Palaeontological Heritage Impact Assessment for N2 Upgrade Between Grahamstown and Fish River
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Chapter 2 Paleozoic Stratigraphy of the Grand Canyon
CHAPTER 2 PALEOZOIC STRATIGRAPHY OF THE GRAND CANYON PAIGE KERCHER INTRODUCTION The Paleozoic Era of the Phanerozoic Eon is defined as the time between 542 and 251 million years before the present (ICS 2010). The Paleozoic Era began with the evolution of most major animal phyla present today, sparked by the novel adaptation of skeletal hard parts. Organisms continued to diversify throughout the Paleozoic into increasingly adaptive and complex life forms, including the first vertebrates, terrestrial plants and animals, forests and seed plants, reptiles, and flying insects. Vast coal swamps covered much of mid- to low-latitude continental environments in the late Paleozoic as the supercontinent Pangaea began to amalgamate. The hardiest taxa survived the multiple global glaciations and mass extinctions that have come to define major time boundaries of this era. Paleozoic North America existed primarily at mid to low latitudes and experienced multiple major orogenies and continental collisions. For much of the Paleozoic, North America’s southwestern margin ran through Nevada and Arizona – California did not yet exist (Appendix B). The flat-lying Paleozoic rocks of the Grand Canyon, though incomplete, form a record of a continental margin repeatedly inundated and vacated by shallow seas (Appendix A). IMPORTANT STRATIGRAPHIC PRINCIPLES AND CONCEPTS • Principle of Original Horizontality – In most cases, depositional processes produce flat-lying sedimentary layers. Notable exceptions include blanketing ash sheets, and cross-stratification developed on sloped surfaces. • Principle of Superposition – In an undisturbed sequence, older strata lie below younger strata; a package of sedimentary layers youngs upward. • Principle of Lateral Continuity – A layer of sediment extends laterally in all directions until it naturally pinches out or abuts the walls of its confining basin. -
New Heterodontosaurid Remains from the Cañadón Asfalto Formation: Cursoriality and the Functional Importance of the Pes in Small Heterodontosaurids
Journal of Paleontology, 90(3), 2016, p. 555–577 Copyright © 2016, The Paleontological Society 0022-3360/16/0088-0906 doi: 10.1017/jpa.2016.24 New heterodontosaurid remains from the Cañadón Asfalto Formation: cursoriality and the functional importance of the pes in small heterodontosaurids Marcos G. Becerra,1 Diego Pol,1 Oliver W.M. Rauhut,2 and Ignacio A. Cerda3 1CONICET- Museo Palaeontológico Egidio Feruglio, Fontana 140, Trelew, Chubut 9100, Argentina 〈[email protected]〉; 〈[email protected]〉 2SNSB, Bayerische Staatssammlung für Paläontologie und Geologie and Department of Earth and Environmental Sciences, LMU München, Richard-Wagner-Str. 10, Munich 80333, Germany 〈[email protected]〉 3CONICET- Instituto de Investigación en Paleobiología y Geología, Universidad Nacional de Río Negro, Museo Carlos Ameghino, Belgrano 1700, Paraje Pichi Ruca (predio Marabunta), Cipolletti, Río Negro, Argentina 〈[email protected]〉 Abstract.—New ornithischian remains reported here (MPEF-PV 3826) include two complete metatarsi with associated phalanges and caudal vertebrae, from the late Toarcian levels of the Cañadón Asfalto Formation. We conclude that these fossil remains represent a bipedal heterodontosaurid but lack diagnostic characters to identify them at the species level, although they probably represent remains of Manidens condorensis, known from the same locality. Histological features suggest a subadult ontogenetic stage for the individual. A cluster analysis based on pedal measurements identifies similarities of this specimen with heterodontosaurid taxa and the inclusion of the new material in a phylogenetic analysis with expanded character sampling on pedal remains confirms the described specimen as a heterodontosaurid. Finally, uncommon features of the digits (length proportions among nonungual phalanges of digit III, and claw features) are also quantitatively compared to several ornithischians, theropods, and birds, suggesting that this may represent a bipedal cursorial heterodontosaurid with gracile and grasping feet and long digits. -
