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001-012 Primeras Páginas
PUBLICACIONES DEL INSTITUTO GEOLÓGICO Y MINERO DE ESPAÑA Serie: CUADERNOS DEL MUSEO GEOMINERO. Nº 9 ADVANCES IN TRILOBITE RESEARCH ADVANCES IN TRILOBITE RESEARCH IN ADVANCES ADVANCES IN TRILOBITE RESEARCH IN ADVANCES planeta tierra Editors: I. Rábano, R. Gozalo and Ciencias de la Tierra para la Sociedad D. García-Bellido 9 788478 407590 MINISTERIO MINISTERIO DE CIENCIA DE CIENCIA E INNOVACIÓN E INNOVACIÓN ADVANCES IN TRILOBITE RESEARCH Editors: I. Rábano, R. Gozalo and D. García-Bellido Instituto Geológico y Minero de España Madrid, 2008 Serie: CUADERNOS DEL MUSEO GEOMINERO, Nº 9 INTERNATIONAL TRILOBITE CONFERENCE (4. 2008. Toledo) Advances in trilobite research: Fourth International Trilobite Conference, Toledo, June,16-24, 2008 / I. Rábano, R. Gozalo and D. García-Bellido, eds.- Madrid: Instituto Geológico y Minero de España, 2008. 448 pgs; ils; 24 cm .- (Cuadernos del Museo Geominero; 9) ISBN 978-84-7840-759-0 1. Fauna trilobites. 2. Congreso. I. Instituto Geológico y Minero de España, ed. II. Rábano,I., ed. III Gozalo, R., ed. IV. García-Bellido, D., ed. 562 All rights reserved. No part of this publication may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopy, recording, or any information storage and retrieval system now known or to be invented, without permission in writing from the publisher. References to this volume: It is suggested that either of the following alternatives should be used for future bibliographic references to the whole or part of this volume: Rábano, I., Gozalo, R. and García-Bellido, D. (eds.) 2008. Advances in trilobite research. Cuadernos del Museo Geominero, 9. -
Phylogenetic Analysis of the Olenellina Walcott, 1890 (Trilobita, Cambrian) Bruce S
2 j&o I J. Paleont., 75(1), 2001, pp. 96-115 Copyright © 2001, The Paleontological Society 0022-3360/01 /0075-96$03.00 PHYLOGENETIC ANALYSIS OF THE OLENELLINA WALCOTT, 1890 (TRILOBITA, CAMBRIAN) BRUCE S. LIEBERMAN Departments of Geology and Ecology and Evolutionary Biology, University of Kansas, Lindley Hall, Lawrence 66045, <[email protected]> ABSTRACT—Phylogenetic analysis was used to evaluate evolutionary relationships within the Cambrian suborder Olenellina Walcott, 1890; special emphasis was placed on those taxa outside of the Olenelloidea. Fifty-seven exoskeletal characters were coded for 24 taxa within the Olenellina and two outgroups referable to the "fallotaspidoid" grade. The Olenelloidea, along with the genus Gabriellus Fritz, 1992, are the sister group of the Judomioidea Repina, 1979. The "Nevadioidea" Hupe, 1953 are a paraphyletic grade group. Four new genera are recognized, Plesionevadia, Cambroinyoella, Callavalonia, and Sdzuyomia, and three new species are described, Nevadia fritzi, Cirquella nelsoni, and Cambroinyoella wallacei. Phylogenetic parsimony analysis is also used to make predictions about the ancestral morphology of the Olenellina. This morphology most resembles the morphology found in Plesionevadia and Pseudoju- domia Egorova in Goryanskii and Egorova, 1964. INTRODUCTION group including the "fallotaspidoids" plus the Redlichiina, and HE ANALYSIS of evolutionary patterns during the Early Cam- potentially all other trilobites. Where the Agnostida fit within this T brian has relevance to paleontologists and evolutionary bi- evolutionary topology depends on whether or not one accepts the ologists for several reasons. Chief among these are expanding our arguments of either Fortey and Whittington (1989), Fortey (1990), knowledge of evolutionary mechanisms and topologies. Regard- and Fortey and Theron (1994) or Ramskold and Edgecombe ing evolutionary mechanisms, because the Cambrian radiation (1991). -
