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TheThe AmericanAmerican GARDENERGARDENER® TheThe MagazineMagazine ofof thethe AAmericanmerican HorticulturalHorticultural SocietySociety May / June 2016 go green with Native Ferns Colorful Geums Weed-Control Strategies That Work Versatile Woody Vines contents Volume 9 5 , Number 3 · May I June 2016 FEATURES DEPARTMENTS 5 NOTES FROM RIVER FARM 6 MEMBERS' FORUM 8 NEWS FROM THE AHS AHS recognizes flower show exhibits with its Environmental Award, new initiative launched ro encourage interest in honicultural careers, highlights ofAHS Travel Study ro Ponugal. I2 AHS NEWS SPECIAL Preview of 2016 National Children & Youth Garden Symposium in Columbia, South Carolina. 42 GARDEN SOLUTIONS Tackling plant mites. 44 TRAVELER'S GUIDE TO GARDENS Betty Ford Alpine Gardens in Vail, Colorado. 46 HOMEGROWN HARVEST Good-for-you blueberries. 14 PERENNIAL VINES FOR TEMPERATE GARDENS BY ANDREW BUNTING 48 GARDENER'S NOTEBOOK New plant species Woody-stemmed vines bring their attractive foliage and flowers discovered, citizen to eye level and beyond. scientists needed for tick research, bees get a break 20 A SYMPHONY OF GEUMS BY MARTY WINGATE from a class of pesticides, Orchestrate a harmonious garden with the colorful blooms and cacti used ro purify water, National Gardening Association reconfigured, 2016 tidy foliage of geums. Arrhur Hoyt Scott Medal and Award winner announced, in memoriam for 24 HABITAT HEDGEROWS BY KRIS WETHERBEE garden communicarors Suzanne Frutig Bales and Mel Banholomew. By adapting the traditional hedgerow concept, it's easy to create habitat for wildlife in gardens of any size. 52 GREEN GARAGE Handy hand tools. 3 0 NATIVE FERNS BY C. CO LSTON BURRELL 54 BOOK REVIEWS Endemic to North America, these adaptable species add texture All the Presidents' Gardens, The Cabaret of and quiet grace to shady spots and contemplative nooks. Plants, and Planting in a Post-Wild World. 3 6 WIN THE WAR ON WEEDS BY THOMAS CHRISTOPHER 56 REGIONAL HAPPENINGS Get a grip on garden interlopers with these proven, ecological­ 60 PRONUNCIATIONS AND HARDINESS ly-based strategies to control weeds. AND HEAT ZONES 62 PLANT IN THE SPOTLIGHT ON THE COVER: Ostrich fern and Chrisrmas fern combine to create a serene vignette in this Maryland Bush anemone (Carpenteria californica). shade garden designed by MaryJo Messenger. Photograph by Neil Soderstrom MAY I JUNE 2016 3 Free App AMERICAN HORTICULTURAL SOCIETY HOMEGROWN Making America a Nation of Gardeners, a Land of Gardens With Bonnie Plants Board of Directors CHAIR Amy Bolton Falls Church, Virginia FIRST VICE CHAIRMAN Jane Diamantis McDonald, Tennessee SECOND VICE CHAIRMAN Mary Pat Matheson Atlanta, Georgia SECRETARY Nancy Hargroves Manakin Sabot, Virginia TREASURER J. Landon Reeve, IV Woodbine, Maryland IMMEDIATE PAST CHAIR Harry A. Rissetto, Esq. Falls Church, Virginia EXECUTIVE COMMITTEE Henrietta Burke Alexandria, Virginia Marcia Zech Mercer Island, Washington Skipp Calvert Alexandria, Virginia Q Tim Conlon Dubuque, Iowa Q Gay Estes Houston, Texas Tom Johnson Washington, D.C. Q Louis B. Lynn Columbia, South Carolina Nancy Ross Englewood, Florida Holly Shimizu Glen Echo, Maryland Q Ed Snodgrass Street, Maryland Q Erich Veitenheimer Alexandria, Virginia EXECUTIVE DIRECTOR Tom Underwood PRESIDENT EMERITUS Katy Moss Warner President’s Council The President’s Council is comprised of dedicated members whose annual support makes many of the Society’s programs possible, from youth gardening activities to horticultural awards programs. FOUNDER’S CIRCLE ($25,000+) Mr. and Mrs. George Diamantis Q Ms. Katy Moss Warner Q Mr. and Mrs. Klaus Zech LIBERTY HYDE BAILEY CIRCLE ($10,000-$24,999) Mr. and Mrs. Richard Davison Q Mrs. Elisabeth C. Dudley Q Mr. and Mrs. Neil Morris Q bonnieplants.com/app Mr. and Mrs. Harry A. Rissetto, Esq. HAUPT CIRCLE ($5,000-$9,999) Ms. Amy Bolton and Mr. Philip Schoene