Geology and Sedimentology of the Upper Devonian Escuminac
Document généré le 2 oct. 2021 16:46 Atlantic Geology Geology and sedimentology of the Upper Devonian Escuminac Formation, Quebec, and evaluation of its paleoenvironment: lacustrine versus estuarine turbidite sequence Reinhard Hesse et Hemdat Sawh Volume 28, numéro 3, november 1992 Résumé de l'article La Formation d'Escuminac du Groupe de Miguasha du DeVonien sup£rieur, URI : https://id.erudit.org/iderudit/ageo28_3art05 dans la partie ouest de la baie des Chaleurs, Quebec, ceJcbre pour sa faune de poissons fossile, est une sequence de turbidites dont l'environnemcnt de Aller au sommaire du numéro deposition a fait l'objet de discussions. Les interpretations varient suivant les diffeients auteurs entre lacustre ou saumatre amarin cotier. C'estpossiblement d'origine estuarienne. Son epaisseur totale de 117 m a €l6 divis£e en huit Éditeur(s) unite's lithostratigraphiques sur la base de variations verticales dans le rapport gres:shale. Un milieu lacustre est possible en regard de 1'association avec les Atlantic Geoscience Society formations fluviatiles en-dessous (Formation de Fleurant) et au-dessus (Formation de Bonaventure), bien qu'elles soient scparees de TEscuminac par ISSN une inconformite' ou une discordance angulaire. L'absence de toute faune marine invert£br£e & coquillc est un argument ne'gatif favorisant 0843-5561 (imprimé) indirectement une origine lacustre. Des conditions d'eau de fond 1718-7885 (numérique) periodiquement stagnante, indiqules par la preservation de laminites, sont compatibles avec un environnement de lac. La faune de poissons fossile, Découvrir la revue cependant, contient des elements qui pointent vers des environnement saumatres ou marins. Les donnfies gdochimiques semblent aussi appuyer des conditions saumatres, bien qu'il ne soit pas clair jusqu'a quel point la signature Citer cet article geochimique ait pu etre h£ritec d'argiles marines plus anciennes. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Earth-Science-Conservation-No.-029-June-1991.Pdf
Editorial Contents Dawn ofa new era I Main features I Chris Stevens, English Nature As of 1 April the Government's new The Strategy launched at Westtninster 3 Earth science conservation in Great arrangements for nature conservation Britain - a strategy, to give it its full came into effect, and statutory duties The RIGS initiative - full speed ahead title, was formally launched on 5 for England, Scotland and Wales are Mike Harley reports on the astonishing pace at which RIGS December 1990. ,The sening was the now discharged via three new national schemes are growing 5 Hoare Memorial Hall in Church agencies - English Nature, Nature RockWATCH - fossils, fun and lots more! House, Westminster - a slightly more Conservancy Council for Scotland and Wayne Talbot and Beverley Halstead ask what's in it for the kids? 10 geological setting than its name Countryside Council for Wales (for would suggest, with lots of Ancaster further details see page 9). The Geological Society and conservation and Purbeck limestone internally and Coos Wilson explains the role of the Conservation Committee 22 A future for our magazine a splendid flint-clad exterior. Ifthe 'Strategy' is new to you, you will find The new agencies have agreed to I Protecting our earth science resource I a summary of it in Earth science produce Earth science conservation as a conservation) 27 and 28. joint effort. This is encouraging news What is the earth science resource that we are so keen to protect? to all who believe that the magazine has These articles illustrate just three aspects of this rich heritage A full house! a useful role to play. -
Evolúcia Ekosystémov
