THE MAMMAL FAUNA OF THE EARLY EOCENE BLACKHEATH FORMATION OF ABBEY WOOD, LONDON

CONTENTS Page INTRODUCTION 1 ACKNOWLEDGEMENTS 1 INSTITUTIONAL ABBREVIATIONS 2 HISTORY OF RESEARCH .. 2 STRATIGRAPHY 4 SYSTEMATIC DESCRIPTIONS 11

Page Page Subclass ALLOTHERIA Marsh, 1880 11 Genus Macrocranion Weitzel, 1949 .. .. 38 Order MULTITUBERCULATA Cope, 1884b .. 11 Genus indet 41 Family NEOPLAGIAULACIDAE Grandordcr ARCHONTA Gregory, 1910.. .. 43 Amcghino,1890 11 Order CHIROPTERA Blumcnbach, 1779 .. 43 Genus Ectypodus Matthew & Granger, Family EPPSINYCTERIDIDAE nov. .. .. 43 1921 11 Genus Eppsinycteris Hooker, 1996b .. .. 43 Subclass TREGHNOTHERIA McKenna, 1975 .. ..14 Family and genus indet. 44 Infraclass THERIA Parker & Haswcll, 1897 14 Order uncertain 44 Supercohort METATHERIA Huxley, 1880 .. ..14 Family ADAPISORICULIDAE Van Valen, Cohort MARSUPIALIA Illiger, 1811 14 1967.! 44 Family HERPETOTHERITDAE Trouessart, 1879 14 Genus Wyonycteris Gingerich, 1987 .. .. 44 Genus Amphiperatherium Eilhol, 1879 .. .. 14 Genus Remiculus Russell, 1964? 45 Order DIDELPHIMORPHIA Gill, 1872 17 Family NYCTITHERIIDAE Simpson, 1928 47 Family PERADECTIDAE Crochet, 1979 .. ..17 Genus Leplacodon Matthew & Granger, Genus Peradectes Matthew & Granger, 1921 .. 17 1921 47 Supercohort EUTHERIA Huxley, 1880.. 17 Order PLESIADAPIFORMES Simons, 1972 48 Cohort PLACENTALIA Owen, 1837 17 Family PAROMOMYIDAE Simpson, 1940 48 Order PANTOLESTA McKenna, 1975 17 Genus Arcius Godinot, 1984 48 Family PANTOLESTIDAE Cope, 1884a .. ..17 Family PLESIADAPIDAE Trouessart, 1897 49 Genus Palaeosinopa Matthew, 1901 17 Genus Platychoerops Charlesworth, 1855.. 49 Order LEPTICTIDA McKenna, 1975? 20 Order PRIMATES Linnaeus, 1758 50 Family LEPTICTIDAE Gill, 1872? 20 Family OMOMYIDAE Trouessart, 1879 .. 50 Genus indet 20 Subfamily MICROCHOERINAE Grandordcr ANAGALIDA Szalay & McKenna, Lydekker, 1887 50 1971 22 Genus Melaneremia Hooker, 2007 .. .. 50 Order TILLODONTIA Marsh, 1875 22 Superfamily ADAPOIDEA Trouessart, 1879 52 Family ESTHONYCHIDAE Cope, 1883 .. ..22 Family NOTHARCTIDAE Trouessart, 1879 Genus Plesiesthonyx Lemoine, 1891 22 (sensu stricto) 52 Order RODENTIA Bowdich, 1821 24 Genus Cantius Simons, 1962 52 Family 'PARAMYIDAE' Miller & Gidley, 1918 24 Grandorder FERAE Linnaeus, 1758 57 Subfamily AILURAVINAE Michaux, 1968 .. 24 Order DIDELPHODONTA McKenna, 1975 57 Genus Ailuravus Riitimeyer, 1891 24 Family CIMOLESTIDAE Marsh, 1889.. .. 57 Genus Meldimys Michaux, 1968? 28 GenusDidelphodus Cope, 1882b 57 Genus indct 28 Order APATOTHERIA Scott &Jepsen, 1936 61 Subfamily PSEUDOPARAMYINAE Michaux, Family APATEMYIDAE Matthew, 1909 .. 61 1964 29 Genus Apatemys Marsh, 1872 61 Genus Pseudoparamys Michaux, 1964 29 Order PANTODONTA Cope, 1873 64 Subfamily MICROPARAMYINAE Wood, Family CORYPHODONTIDAE Marsh, 1962 (J«WK Dawson 2001) 30 1876b 64 Genus Sparnacomys Hartenberger, 1971 .. .. 30 Genus Coryphodon Owen, 1845 64 Genus Pantrogna Hartcnberger, 1971 .. .. 33 Order CREODONTA Cope, 1875b 73 Family incertae sedis 34 Family OXYAENIDAE Cope, 1877 73 Genus undet 34 Subfamily AMBLOCTONINAE Cope, Grandordcr LIPOTYPHLA Haeckel, 1866 .. ..36 1877 73 Order ERINACEOMORPHA Gregory, 1910 .. 36 Genus Palaeonictis de Blainville, 1842 .. 73 Family AMPHILEMURIDAE Hill, 1953 .. ..36 Subfamily OXYAENINAE Cope, 1877 .. 79 Genus Neomalronella Russell, Louis & Savage, Genus Oxyaena Cope, 1874 79 1975 ' .. 36 Family HYAENODONTIDAE Leidy, 1869 .. 86 Page Page Genus Prototomus Cope, 1874 86 Family 'DIACODEXEIDAE'Gazin, 1955.. .. 104 Order CARNIVORA Bowdich, 1821 .. 89 Genus Diacodexis Cope, 1882c 104 Family VIVERRAVIDAE Wortman & Order PANAMERIUNGUI ATA Muizon & Matthew, 1899 89 Cifclli, 2000 109 Genus\ iverravus Marsh, 1872 89 Family HYOPSODONTIDAE Troucssart, Family 'MIACIDAE' Cope, 1880a .. .. 91 1879.! 109 Genus Miacis Cope, 1872d 91 Genus Hyopsodus Leidy, 1870 109 Genus Uintacyon Lcidy, 1872 95 Genus Lessnessina Hooker, 1979 113 Grandorder UNGULATA Linnaeus, 1766.. 97 Order THENACODONTA' McKenna, 1975 114 Order ARCTOCYONIA Van Valen, 1969 97 Family 'PHENACODONTIDAE' Cope, Family ARCTOGYONIDAE Giebel, 1855 97 1881b 114 Subfamily ARCTOCYONINAE Giebel, Genus Phenacodus Cope, 1873 114 1855 97 Order PERISSODACTYLA Owen, 1848 ..117 Genus Arctocyonides Lemoine, 1891 .. 97 Infraorder EUPERISSODACTYTA Hooker Order MESONYCHIA Matthew, 1937 .. 100 & Dashzeveg, 2004 117 Family MESONYCHIDAE Cope, 1875b 100 Superfamily EQUOIDEA Gray, 1821 .. .. 117 Genus Pachyaena Cope, 1874 100 Family EQUIDAE Gray, 1821? 117 Order ARTIODACTYLA Owen, 1848 .. .. 104 Genus Pliolophus Owen, 1858 117

CORRELATION 129 PALAEOENVIRONMENT 132 HABTTAT OF THE MAMMALS 132 ASPECTS OF TAPHONOMY 136 INTERCONTINENTAL DISPERSAL 138 REFERENCES 140 APPENDIX 154 INDEX 158