Q Mr. and Mrs. Leonard Calvert, III Q Mr. James R. Cargill, II Q Mr. and Mrs. Timothy Conlon Q Mr. Joseph Errington and Mr. William Pullen Q Ms. Catherine M. Hayes Q Mr. and Mrs. Thomas E. Johnson Gifts by Will or Trust Q Mr. Mark Olson Q Mr. and Mrs. J. Landon Reeve, IV benefit you and the SUSTAINER’S CIRCLE ($2,500-$4,999) Mrs. Leslie S. Ariail Q Mrs. Lynda A. Bachman Q Mr. and Mrs. Andy Daniel Q Ms. Julie Ernest Q Mr. and Mrs. Carl Estes Q Dr. Amy Goldman Fowler Q Mr. and Mrs. Joel Goldsmith Q Mr. Thomas Gibian and Ms. Christina Grady Q Dr. and American Horticultural Mrs. William O. Hargrove Q Mr. and Mrs. Albert Huddleston Q Ms. Erika Huddleston Q Dr. and Mrs. Louis B. Lynn Q Mr. and Mrs. Frank Society. Nicolai Q Mr. David D. Parrish Q Dr. Erich E. Veitenheimer and Mr. Andrew Cariaso Q Ms. Katherine J. Ward COUNCIL MEMBER’S CIRCLE ($1,000-$2,499) Ms. E. Pauline Adams Q Ms. Kathleen W. Arnold Q Mr. and Mrs. Robert Baillie Q Mrs. Sallie S. Barnes Q Dr. and Mrs. William E. Barrick Q Mr. Robert A. Bartlett, Jr. Q Mrs. Ritchie Battle Q Mrs. Katherine M. Belk Q Dr. Sherran Blair Q Mr. and Mrs. Michael Bradshaw Q Mrs. Henrietta Burke Q Mr. and Mrs. Allen W. Bush Q Mrs. Torrey Matheson Cooke Q Dr. Karen Davis and Mr. Richard Davis Q Ms. Catherine Eberbach Q Ms. Katherine B. Edwards and Mr. John A. Ronveaux Q Dr. Carol K. Emerson Q Ms. Megan Evans and Mr. Howard M. Tucker Q Ms. Inger Fair Q Mr. and Mrs. A. Michael Gellman Q Mr. Gerald T. Halpin Q Mr. and Mrs. Herbert F. Hargroves Q Mrs. Martha Harris Q Mr. and Mrs. Alfred Heiler Q Mr. and Mrs. Paul A. Hess Q Ms. Nancy Hockstad Q Ms. Jadwiga Hoffmann Q Ms. Patricia Kitchings Q Mr. Charles J. and Dr. Dancy Kittrell Q Mrs. Virginia Korteweg Q Ms. Mary A. Lambert Q Mrs. Carolyn Marsh Lindsay Q Mrs. Mary T. McConnell Q Mr. and Mrs. John A. McMurtrie Q Ms. Elizabeth D. Miller Q Mr. and Mrs. Peter Morris Q Mr. and Mrs. Robert Murray Q Mr. Arnold Orr Q Mr. and Mrs. Al Osman Q Ms. Julie Overbeck Q Mrs. Lynn C. Rhomberg Q Ms. Stephanie L. Rodden and Mr. John Cienki Q Mr. and Mrs. Donald H. Ross Q Mr. and Mrs. James A. Runde Q Mr. and Mrs. Osamu Shimizu Q Mr. and Mrs. Charles Henry Smith, Jr. Q Mr. Howard M. Tucker and Ms. Megan Evans Q Mr. and Mrs. Tom Underwood Q Mr. Joe Viar, Jr. and Ms. Bonnie Christ Q Mrs. Angela M. Vikesland Q Mr. and Mrs. Robert D. Volk Q Mr. and Mrs. Charles F. Walton Q Ms. Elizabeth M. Wehrle Q Mrs. Dudley B. White Q Mrs. Corinne Winburn HONORARY PRESIDENT’S COUNCIL (in memoriam) Ms. Louise Fruehling Q Mrs. Enid Haupt Q Mrs. John A. Lutz Q Mr. and Mrs. Bruce Miller Ms. Wilma L. Pickard Corporate Members Bonnie Plants Q The Care of Trees Q Chapel Valley Landscape Company Q Corona, Inc. The Espoma Company Q Osmocote Q OXO For more information, please contact us at Horticultural Partners [email protected] America In Bloom Q Bellingrath Gardens & Home Q The Colonial Williamsburg Foundation Q Cox Arboretum Metropark or (703) 768-5700. Q Friends of Fellows Riverside Gardens Q The Gardeners of America/Men’s Garden Clubs of America Q The Omni Homestead Q Inniswood Garden Society Q Wegerzyn Gardens Foundation 4 the American Gardener The American NOTES CFROM RIVER FARM GARDENER EDITOR David J. Ellis MANAGING EDITOR AND ART DIRECTOR Mary Yee ASSOCIATE EDITOR Viveka Neveln ATIONAL POLLINATOR WEEK is June 20 to 26, which is a great time EDITORIAL INTERN to stop and think about what we as gardeners can do—individually Mary Chadduck and collectively—to counteract the current decline in the number and CONTRIBUTING EDITOR N health of pollinators. Last summer, the American Horticultural Society became a Rita Pelczar part of the National Pollinator Garden Network (NPGN). The NPGN represents CONTRIBUTING WRITER the interests of the garden, pollinator, and conservation communities, which are Carole Ottesen collaborating to combat this trend. EDITORIAL ADVISORY BOARD One proven way to reverse the dwindling pollinator