Smithsonian Miscellaneous Collections
SMITHSONIAN MISCELLANEOUS COLLECTIONS VOLUME 53, NUMBER 6 CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.-0LENELLUS AND OTHER GENERA OF THE MESONACID/E With Twenty-Two Plates CHARLES D. WALCOTT (Publication 1934) CITY OF WASHINGTON PUBLISHED BY THE SMITHSONIAN INSTITUTION AUGUST 12, 1910 Zl^i £orb (gaitimovt (pnee BALTIMORE, MD., U. S. A. CAMBRIAN GEOLOGY AND PALEONTOLOGY No. 6.—OLENELLUS AND OTHER GENERA OF THE MESONACID^ By CHARLES D. WALCOTT (With Twenty-Two Plates) CONTENTS PAGE Introduction 233 Future work 234 Acknowledgments 234 Order Opisthoparia Beecher 235 Family Mesonacidas Walcott 236 Observations—Development 236 Cephalon 236 Eye 239 Facial sutures 242 Anterior glabellar lobe 242 Hypostoma 243 Thorax 244 Nevadia stage 244 Mesonacis stage 244 Elliptocephala stage 244 Holmia stage 244 Piedeumias stage 245 Olenellus stage 245 Peachella 245 Olenelloides ; 245 Pygidium 245 Delimitation of genera 246 Nevadia 246 Mesonacis 246 Elliptocephala 247 Callavia 247 Holmia 247 Wanneria 248 P.'edeumias 248 Olenellus 248 Peachella 248 Olenelloides 248 Development of Mesonacidas 249 Mesonacidas and Paradoxinas 250 Stratigraphic position of the genera and species 250 Abrupt appearance of the Mesonacidse 252 Geographic distribution 252 Transition from the Mesonacidse to the Paradoxinse 253 Smithsonian Miscellaneous Collections, Vol. 53, No. 6 232 SMITHSONIAN MISCELLANEOUS COLLECTIONS VOL. 53 Description of genera and species 256 Nevadia, new genus 256 weeksi, new species 257 Mcsonacis Walcott 261 niickwitzi (Schmidt) 262 torelli (Moberg) 264 vermontana -
New Information on Olenelline Trilobites from the Early Cambrian Sekwi Formation, Northwestern Canada
KU ScholarWorks | http://kuscholarworks.ku.edu Please share your stories about how Open Access to this article benefits you. New information on olenelline trilobites from the Early Cambrian Sekwi Formation, Northwestern Canada by Bruce S. Lieberman 2010 This is the author’s accepted manuscript version of the article, made available with the permission of the publisher. The original published version can be found at the link below. Lieberman, Bruce S. 2010. “New information on olenelline trilobites from the Early Cambrian Sekwi Formation, northwestern Canada.” Canadian Journal of Earth Sciences (47):1445-1449. Published version: http://www.nrcresearchpress.com/doi/ abs/10.1139/E10-073#.U21JO4FdXUL Terms of Use: http://www2.ku.edu/~scholar/docs/license.shtml KU ScholarWorks is a service provided by the KU Libraries’ Office of Scholarly Communication & Copyright. For Review Purposes Only/Aux fins d'examen seulement 1 1 New information on olenelline trilobites from the Early Cambrian Sekwi Formation, 2 Northwestern Canada 3 4 Francine R. Abe 5 Department of Ecology and Evolutionary Biology, University of Kansas, 1345 Jayhawk 6 Boulevard, Lawrence, Kansas 66045, USA, Tel: +1 (785) 864-3369, Fax: +1 (785) 864-5335 7 <[email protected]> 8 Bruce S. Lieberman 9 Department of Geology and Natural History Museum and Biodiversity Research Center, 10 University of Kansas, 1475 Jayhawk Boulevard, 120 Lindley Hall, Lawrence, Kansas 66045, 11 USA, <[email protected]> 12 Michael C. Pope 13 Department of Geology and Geophysics, Texas A&M University, College Station, Texas 77843, 14 USA, <[email protected]> 15 Kelly Dilliard 16 Department of Physical Sciences and Mathematics, Wayne State College, 1111 Main Street, 17 Wayne, Nebraska 68787, USA, <[email protected]> 18 For Review Purposes Only/Aux fins d'examen seulement 2 19 Abstract 20 A new species of olenelline trilobite, Nevadella keelensis, is described from the lower Cambrian Sekwi 21 Formation, Mackenzie Mountains, Canada. -
