyoides 251 willmanni, Xystonotus 362, 365 wolgaensis, Hydrochoreutes 192 Woolastookia 49, 50, 53, 57, 71 Wuria 259

X Xenolimnesia 166 Xenomomonia 369 Xystonotus 361, 362, 365

Y yaroslavlensis, Tiphys 219 ypsilophora, Unionicola 230