populations is to increase the CHAIR Ethne Clarke number of pollinator-friendly gardens. Each new garden, whether big or small, across Colorado Springs, Colorado the country means more nectar and pollen Linda Askey sources, more welcoming habitat areas, Birmingham, Alabama and more people dedicated to pollina- Julie Chai Mountain View, California tor-friendly sustainable gardening practic- Mary Irish es. It’s a simple way to make a difference. San Antonio, Texas A signature program of the NPGN is Panayoti Kelaidis the Million Pollinator Garden Challenge, Denver, Colorado which was launched last June. Since then, Charlie Nardozzi tens of thousands of pollinator gardens Burlington, Vermont have been created, including our very own Denny Schrock wildlife garden here at River Farm, thanks Ames, Iowa to our 2015 horticulture interns Sam Guc- 2015 interns Sam Guccione and Stacee Jessica Walliser cione and Stacee Snyder. These two young Snyder in River Farm’s wildlife garden Pittsburgh, Pennsylvania people spent the summer researching and Kris Wetherbee Oakland, Oregon locating the most appropriate plants to include, creating a design, and putting in the sweat equity to bring their plan to fruition. This garden has been registered as one of CONTACT US The American Gardener the million pollinator gardens in this country, and I encourage you to also create and 7931 East Boulevard Drive register your own pollinator gardens at www.millionpollinatorgardens.org. Alexandria, VA 22308 There are many ways to embrace pollinators in your garden; the good news (703) 768-5700 is they don’t require a huge amount of room or a big investment. Pollinator gar- EDITORIAL E-MAIL: [email protected] dens are easy to create, and they are attractive as well as functional.
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  • Infrageneric Revision of the Fern Genus Deparia (Athyriaceae, Aspleniineae, Polypodiales)

    Infrageneric Revision of the Fern Genus Deparia (Athyriaceae, Aspleniineae, Polypodiales)

    Systematic Botany (2018), 43(3): pp. 645–655 © Copyright 2018 by the American Society of Plant Taxonomists DOI 10.1600/036364418X697364 Date of publication August 10, 2018 Infrageneric Revision of the Fern Genus Deparia (Athyriaceae, Aspleniineae, Polypodiales) Li-Yaung Kuo,1,7 Atsushi Ebihara,2 Tian-Chuan Hsu,3 Germinal Rouhan,4 Yao-Moan Huang,5 Chun-Neng Wang,1,6,8 Wen-Liang Chiou,3 and Masahiro Kato2 1Institute of Ecology and Evolutionary Biology, National Taiwan University, Taipei 10617, Taiwan 2Department of Botany, National Museum of Nature and Science, Amakubo 4-1-1, Tsukuba, Ibaraki 305-0005, Japan 3Botanical Garden Division, Taiwan Forestry Research Institute, Taipei 10066, Taiwan 4Mus´eum national d’Histoire naturelle, Institut de Syst´ematique, Evolution, Biodiversit´e ((ISYEB) CNRS, Sorbonne Universit´e EPHE), Herbier national, 16 rue Buffon CP39, F-75005 Paris, France 5Silviculture Division, Taiwan Forestry Research Institute, Taipei 10066, Taiwan 6Department of Life Science, National Taiwan University, Taipei 10617, Taiwan 7Current address: Boyce Thompson Institute, Ithaca, New York 14853, USA ([email protected]) 8Author for correspondence ([email protected]) Communicating Editor: Sven Buerki Abstract—Current molecular phylogenetic analyses support the monophyly and circumscription of the athyrioid fern genus Deparia (Athyr- iaceae), which includes previously recognized genera including Athyriopsis, 3Depazium, Dictyodroma, Dryoathyrium (5 Parathyrium), Lunathyrium, and Neotriblemma (5 Triblemma Ching), and 3Neotribleparia. This broad generic concept has been adopted in several recent taxonomic treatments, including the Pteridophyte Phylogeny Group I. However, the infrageneric taxonomy of Deparia still needs further revision. In this study, we provide a new infrageneric classification with five sections and three subsections based on the phylogenetic evidence.