DOCSLIB.ORG

Recovery of Brachiopod and Ammonoid Faunas Following the End-Permian Crisis: Additional Evidence from the Lower Triassic of the Russian Far East and Kazakhstan

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Recovery of Brachiopod and Ammonoid Faunas Following the End-Permian Crisis: Additional Evidence from the Lower Triassic of the Russian Far East and Kazakhstan Journal of Earth Science, Vol. 25, No. 1, p. 1–44, February 2014 ISSN 1674-487X Printed in China DOI: 10.1007/s12583-014-0398-6 Recovery of Brachiopod and Ammonoid Faunas Following the End-Permian Crisis: Additional Evidence from the Lower Triassic of the Russian Far East and Kazakhstan Yuri D Zakharov*, Alexander M Popov Far Eastern Geological Institute, Far Eastern Branch, Russian Academy of Sciences, Stoletiya Prospect 159, Vladivostok 690022, Russia ABSTRACT: After the End-Permian mass extinction, ammonoids reached levels of taxonomic diver- sity higher than in the Changhsingian by the Dienerian Substage of the Induan. However, brachiopods exhibit a prolonged delay in recovery, and their taxonomic diversity had not recovered to Late Permian levels even by the Olenekian. The differential patterns of recovery between these two clades may reflect fundamental differences in physiology and behavior. Brachiopods were benthic organisms that were dependent on specific trophic sources, and their general reduction in size during the Early Triassic may have been a response to a relative paucity of food resources. In contrast, ammonoids were sluggish- nektic organisms that utilized a wider range of trophic sources and that suffered no comparable size decrease during the Early Triassic. Brachiopods may have been at a disadvantage also due to vulner- abilities associated with their larval stage, during which they had to locate a suitable substrate for set- tlement. In contrast, ammonoids had no larval stage and juveniles may have been dispersed widely into favorable habitats. These factors may account for differences in the relative success of ammonoids and brachiopods at high-latitude regions following the End-Permian mass extinction: ammonoids success- fully recolonized the Boreal region during the Early Triassic whereas brachiopods did not. KEY WORDS: Lower Triassic, South Primorye, Kazakhstan, brachiopod, ammonoid, biotic recovery. 1 INTRODUCTION Early Triassic (mainly Olenekian) ammonoids are abundant The Permian-Triassic boundary (PTB) is associated with and diverse in many regions of the world, but the most diverse the largest extinction event of Phanerozoic time (e.g., Kozur, Olenekian brachiopod faunas have been discovered in the for- 2007; Yin and Zhang, 1996; Erwin, 1994). Early Triassic suc- mer USSR area (South Primorye and Mangyshlak). The goal of cessions of both benthonic and nektonic marine organisms this work is to analyze taxonomic diversity patterns of Early contain important information about the postextinction biotic Triassic articulated brachiopods and ammonoids from the for- recovery interval. However, in spite of significant progress in mer USSR (Fig. 1) in order to reconstruct biotic recovery pat- the past decades in investigation of Early Triassic brachiopods terns after the End-Permian mass extinction. (e.g., Ruban, 2009; Chen et al., 2006, 2005a, b; Shen and Shi, 1996; Shen and He, 1994; Xu and Grant, 1994; Jordan, 1993; 2 MATERIALS MacFarlan, 1992; Chen, 1983; Xu and Liu, 1983) and ammon- Original materials on Early Triassic invertebrates used for oids (e.g., Brühwiler et al., 2012a, b, 2010a, b, c, 2008; Smy- our investigation were obtained by the authors from Induan shlyaeva and Zakharov, 2012; Ware et al., 2011; Guex et al., and Olenekian sections of South Primorye during the last five 2010; Brayard et al., 2009a, b, c, 2007a, b, 2006a, b; Shigeta et years. Investigation of Olenekian brachiopods from Man- al., 2009; Brayard and Bucher, 2008; Mu et al., 2007; Shevyrev, gyshlak, Kazakhstan was made on the basis of Zakharov Y D’s 2006, 1986; Krystyn et al., 2003; Ermakova, 2002; Zakharov, collection from his Kara-Tau expedition in 1978. 