1 UNIVERZITA KOMENSKÉHO V BRATISLAVE PRÍRODOVEDECKÁ FAKULTA EVOLÚCIA EKOSYSTÉMOV JOZEF KLEMBARA Katedra ekológie BRATISLAVA, 2014 2 ©2014, Jozef Klembara Univerzita Komenského v Bratislave Prírodovedecká fakulta Katedra ekológie 84215 Bratislava, Slovensko Telefón: +421 2/60296257 1. vydanie ISBN 978-80-223-3682-6 Recenzenti: Mgr. Andrej Čerňanský, PhD RNDr. Jana Ciceková, PhD. Mgr. Ivan Bartík 3 OBSAH Evolúcia ekosystémov 1 - GLOBÁLNA BLOKOVÁ TEKTONIKA A KONTINENTÁLNY DRIFT ........................................................................................................................ 4 1. 1 - Globálna bloková tektonika .................................................................................... 5 1.2 - Príčiny pohybu litosferických blokov ...................................................................... 10 1.3 - Kontinentálny drift ................................................................................................... 10 2 - EVOLÚCIA EKOSYSTÉMOV ....................................................................... 12 2.1 - Prekambrium (4600 – 545 mil. r.) ........................................................................... 15 2.2 - Prvohory - Paleozoikum (545 – 245 mil. r.) ............................................................ 17 2.2.1 – Kambrium .............................................................................................................. 18 2.2.2 – Ordovik ................................................................................................................. -
Maceral Types and Quality of Coal in the Tuli Coalfield: a Case
applied sciences Article Maceral Types and Quality of Coal in the Tuli Coalfield: A Case Study of Coal in the Madzaringwe Formation in the Vele Colliery, Limpopo Province, South Africa Elelwani Denge * and Christopher Baiyegunhi Department of Geology and Mining, University of Limpopo, Private Bag X1106, Sovenga 0727, South Africa; [email protected] * Correspondence: [email protected] Featured Application: Authors are encouraged to provide a concise description of the specific application or a potential application of the work. This section is not mandatory. Abstract: The Madzaringwe Formation in the Vele colliery is one of the coal-bearing Late Palaeozoic units of the Karoo Supergroup, consisting of shale with thin coal seams and sandstones. Maceral group analysis was conducted on seven representative coal samples collected from three existing boreholes—OV125149, OV125156, and OV125160—in the Vele colliery to determine the coal rank and other intrinsic characteristics of the coal. The petrographic characterization revealed that vitrinite is the dominant maceral group in the coals, representing up to 81–92 vol.% (mmf) of the total sample. Collotellinite is the dominant vitrinite maceral, with a total count varying between 52.4 vol.% (mmf) and 74.9 vol.% (mmf), followed by corpogelinite, collodetrinite, tellinite, and pseudovitrinite with a Citation: Denge, E.; Baiyegunhi, C. count ranging between 0.8 and 19.4 vol.% (mmf), 1.5 and 17.5 vol.% (mmf), 0.8 and 6.5 vol.% (mmf) Maceral Types and Quality of Coal in the Tuli Coalfield: A Case Study of and 0.3 and 5.9 vol.% (mmf), respectively. The dominance of collotellinite gives a clear indication Coal in the Madzaringwe Formation that the coals are derived from the parenchymatous and woody tissues of roots, stems, and leaves. -
Guides Level Ii Manual 2005 December
