Gontents

ix ...... ' Prefacetovolume5. xYil Contributors nil On using the Manual Part A I Road map of the phylum ' ' æ the phylum Actinobacteria ' ' Taxonomic outline of 33 phyl' nov' Phylum XXYI. Actinobacteria 34 . - - Class l. Actinobacteria 35 Order l. ActinomYcetales' ' 36 l. ActinomYcetaceae Family 4i2 Genus l. ActinomYces 1(p Genus ll. Actinobaculum . 1t4 Genus lll. Arcanobacterium 16 lY. Mobiluncus. . . . Genus 139 Genus Y. Varibaculum ord. nov' ldl Order ll. Actinopolysporales 169 Family l. ActinoPolYsPoraceae r63 l. ActinoPolYsPora . Genus ' ' 1?to Order lll. Bifidobacteriales lTl L Bifidobacteriaceae Family 171 Genus L Bifidobacterium . âË Genus ll. Aeriscardovia - - - Nf Genus lll. Alloscardovia . - - N Genus lY. Gardnerella ' ' - tll Genus Y. Metascardovia' . - ztg Genus YL Parascardovia 211 Genus Yll. Scardovia zË Order lV. Catenulisporales ord. nov' 26 Family l. CatenulisPoraceae z6 Genus L CatenulisPora- - - - Ên Family ll. ActinosPicaceae - ap, Genus l. ActinosPica É6 OrderV. Corynebacterialesord' nov' ' ' 2tu Family l. Corynebacteriaceae - 2Æ Genus l. CorYnebacterium æ Genus ll.Turicella. 3tl Family ll. Dietziaceae - - - . fln Dietzia Genus L 312 Family lll. MYcobacteriaceae 3t2 Genus l. MYcobacterium .

d xtl CONTENTS

Family lY. Nocardiaceae. . 376 Genus L Nocardia. 376 Genus ll. Gordonia 419 Genus lll. Millisia 435 Genus lY. Rhodococcus. . 437 Genus Y. Skermania 464 Genus Y L S maragdicoccus 467 Genus Yll. Williamsia. . . . 470 Family V. Segniliparaceae. 497 Genus l. Segniliparus 497 Family Y L Tsu kamu rellaceae 500

Genus L Tsukamurella. . . . 500 Order Vl. Frankiales ord. nov 509 Family L Frankiaceae . . . 512 Genus L Frankia 512 Family ll. Acidothermaceae 520 Genus l. Acidothermus. . . . 520 Family lll. Cryptosporangiaceae 522 Genus L Cryptosporangium 522 Genus lncertae sedis l. Fodinicola 525 Family lY. Geodermatophilaceae . . . . 528 Genus L Geodermatophilus 528 Genus ll. Blastococcus. . . 531 Genus lll. Modestobacter. 536 Famif y Y. N akamu rel laceae. 539 Genus l. Nakamurella. . . . 540 Genus ll. Humicoccus . . . 542 Family Yl. Sporichthyaceae. . 544 Genus L Sporichthya 544 OrderVlf . Glycomycetalesord. nov. . . 546 Family l. Glycomycetaceae. 547 GenusLGlycomyces.... 547 Genus ll. Stackebrandtia . 553 Order Vlll. Jiangellales ord. nov. 555 Family L Jiangellaceae . . 557 Genus L Jiangella. 557 Genus ll. Haloactinopolyspora 560 Order lX. Kineosporiales ord. nov. . . 561 Family L Kineosporiaceae. . 562 Genus L Kineosporia 562 Genus ll. Kineococcus. . . 565 Genus lll. Quadrisphaera. 567 Order X. Micrococcales. . . 569 Family l. Micrococcaceae . 571 Genus l. Microcoecus. . . . 571 Genus ll. Acaricomes. . . . 577 Genus lll. Arthrobacter. . . 578 Genus lV. Citricoccus. . . . 624 Genus Y. Kocuria 626 Genus Yl. Nesterenkonia . 636 Genus Yll. Fienibacterium 643 Genus Ylll. Rothia 646 Genus lX. Yaniella ti5o Genus X. Zhihengliuella. . 653 CONTENTS xiii

Family ll. Beutenbergiaceae 667

Genus L Beutenbergia. . . . 669 Genus ll. Miniimonas . . . . 671 Genus lll. Salana 673 Genus lY. Serinibacter . . . 677