1997, 1996, 1978; Dagys and Ermakova, 1996, 1988; Krystyn and Orchard, 1996; Waterhouse, 1996; Tozer, 1994; Guex, 3 LOWER TRIASSIC AMMONOID AND BRACHIO- 1978), our knowledge of these important invertebrate groups, POD BIOSTRATIGRAPHY especially Mesozoic-type Brachiopodaremains incomplete. 3.1 South Primorye (Russian Far East) The Early Triassic epiplatformal terrigenous assemblage *Corresponding author: [email protected] of South Primorye (Fig. 1) lies unconformably on the Upper © China University of Geosciences and Springer-Verlag Berlin Palaeozoic and more ancient marine and continental sedimen- Heidelberg 2014 tary and volcano-sedimentary deposits, volcanics and grani- toids (Zakharov et al., 2010; Zakharov, 1978, 1968). In many Manuscript received June 11, 2012. areas of South Primorye (e.g., Russian Island, Ussuri and Amur Manuscript accepted September 28, 2012. gulfs, Abrek Bay and Artyom area), the bulk of the Induan or Zakharov, Y. D., Popov, A. M., 2014. Recovery of Brachiopod and Ammonoid Faunas Following the End-Permian Crisis: Addi- tional Evidence from the Lower Triassic of the Russian Far East and Kazakhstan. Journal of Earth Science, 25(1): 1–44, doi:10.1007/s12583-014-0398-6 2 Yuri D Zakharov and Alexander M Popov Induan to earliest Olenekian Lazurnaya Bay Formation con- posed only on Russian Island and consists of sandstone with sists of shallow-marine shelf sediments. These are predomi- thick lenses of sandy limestone. It is represented by a single nantly conglomerate and sandstone with lenses of sandy zone, the Tirolites-Amphistephanites Zone (Figs. S3 and S6). limestone-coquina dominated by bivalves. The lower part of The Zhitkov Cape Formation of Smithian to Spathian age the Induan is represented by the Tompophiceras ussuriense represents deeper shelf facies characterised by the dominance of Zone, its upper part corresponds to the Gyronites subdharmus mudstone and siltstone with numerous calcareous-marly nodules Zone (Figs. S1 and S2). and lenses and thin, irregular beds of sandstone. The lower part The Tobizin Cape Formation, of Smithian age, is present of the formation, which is found in the West Ussuri Gulf-Strelok on Russian Island and on the western coast of Ussuri Gulf, Strait area (Figs. S1 and S7), is represented by the “Hedenstroe- where it conformably overlies the Lazurnaya Bay Formation. It mia” bosphorensis and Anasibirites nevolini zones and is is represented by sandstone with lenses of coquinoid calcare- Smithian in age. The upper part of the formation was formed in ous sandstone with mainly cephalopod and bivalve remains. this area during the Early (Tirolites-Amphistephanites Zone) and On Russian Island, the following zones can be recognized: Late (Neocolumbites insignis and Subfengshanites multiformis “Hedenstroemia” bosphorensis below and Anasibirites nevolini zones) Spathian. On Russian Island (Figs. S5–S7) and the west- above (Figs. S3 and S4). ern coast of Amur Gulf (Zakharov et al., 2005), only the upper The Schmidt Cape Formation of early Spathian age is ex- Olenekian Zhitkov Cape Formation is present. However, the Razdol naia ° 60° 80° N 80° N 60° 44 N ’ ()a USSURIYSK Rakovka Rakovka ° hka ac 40 N Komarovkani Iv 2 Barsukovka ° Kaimanovka 40 N aia 1 Perevozn Study area 1 000 km Borisovka Razdol'noye Razdol'naya 70° 90° 110° E a Zavodskoy k Bol. Kiparisovka ov yom rt A Artyom Vol’no-Nadezhdinskoe Zanadvorovka Steklyanukha Tavrichanka Uglovoe Smolyaninovo Bogataya Sukhodol