Re-Evaluation of the Stratigraphically Important Olenellid Trilobite Holmia Cf. Mobergi from the Cambrian Series 2, Stage 3
NORWEGIAN JOURNAL OF GEOLOGY Vol 99 Nr. 1 https://dx.doi.org/10.17850/njg99-1-04 Re-evaluation of the stratigraphically important olenellid trilobite Holmia cf. mobergi from the Cambrian Series 2, Stage 3 and its implications for the lower Cambrian stratigraphy in the Mjøsa area, Norway Magne Høyberget1, Jan Ove R. Ebbestad2 & Bjørn Funke3 1Rennesveien 14, N–4513 Mandal, Norway. 2Museum of Evolution, Uppsala University, Norbyvägen 16, SE–752 36 Uppsala, Sweden. 3Gjelleråsveien 10, N–1481 Hagan, Norway. E-mail corresponding author (Magne Høyberget): [email protected] The olenellid trilobite Holmia cf. mobergi, known from a single cephalon in the upper lower Cambrian strata from a river section in Flagstadelva, Hamar, has played a significant stratigraphic role in interpreting the lower Cambrian informal Series 2, Stage 3 in the Mjøsa area, Norway, since its discovery in the early 1950s. It was considered one of the oldest trilobite taxa in the lower Cambrian of Scandinavia, but the stratigraphic level and biozonation of the cephalon were problematic and a matter of discussion for decades. Moreover, organic-walled microfossil biostratigraphy questioned the supposed age of the trilobite. New specimens of this taxon collected from the type locality show that the species occurs at a different stratigraphic level than first reported, prompting a new description of the species and a re-evaluation of the taxon’s biostratigraphic significance. Holmia cf. mobergi is compared with new and well-preserved topotype material of Holmia inusitata, a very rare taxon hitherto known from one single outcrop in an autochthonous setting in Norway. Holmia cf. -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
Tentative Correlation of Cambrian Series 2 Between South China and Other Continents
Tentative correlation of Cambrian Series 2 between South China and other continents JINLIANG YUAN, XUEJIAN ZHU, JIHPAI LIN & MAOYAN ZHU The apparent absence, in Cambrian Series 2, of widespread and rapidly evolving organisms comparable to the later agnostids, graptolites, conodonts, ammonites, or planktonic foraminifers, has prevented a consistent intercontinental biostratigraphy. Occasional genera, and (rarely) species, of trilobites, archaeocyathans and other groups may be found in more than one region. Nevertheless, based on the complete trilobite and archaeocyathan successions in the shallow water Yangtze Platform and in deeper water Jiangnan Slope environment, correlation of Cambrian Series 2 between South China and other continents is discussed in detail. The oldest trilobite Parabadiella and Tsunyidiscus in South China can be correlated with the oldest trilobite Abadiella in Australia, Profallotapis in Siberia and Eofallotaspis in Morocco. • Key words: Cambrian Series 2, correlation, South China, other continents. YUAN, J.L., ZHU, X.J., LIN,J.P.&ZHU, M.Y. 2011. Tentative correlation of Cambrian Series 2 between South China and other continents. Bulletin of Geosciences 86(3), 397–404 (2 tables). Czech Geological Survey. Prague, ISSN 1214-1119. Manuscript received December 17, 2010; accepted in revised form July 1, 2011; published online Septem- ber 22, 2011; issued September 30, 2011. Jinliang Yuan, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China; [email protected] • Xuejian Zhu, Jihpai Lin & Maoyan Zhu, State Key Laboratory of Palaeontology and Stratigra- phy, Nanjing Institute of Geology and Palaeontology,Chinese Academy of Sciences, Nanjing, 210008, China International correlation of Cambrian Series 2 has been a nomenclature lead to paucity of biostratigraphically useful major problem since the concept of four series within the data in key regions (Palmer 1998, Geyer 2001). -