GUIDING LEVEL II A TRAINING MANUAL DESIGNED TO ASSIST WITH PREPARATION FOR THE FGASA LEVEL II AND TRAILS GUIDE EXAMS All rights reserved. No part of the material may be reproduced or utilized in any form or by any means, electronic or mechanical including photocopying, recording or by an information storage retrieval system, without the written permission of Lee Gutteridge. (INCLUDING MORE THAN FOUR HUNDRED PHOTOS AND DIAGRAMS) COMPILED BY LEE GUTTERIDGE THIS STUDY MATERIAL CONFORMS TO THE SYLLABUS SET BY FGASA FOR THE LEVEL II EXAMS AND IS APPROVED BY PROFESSOR W.VAN HOVEN OF THE CENTRE FOR WILDLIFE MANAGEMENT AT THE UNIVERSITY OF PRETORIA P.O. Box 441, Mookgopong, 0560, Limpopo, South Africa. Cell 083 667 7586 2 LEVEL TWO TRAINING MANUAL This manual has been compiled from the perspective of a guide in the field. In writing it I asked myself what can I use on a game drive, or game walk as regards information. These aspects covered in this manual will give the guide good, interesting and factual information for direct discussion with the guest. No one book will cover every aspect so here I have included sections on the following topics. 1. Ecology 2. Mammals 3. Birds 4. Reptiles and Amphibians 5. Astronomy 6. Botany 7. Insects, Arachnids and their relatives 8. Geology and Climatology 9. Fish 10. Survival 11. AWH and VPDA The problem for guides is not always finding the answers, but also what is the question to be researched in the first place? It is difficult for a guide to pre-empt what guests will ask them over their guiding careers, but many of the questions and answers which will come into play have been covered here. -
Archaeopteris Is the Earliest Known Modern Tree
letters to nature seawater nitrate mapping systems for use in open ocean and coastal waters. Deep-Sea Res. I 43, 1763± zones as important sites for the subsequent development of lateral 1775 (1996). 26. Obata, H., Karatani, H., Matsui, M. & Nakayama, E. Fundamental studies for chemical speciation in organs; and wood anatomy strategies that minimize the mechani- seawater with an improved analytical method. Mar. Chem. 56, 97±106 (1997). cal stresses caused by perennial branch growth. Acknowledgements. We thank G. Elrod, E. Guenther, C. Hunter, J. Nowicki and S. Tanner for the iron Archaeopteris is thought to have been an excurrent tree, with a analyses and assistance with sampling, and the crews of the research vessels Western Flyer and New Horizon single trunk producing helically arranged deciduous branches for providing valuable assistance at sea. Funding was provided by the David and Lucile Packard 7 Foundation through MBARI and by the National Science Foundation. growing almost horizontally . All studies relating to the develop- ment of Archaeopteris support the view that these ephemeral Correspondence and requests for materials should be addressed to K.S.J. (e-mail: johnson@mlml. calstate.edu). branches arise from the pseudomonopodial division of the trunk apex8±11. Apical branching also characterizes all other contempora- neous non-seed-plant taxa including those that had also evolved an arborescent habit, such as lepidosigillarioid lycopsids and cladoxy- Archaeopteris is the earliest lalean ferns. This pattern, which can disadvantage the tree if the trunk apex is damaged, contrasts with the axillary branching knownmoderntree reported in early seed plants12. Analysis of a 4 m-long trunk from the Famennian of Oklahoma has shown that Archaeopteris may Brigitte Meyer-Berthaud*, Stephen E. -
The Stratigraphy of the Ohio Range, Antarctica
This dissertation has been 65—1200 microfilmed exactly as received LONG, William Ellis, 1930- THE STRATIGRAPHY OF THE OHIO RANGE, ANTARCTICA. The Ohio State University, Ph.D., 1964 G eology University Microfilms, Inc., Ann Arbor, Michigan THE STRATIGRAPHY OF THE OHIO RANGE, ANTARCTICA DISSERTATION Presented in Partial Fulfillment of the Requirements for the Degree Doctor of Philosophy in the Graduate School of The Ohio State University By William Ellis Long, B.S., Rl.S. The Ohio State University 1964 Approved by A (Miser Department of Geology PLEASE NOTE: Figure pages are not original copy* ' They tend tc "curl11. Filled in the best way possible. University Microfilms, Inc. Frontispiece. The Ohio Range, Antarctica as seen from the summit of ITIt. Glossopteris. The cliffs of the northern escarpment include Schulthess Buttress and Darling Ridge. The flat area above the cliffs is the Buckeye Table. ACKNOWLEDGMENTS The preparation of this paper is aided by the supervision and advice of Dr. R. P. Goldthwait and Dr. J. M. Schopf. Dr. 5. B. Treves