Vladivostok Bol’shoi Kamen’ 1 G Amur Gulf 5 Ussuri ulf (b) 6 43° 43° 3 8 Popov Isl. Russian Island Fokino 4 2 Reyneke Isl. 7 Dunai Sea of Japan Putyatin Isl. 0 15 km Askol’d Isl. 132°E Figure 1. (a) Location of study areas in the former USSR. 1. South Primorye and 2. Mangyshlak. (b) Study sections in South Pri- morye area. 1. Seryj-Tri Kamnya; 2. Abrek; 3. Konechnyj; 4. Tobizin; 5. Ayax-Balka; 6. Zhitkov; 7. Golyj; 8. Paris. Recovery of Brachiopod and Ammonoid Faunas Following the End-Permian Crisis 3 Olenekoceras-bearing part of the Spathian on the western coast gen. and sp. nov. B), Norellidae (Piarorhynchella? sp. nov.) of Amur Gulf is characterised mainly by fine-grained, striped and Rhynchonellidae (Abrekia sulcata, Rhynchonellidae gen. sandstone (Atlasov Cape Formation). and sp. nov.). For detailed biostratigraphic distribution data, The Karazin Cape Formation represents the Anisian eve- see Figs. S1 and S2. Among these brachiopod families, the rywhere in South Primorye. No benthic forms of allochthonous Pontisiidae and the Wellerellidae are Paleozoic-type ones, but origin have been found in this formation and therefore it seems the Norellidae and the Rhynchonellidae seem to have origi- to have accumulated under offshore environmental conditions. nated during the Early Triassic (Induan). It composed mainly of fucoid sandstone with large calcareous Newly found Dienerian brachiopod shells in South Pri- septarian nodules. The Lower Anisian in South Primorye is morye, including specimens of Abrekia, first described by A. S. represented by the Ussuriphyllites amurensis and Leiophyllites Dagys (see Supplement), are generally very small as compared pradyumna zones (Figs. S5–S7). with Late Permian ones (Tables S2–S4)—their maximal shell The detailed lithological succession and distribution of length is only 7.8 mm. brachiopod and ammonoid species in the Lower Triassic of South Primorye are shown in some figures (Figs. S1 and S8). 3.1.3 “Hedenstroemia” bosphorensis Zone (Smithian) Figure 2 illustrates the stratigraphic distribution of brachiopods The early Smithian “Hedenstroemia” bosphorensis Zone and ammonoids at the family and order levels (within the Up- has been subdivided into a lower unit (“Gyronites separatus” per Changhsingian-Spathian interval). (=Ussuriflemingites abrekensis) beds) and an upper unit (Eu- flemingites prynadai beds) (Zakharov, 1997). The
Load more

Recommended publications
  • Construction, Main Projects
    1520 GROUP OF COMPANIES PRESENTATION ABOUT COMPANY The 1520 Group of Companies is one of the largest production and construction holdings in Russia concentrated on the design and construction of railways. Group’s global reach covers the entire Russian railway track with a track gauge of 1,520 mm — from the Baltics to the Pacific Ocean. The Group employs over 20 thousand people. The 1520 Group of Companies focuses on complex construction, renovation and overhaul of railway infrastructure facilities of any complexity level, design and survey work, production of structures, assemblies and parts, construction and maintenance of highways. The 1520 Group of Companies is participating in the modernization and development of the «Eastern Polygon» — the Trans-Siberian and Baikal-Amur Mainlines, in the renovation and development of the Moscow Transport Hub as well as in other major projects intended for the integrated development of the railway infrastructure in the Russian Federation. The 1520 Group of Companies is a highly professional team of managers, designers, engineers and experts specialized in complicated construction- related fields who have unique experience in the railway and road construction in any climatic conditions. The 1520 Group of Companies is headed by Alexey Krapivin. 2 The 1520 GROUP OF COMPANIES is comprised of the following companies and enterprises: PROJECT DIVISION: ROSZHELDORPROJECT Moscow and 18 branches across Russia LENGIPROTRANS Saint-Petersburg DALGIPROTRANS Khabarovsk CONSTRUCTION DIVISION: BAMSTROYMEKHANIZATZIYA
