Madre de Dios, PERÚ 1 Dung () of the Amarakaeri Communal Reserve Gorky Valencia1, Alfonso Alonso2, Rolando Coronel1 and Tremaine Gregory2 1Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016

1 Ateuchus connexus 2 Canthidium basipunctatum 3 Canthidium lentum 4 Canthidium onitoides

SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

5 Canthon aberrans 6 Canthon aequinoctialis 7 Canthon femoralis 8 Canthon fulgidus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

9 Coprophanaeus larseni (M) 10 Coprophanaeus telamon (F) 11 Coprophanaeus telamon (M) 12 Deltochilum amazonicum SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

13 Deltochilum carinatum 14 Deltochilum howdeni 15 Deltochilum larseni 16 Deltochilum orbiculare (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

17 Deltochilum orbiculare (M) 18 Deltochilum peruanum (F) 19 Deltochilum peruanum (M) 20 Dendropaemon attalus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

Madre de Dios, PERÚ 2 Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve Gorky Valencia1, Alfonso Alonso2, Rolando Coronel1 and Tremaine Gregory2 1Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016

21 Dichotomius batesi 22 Dichotomius cf. fonsecae (M) 23 Dichotomius mamillatus (F) 24 Dichotomius mamillatus (M)

SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

25 Dichotomius ohausi (F) 26 Dichotomius ohausi (M) 27 Dichotomius prietoi 28 Dichotomius robustus SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

29 Dichotomius worontzowi 30 Eurysternus caribaeus 31 Eurysternus foedus 32 Eurysternus hypocrita SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

33 Eurysternus lanuginosus 34 Eurysternus nigrovirens 35 Eurysternus plebejus 36 Eurysternus vastiorum SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

37 Eurysternus wittmerorum 38 Ontherus alexis 39 Onthophagus haematopus (F) 40 Onthophagus haematopus (M) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

Madre de Dios, PERÚ 3 Dung Beetles (Scarabaeinae) of the Amarakaeri Communal Reserve Gorky Valencia1, Alfonso Alonso2, Rolando Coronel1 and Tremaine Gregory2 1Museo de Historia Natural, Universidad Nacional San Antonio Abad del Cusco, 2Smithsonian Conservation Biology Institute (SCBI). Photos: Gorky Valencia. Produced by: Sulema Castro. Thanks to: SERNANP, ECA-RCA, Juan Carlos Altamirano, Sulema Castro, Jessica Deichmann, Erica Dholoo, Agustín Fernández Zumaeta, Liber García, Wenister García, Guillermo Knell, Johny Nayori Menkorie, Raúl Quenticuari, Santos Quispe and David Federico Vargas. Funding: SCBI, Hunt Oil Exploration and Production Company of Peru. (F) Female, (M) Male. Tick marks in images indicate 1 millimeter. © All rights reserved to the Smithsonian Conservation Biology Institute [fieldguides.fieldmuseum.org] [778] version 1 06/2016

41 Onthophagus onorei (F) 42 Onthophagus onorei (M) 43 Onthophagus osculatii (F) 44 Onthophagus osculatii (M)

SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

45 Onthophagus rubrescens (F) 46 Onthophagus xanthomerus (F) 47 Onthophagus xanthomerus (M) 48 Oxysternon conspicillatum (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

49 Oxysternon conspicillatum (M) 50 Oxysternon silenus (F) 51 Oxysternon silenus (M) 52 Oxysternon spiniferum (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

53 Oxysternon spiniferum (M) 54 bispinus (F) 55 Phanaeus bispinus (M) 56 Phanaeus cambeforti (F) SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE

57 Phanaeus cambeforti (M) 58 Scybalocanthon aereus 59 Sylvicanthon bridarollii 60 Uroxys gorgon SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE SCARABAEIDAE