Family lll. Bogoriellaceae . 678 Genus L Bogoriella 679 Genus ll. Georgenia 681 Family lV. Brevibacteriaceae. 685 Genus l. Brevibacterium. . 68s Family V. Cell u lomonadaceae. 701 Genus l. Cellulomonas. . . 702 Genus ll. lncertae sedis Actinotalea. . 710 Genus lll. lncertae sedis Demequina. 712 Genus lV. Oerskovia 713 Genus Y. Tropheryma. . . . . 717 Family V L De rmabacte raceae. 727 GenusLDermabacter... 727 Genus ll. Brachybacterium. 730 Famify Yll. Dermacoccaceae 738 Genus L Dermacoccus. . . . 798 Genus ll. Demetria 742 Genus lll. Kytococcus. . . . 744 Family Vlll. Dermatophilaceae 748 Genus L Dermatophilus . . 749 Genus ll. Kineosphaera . . 752 Family lX. I ntrasporangiaceae 754 Genus l. lntrasporangium. 759 Genus ll. Arsenicicoccus 761 Genus lll. Humihabitans. . 764 Genus lV. Janibacter . . . . 765 Genus V. Knoellia 769 Genus Yl. Kribbia 773 Genus Vll. Lapillicoccus . . 774 Genus V lll. O rnithi n icoccus 775 Genus lX. Ornithinimicrobium 777 Genus X. Oryzihumus . . . 781 Genus Xl. Phycicoccus. . . 782 Genus Xll. Serinicoccus. . 78ô Genus Xlll. Terrabacter . . 788 Genus XlV. Terracoccus. . 790 Genus XV. Tetrasphaera . 792 Family X. Jonesiaceae . . 802 Genus l. Jonesia 802 Family Xl. Microbacteriaceae 807 Genus l. Microbacterium . 814 Genus ll. Agreia 852 Genus lll. Agrococcus . . . 855 Genus lV. Agromyces. . . . 862 GenusV. Clavibacter . . . . 877 Genus Yl. Cryobacterium 883 Genus Yll. Curtobacterium. 887 Genus Y lll. Frigoribacte ri um. 895 Genus lX. Frondihabitans. 898 xtv CONTENTS

Genus X. Gulosibacter. . 900 Genus Xl. Humibacter . . . 904 Genus Xll. Labedella 906 GenusXll].Leifsonia.... 907 Genus XlV. Leucobacter. . . 923 GenusXV.Microcella.... 933 Genus XVl. Microterricola 934 Genus XVll. Mycetocola. . 936 Genus XVllL Okibacterium. 940 Genus XlX. Phycicola 942 Genus XX. Plantibacter . . 943 Genus XXl. Pseudoclavibacter. 949 Genus XXll. Rathayibacter. . . 953 Genus XXlll. Rhodoglobus 964 Genus XXIY. Salinibacterium 969 GenusXXV.Subtercola.... 973 Genus XXVI. Yonghaparkia 975 FamilyXll.Promicromonosporaceae.... 995 Genus L Promicromonospora. 995 Genus ll. Cellulosimicrobium 1002 Genus lll.lsoptericola 1006 Genus lY. Myceligenerans . 1010 Genus V. Xylanibacterium. 1012 Genus Vl. Xylanimicrobium . . 1015 Genus Yll. Xylanimonas. . 1016 Family Xlll. Rarobacteraceae 1019 Genus l.Rarobacter..... 1O2O Family XlV. Ruaniaceae . 1022 Genus l. Ruania 1024 Genus.ll. Haloactinobacterium. 1025 Family XV. Sanguibacteraceae. 1027 Genus LSanguibacter... 1030 Part B

Order Xl. Micromonosporales ord. nov.. . 1035 Family L Micromonosporaceae. 1035 Genus 1. Micromonospora 1039 Genus ll. Actinocatenispora !057 Genus lll. Actinoplanes . , 1058 Genus lV. Asanoa. 1088 Genus V. Catellatospora. . . 1090 Genus Vl. Catenuloplanes . . 1092 Genus Vll. Couchioplanes . . 1094 Genus Ylll. Dactylosporangium 1096 Genus lX.Krasilnikovia.... 1106 GenusX.Longispora.... 1107 Genus Xl. Luedemannella 1110 Genus Xll. Pilimelia 1111 Genus Xlll. Polymorphospora 1117 Genus XlV. Salinispora. . . 1118 Genus XV. Spirilliplanes. . 1123 Genus XVL Venucosispora . . 1',?4 Genus XVll. Virgisporangium. 1127 Order Xll. ord. nov. 1137 Family l. 1138 Genus l. Propionibacterium 1138 CONTENTS xv