    [Show full text]
  • Eichwald, Eduard. Lethaea Rossica Ou Paléontologie De La Russie . Second Volume, Période Moyenne. Seconde Section. 1995
    S и. 1(116.0 ВСЕСОЮЗНЫЙ НАУЧНО ИССЛЕДОВАТЕЛЬСКИЙ ГЕОЛОГИЧЕСКИЙ ИНСТИТУТ (ВСЕГЕИ ) Д.Д .К И ПАРИ С ОБА ПАЛЕОНТОЛОГИЧЕСКОЕ ОБОСНОВАНИЕ СТРАТИГРАФИИ ТРИАСОВЫХ ОТЛОЖЕНИЙ ПРИМОРСКОГО КРАЯ I шшишшшшшшшиивш/шшшш МИНИСТЕРСТВО ГЕОЛОГИИ СССР ВСЕСОЮЗНЫЙ НАУЧНО-ИССЛЕДОВАТЕЛЬСКИИ /' ГЕОЛОГИЧЕСКИИ ИНСТИТУТ (ВСЕГЕИ) НОВАЯ СЕРИЯ ' ТОМ 181 Л. Д. КИПАРИСОВА ПАЛЕОНТОЛОГИЧЕСКОЕ ОБОСНОВАНИЕ СТРАТИГРАФИИ ТРИАСОВЫХ ОТЛОЖЕНИЙ ПРИМОРСКОГО КРАЯ Часть 2 ПОЗДНЕТРИАСОВЫЕ ДВУСТВОРЧАТЫЕ МОЛЛЮСКИ И ОБЩАЯ СТРАТИГРАФИЯ ИЗДАТЕЛЬСТВО «НЕДРА» МОСКВА 1972 УДК 551.7(571.63) Кипарисова Л. Д. Палеонтологическое обоснование стратигра- фии триасовых отложений Приморского края. Часть 2. Позднетриасовые двустворчатые моллюски и общая стратиграфия. ВСЕГЕИ, нов. серия, т. 181, М., «Недра», 1972 г., 248 с. Работа представляет собой вторую часть монографии «Палеонтологи- ческое обоснование стратиграфии триасовых отложений Приморского края». Первая часть монографии — «Головоногие моллюски» опубликована в 1961 г. Книга содержит описание двустворчатых моллюсков позднего триаса. Кроме того, в ней приводится описание опорных разрезов и общая схема расчленения триасовых отложений Приморского края. Позднетриасовые двустворчатые моллюски Приморского края описы- ваются впервые; впервые анализируется их состав, стратиграфическое и географическое распространение. Устанавливается их общность с позднетриасовыми двустворками Япо- нии и Северо-Востока СССР; только самый ранний комплекс, характери- зующий тегючинскне известняки, имеет явно альпийский характер. Дается схема корреляции триасовых отложений
    [Show full text]
  • Anisian (Middle Triassic) Ammonoids from North America: Quantitative Biochronology and Biodiversity
    Anisian (Middle Triassic) ammonoids from North America: quantitative biochronology and biodiversity Claude Monnet and Hugo Bucher Paläontologisches Institut und Museum, Universität Zürich, Karl Schmid Strasse 4, CH-8006 Zürich, Switzerland email: [email protected]; [email protected] ABSTRACT: This study synthesizes and revises the ammonoid zonations as well as their correlation with each other for western Nevada (USA), British Columbia (Canada), and the Sverdrup Basin (Canada) by utilizing the unitary association method. Based on a standard- ized taxonomy, the Anisian in the studied areas contains 13, 10, and 3 zones and a total of 174, 90, and 7 species, respectively. The zonations are correlated by means of a ‘common taxa zonation’, which includes all taxa common to the studied basins. This leads to new and more precise correlations, which are at slight variance with those of the literature. Hence, the Buddhaites hagei Zone (Canada) corre- lates only with the Intornites mctaggarti Subzone (Nevada) and not with the entire Acrochordiceras hyatti Zone (Nevada). The Tetsaoceras hayesi Zone (Canada) appears to correlate with the Unionvillites hadleyi Subzone (Nevada) of the hyatti Zone and not with the Nevadisculites taylori Zone. The Hollandites minor Zone (Canada) more than likely correlates with the taylori Zone (Nevada) rather than the Balatonites shoshonensis Zone as is usually acknowledged. The unitary association method enables us to quantify the diachronism of the studied taxa, which affects about 67% of the genera and 18% of the species common to Nevada and British Columbia. Therefore, this diachronism is significant and its value for correlation should not be overlooked.