Th TRILO the Back to the Past Museum Guide to TRILO BITES
With regard to human interest in fossils, trilobites may rank second only to dinosaurs. Having studied trilobites most of my life, the English version of The Back to the Past Museum Guide to TRILOBITES by Enrico Bonino and Carlo Kier is a pleasant treat. I am captivated by the abundant color images of more than 600 diverse species of trilobites, mostly from the authors’ own collections. Carlo Kier The Back to the Past Museum Guide to Specimens amply represent famous trilobite localities around the world and typify forms from most of the Enrico Bonino Enrico 250-million-year history of trilobites. Numerous specimens are masterpieces of modern professional preparation. Richard A. Robison Professor Emeritus University of Kansas TRILOBITES Enrico Bonino was born in the Province of Bergamo in 1966 and received his degree in Geology from the Depart- ment of Earth Sciences at the University of Genoa. He currently lives in Belgium where he works as a cartographer specialized in the use of satellite imaging and geographic information systems (GIS). His proficiency in the use of digital-image processing, a healthy dose of artistic talent, and a good knowledge of desktop publishing software have provided him with the skills he needed to create graphics, including dozens of posters and illustrations, for all of the displays at the Back to the Past Museum in Cancún. In addition to his passion for trilobites, Enrico is particularly inter- TRILOBITES ested in the life forms that developed during the Precambrian. Carlo Kier was born in Milan in 1961. He holds a degree in law and is currently the director of the Azul Hotel chain. -
An Inventory of Trilobites from National Park Service Areas
Sullivan, R.M. and Lucas, S.G., eds., 2016, Fossil Record 5. New Mexico Museum of Natural History and Science Bulletin 74. 179 AN INVENTORY OF TRILOBITES FROM NATIONAL PARK SERVICE AREAS MEGAN R. NORR¹, VINCENT L. SANTUCCI1 and JUSTIN S. TWEET2 1National Park Service. 1201 Eye Street NW, Washington, D.C. 20005; -email: [email protected]; 2Tweet Paleo-Consulting. 9149 79th St. S. Cottage Grove. MN 55016; Abstract—Trilobites represent an extinct group of Paleozoic marine invertebrate fossils that have great scientific interest and public appeal. Trilobites exhibit wide taxonomic diversity and are contained within nine orders of the Class Trilobita. A wealth of scientific literature exists regarding trilobites, their morphology, biostratigraphy, indicators of paleoenvironments, behavior, and other research themes. An inventory of National Park Service areas reveals that fossilized remains of trilobites are documented from within at least 33 NPS units, including Death Valley National Park, Grand Canyon National Park, Yellowstone National Park, and Yukon-Charley Rivers National Preserve. More than 120 trilobite hototype specimens are known from National Park Service areas. INTRODUCTION Of the 262 National Park Service areas identified with paleontological resources, 33 of those units have documented trilobite fossils (Fig. 1). More than 120 holotype specimens of trilobites have been found within National Park Service (NPS) units. Once thriving during the Paleozoic Era (between ~520 and 250 million years ago) and becoming extinct at the end of the Permian Period, trilobites were prone to fossilization due to their hard exoskeletons and the sedimentary marine environments they inhabited. While parks such as Death Valley National Park and Yukon-Charley Rivers National Preserve have reported a great abundance of fossilized trilobites, many other national parks also contain a diverse trilobite fauna. -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