Genus ll. Aestuariimicrobium 1155 Genus lll.Brooklawnia..,. 1156 Genus lV. Friedmanniella . 1159 Genus V. Granulicoccus . . 1165 Genus Vl. Luteococcus. . . 1166 Genus Yll. Microlunatus . . 1168 GenusVlll.Micropruina... 1173 Genus lX. Propionicicella. 1174 Genus X. Propionicimonas. . 1175 Genus Xl. Propioniferax. , . 1177 Genus Xll. Propionimicrobium 1179 rlr Genus Xlll. Tessaracoccus, 1180 Family ll. Nocardioidaceae. . 1189 rl Genusl.Nocardioides.... 1197 Genus ll. Actinopolymorpha. 1251 Genus lll. Aeromicrobium. . 1258 Genus lY. Kribbella 1268 GenusY.Marmoricola... 1284 Order Xlll. Pseudonocardiales ord. nov. 1301 Family l. Pseudonocardiaceae 1302 Genus l. Pseudonocardia. 1305 Genus ll. Actinoalloteichus. 1323 Genus lll. Actinokineospora 1325 Genus lY. Actinosynnema . 1331 Genus Y. Amycolatopsls. . . 1334 Genus Yl. Crossiella 1359 Genus Vll. Goodfellowiella. 1363 Genus Ylll. Kibdelosporangium 1366 Genus lX. Kutzneria 1371 Genus X. Lechevalieria . . 1375 Genus Xl. Lentzea '1379 GenusXll.Prauserella.... 1384 Genus Xlll. Saccharomonospora 1390 Genus XlY. Saccharopolyspora 1396 Genus XY. Saccharothrix . . 1415 Genus XVl. Streptoalloteichus 1419 Genus XVll. Thermocrispum 1423 Genus XVlll. Umezawaea . 1427 OrderXlV. Streptomycetalesord. nov. .. 1446 Family L Streptomycetaceae 1446 Genusl.Streptomyces.... 1455 Genus lncertaesedlsL Kitasatospora.,.. 1768 Genus lncertae sedis ll. Streptacidiphilus . 1777 Order XV. Streptosporangiales ord. nov. 1805 Family L Streptosporangiaceae 1807 Genus l. Streptosporangium. 181 1 Genus ll. Acrocarpospora . 1825 Genus lll. Herbidospora, . . 1827 Genus lY. Microbispora . . 1831 Genus V. Microtetraspora. 1838 Genus Yl. Nonomuraea . . 1844 Genus Vll. Planobispora. . . 1861 Genus Vlll. Planomonospora 1866 Genus lX. Planotetraspora. 1873 Genus X. Sphaerisporangium 1875 Genus Xl. Thermopolyspora 1880 xvl CONTENTS

Family ll. Nocardiopsaceae 1889 Genus LNocardioPs,s.,.. 1891 Genus ll. Hatoactinos7ora 1907 Genus lll. StrePtomonosPora 1908 Genus lY.Thermobifida.. 1914 Family lll. Thermomonosporaceae. . ' 1929 Genus l. ThermomonosPora 1931 Genus ll-Actinocorattia.... 1935 Genus lll. Actinomadura. . 1940 Genus lY. SPirittosPora. . . 1959 Order XYl. tncertae sedis 1967 Genus LThermobisPora... 1967 1968 Class f l. Acidimicrobira class. nov . . Order l. Acidimicrobiates. 1969 Family L Acidimicrobiaceae . 1970 Genus l. Acidimicrobium. . 1970 Genus ll. Ferrimicrobium . 1972 Genus lll. Ferrithrix 1973 Familyll.lamiaceae.... 1974 Genus l. tamia 1975 Class lll. Coriobacteriiaclass. nov. 1975 Order L Coriobacteriales 1976 Family l. Coriobacteriaceae 1977 Genus l. Coriobacterium . 1977 Genus ll. AtoPobium 1978 Genus 11. Co1insetta 1984 Genus lY. CryPtobacterium 1987 Genus V. Denitrobacterium. 1988 GenusYl.Eggerthetta... 1990 Genus Vll. Otsenetta 1993 Genus ylll. Stackia 1995 Class lV. Nitriliruptoriaclass. nov. 2000 Order l. Nitriliruptorales. . 2001 Family L Nitritiruptoraceae 2001 Genus LNitritiruPtor..... 2001 Order ll.Euzebyate.s.... 2OO2 Family l.Euzebyaceae... 2003 Genus l. Euzebya. :2003 Class V. Rubrobactera class. nov. . . 2OO4 Order l. Rubrobacterales. 2005 Family L Rubrobacteraceae 2006 Genus l. Rubrobacter. . . 2006 ClassVl. Thermoteopftfiiaclass. nov. . . 2010 Order l. Thermoteophitates 2010 Family L Thermoleophitaceae. 2011 Genus l.ThermoteoPhitum. 2011 Order ll. Sotirubrobacterales. 20'14 Family l. Sotirubrobacteraceae 2015 Genus|sotirubrobacter. 2015 Family ll. Conexibacteraceae 2018 Genus LConexibacter... 2019 Family lll. Patutibacteraceae. 2024 Genus LPatutibacter.... 2024

Author index. 2029

lndex of scientific names of Archaeaand 2031