provided petrographic advice and Dir. G. A. Doumani provided information con cerning the invertebrate fossils. Invaluable assistance in the fiBld was provided by Mr. L. L. Lackey, Mr. M. D. Higgins, Mr. J. Ricker, and Mr. C. Skinner. Funds for this study were made available by the Office of Antarctic Programs of the National Science Foundation (NSF grants G-13590 and G-17216). The Ohio State Univer sity Research Foundation and Institute of Polar Studies administered the project (OSURF Projects 1132 and 1258). Logistic support in Antarctica was provided by the United States Navy, especially Air Development Squadron VX6. -
Annual Meeting 2014
The Palaeontological Association 58th Annual Meeting 16th–19th December 2014 University of Leeds PROGRAMME abstracts and AGM papers Public transport to the University of Leeds BY TRAIN: FROM TRAIN STATION ON FOOT: Leeds Train Station links regularly to all major UK cities. You The University campus is a 20 minute walk from the train can get from the station to the campus on foot, by taxi or by station. The map below will help you find your way. bus. A taxi ride will take about 10 minutes and it will cost Leave the station through the exit facing the main concourse. approximately £5. Turn left past the bus stops and walk down towards City Square. Keeping City Square on your left, walk straight up FROM TRAIN STATION BY BUS: Park Row. At the top of the road turn right onto The Headrow, We advise you to take bus number 1 which departs from passing The Light shopping centre on your left. After The Light Infirmary Street. The bus runs approximately every 10 minutes turn left onto Woodhouse Lane to continue uphill. Keep going, and the journey takes 10 minutes. passing Morrisons, Leeds Metropolitan and the Dry Dock You should get off the bus just outside the Parkinson Building. boat pub heading for the large white clock tower. This is the (There is also the £1 Leeds City Bus which takes you from the Parkinson building. train station to the lower end of campus but the journey time is much longer). BY COACH: If you arrive by coach you can catch bus numbers 6,28 or 97 to the University (Parkinson Building). -
V53.2018.PSSA Abstracts
tropical to warm temperate palaeoenvironments, principally in Laurussia and the northern continental blocks. By contrast, the Waterloo Farm Lagerstätte, from which Archaeopteris notosaria was described, was situated in southern Gondwana at high palaeolatitude (currently reconstructed as being within the Devonian Antarctic Circle). This was the first evidence that cosmopolitan lowland forests had achieved a truly global distribution by the end of the Devonian. The environmental implications of this vegetative explosion have been explored by subsequent authors, particularly with regard to climate change and possible causes of the End Devonian Mass Extinction. The original description relied on portions of leafy ‘fronds’, and a single poorly preserved fertile fragment. New material has been provided by excavation of a stratigraphically lower site discovered during roadworks in the mid portion of the Witpoort Formation. Not only does this provide a slightly earlier record for the arrival of Archaeopteris in southern Africa but is also significant in that the new material provides additional anatomical information. It includes impres - sions of two whole ‘fronds’, one infertile and one fertile, as well as a fragment in which the structure of the fertile cones is clearly apparent, revealing the type material to have been incomplete. Bone lesion in a gorgonopsian fossil from the Permian of Zambia: periosteal reaction in a premammalian synapsid Kyle Kato 1, Elizabeth Rega 2, Christian Sidor 3, Adam K. Huttenlocker 1* 1Department of Integrative Anatomical Sciences, Keck School of Medicine, University of Southern California, Los Angeles, California, U.S.A. 2College of Osteopathic Medicine of the Pacific and the Graduate College of Biomedical Sciences, Western University of Health Sciences, Pomona, California, U.S.A.