New Olenelloid Trilobites from the Northwest Territories, Canada
Zootaxa 3866 (4): 479–498 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3866.4.2 http://zoobank.org/urn:lsid:zoobank.org:pub:D06E5477-4D5C-4402-B909-09A2AAFFB556 New olenelloid trilobites from the Northwest Territories, Canada I. WESLEY GAPP1,2 & BRUCE S. LIEBERMAN2 1Chevron U. S. A., Inc., 1400 Smith Street, Houston, TX 77002, USA. E-mail: [email protected] 2Department of Ecology & Evolutionary Biology and Biodiversity Institute, University of Kansas, 1345 Jayhawk Blvd, Dyche Hall, Lawrence, KS 66045, USA. E-mail: [email protected] Abstract The Olenelloidea are a superfamily of early Cambrian trilobites, which have been the subject of several phylogenetic anal- yses and also used to address macroevolutionary questions regarding the nature and timing of the Cambrian radiation. The Sekwi Formation of the Mackenzie Mountains, Northwest Territories, Canada, has yielded numerous species from this clade, and here we present new information that expands on the diversity known from this biogeographically and biostrati- graphically important region. In particular, we describe seven new species, (Olenellus baileyi, Mesonacis wileyi, Ellipto- cephala jaredi, Holmiella taurus, H. domackae, Mummaspis delgadoae, and Bristolia colberti). Also recovered are specimens of Elliptocephala logani, specimens that shared affinities with Olenellus clarki, O. getzi, O. fowleri, and Fri- zolenellus hanseni, and one partial specimen, which appears to be a new species of Bolbolenellus. Key words: Cambrian, Trilobita, Olenelloidea, Northwest Territories Introduction The Olenelloidea Walcott, 1890 is a diverse superfamily of early Cambrian trilobites referable to the suborder Olenellina Walcott, 1890 and have been the focus of much attention in the study of evolutionary tempo and mode during the Cambrian radiation (Fortey et al. -
Nr. 220 CONTRIBUTIONS to the GEOLOGY of the MJØSA DISTRICTS and the CLASSICAL SPARAGMITE AREA in SOUTHERN NORWAY
Nr. 220 CONTRIBUTIONS TO THE GEOLOGY OF THE MJØSA DISTRICTS AND THE CLASSICAL SPARAGMITE AREA IN SOUTHERN NORWAY By STEINAR SKJESETH sammendrag: Bidrag til geologien i Mjøstraktene med en kort foreløpig analyse av stratigrafi og tektonikk i det klassiske sparagmittområde WITH 65 TEXT-FIGURES AND 2 PLATES OSLO 1963 UNIVERSITETSFORLAGET NORGES GEOLOGISKE UNDERSØKELSE NR. 220 CONTRIBUTIONS TO THE GEOLOGY OF THE MJØSA DISTRICTS AND THE CLASSICAL SPARAGMITE AREA IN SOUTHERN NORWAY By STEINAR SKJESETH sammendrag: Bidrag til geologien i Mjøstraktene med en kort foreløpig analyse av stratigrafi og tektonikk i det klassiske sparagmittområde WITH 65 TEXT-FIGURES AND 2 PLATES OSLO 1963 UNIVERSITETSFORLAGET Jo, 54? (W Editor for the Publications of the Geological Survey of Norway: State Geologist Fredrik Hagemann A.W. BRØGGERS BOKTRYKKERI A/S - OSLO Contents. Abstract 5 Acknowledgements 7 Introduction 9 The geological setting of the Mjøsa Districts 12 Historical review 13 STRATIGRAPHY 17 The Sparagmite Group "Eocambrian" 17 Eocambrian rocks within the mapped area Description of localities 19 Autochthonous series 21 Allochthonous series 26 The stratigraphy of the Sparagmite Group Elstad Sparagmite 27 Brøttum Sparagmite 27 Brøttum Shale and Limestone 28 Biri Conglomerate 28 Biri Shale and Limestone 29 Moelv Sparagmite 30 Moelv Conglomerate (= tillite) 31 Ekre Shale 32 Mjøsa Quartz-sandstone formation Vardal Sparagmite and Ringsaker Quartzite 32 The Cambrian System Description of localties Autochthonous series 35 Allochthonous series 44 The Cambrian