    [Show full text]
  • Late Permian to Middle Triassic Palaeogeographic Differentiation of Key Ammonoid Groups: Evidence from the Former USSR Yuri D
    Late Permian to Middle Triassic palaeogeographic differentiation of key ammonoid groups: evidence from the former USSR Yuri D. Zakharov1, Alexander M. Popov1 & Alexander S. Biakov2 1 Far-Eastern Geological Institute, Russian Academy of Sciences (Far Eastern Branch), Stoletija Prospect 159, Vladivostok, RU-690022, Russia 2 North-East Interdisciplinary Scientific Research Institute, Russian Academy of Sciences (Far Eastern Branch), Portovaja 16, Magadan, RU-685000, Russia Keywords Abstract Ammonoids; palaeobiogeography; palaeoclimatology; Permian; Triassic. Palaeontological characteristics of the Upper Permian and upper Olenekian to lowermost Anisian sequences in the Tethys and the Boreal realm are reviewed Correspondence in the context of global correlation. Data from key Wuchiapingian and Chang- Yuri D. Zakharov, Far-Eastern Geological hsingian sections in Transcaucasia, Lower and Middle Triassic sections in the Institute, Russian Academy of Sciences (Far Verkhoyansk area, Arctic Siberia, the southern Far East (South Primorye and Eastern Branch), Vladivostok, RU-690022, Kitakami) and Mangyshlak (Kazakhstan) are examined. Dominant groups of Russia. E-mail: [email protected] ammonoids are shown for these different regions. Through correlation, it is doi:10.1111/j.1751-8369.2008.00079.x suggested that significant thermal maxima (recognized using geochemical, palaeozoogeographical and palaeoecological data) existed during the late Kun- gurian, early Wuchiapingian, latest Changhsingian, middle Olenekian and earliest Anisian periods. Successive expansions and reductions of the warm– temperate climatic zones into middle and high latitudes during the Late Permian and the Early and Middle Triassic are a result of strong climatic fluctuations. Prime Middle–Upper Permian, Lower and Middle Triassic Bajarunas (1936) (Mangyshlak and Kazakhstan), Popov sections in the former USSR and adjacent territories are (1939, 1958) (Russian northern Far East and Verkhoy- currently located in Transcaucasia (Ševyrev 1968; Kotljar ansk area) and Kiparisova (in Voinova et al.
    [Show full text]
  • The Magnetobiostratigraphy of the Middle Triassic and the Latest Early Triassic from Spitsbergen, Arctic Norway Mark W
    Intercalibration of Boreal and Tethyan time scales: the magnetobiostratigraphy of the Middle Triassic and the latest Early Triassic from Spitsbergen, Arctic Norway Mark W. Hounslow,1 Mengyu Hu,1 Atle Mørk,2,6 Wolfgang Weitschat,3 Jorunn Os Vigran,2 Vassil Karloukovski1 & Michael J. Orchard5 1 Centre for Environmental Magnetism and Palaeomagnetism, Geography, Lancaster Environment Centre, Lancaster University, Bailrigg, Lancaster, LA1 4YQ, UK 2 SINTEF Petroleum Research, NO-7465 Trondheim, Norway 3 Geological-Palaeontological Institute and Museum, University of Hamburg, Bundesstrasse 55, DE-20146 Hamburg, Germany 5 Geological Survey of Canada, 101-605 Robson Street, Vancouver, BC, V6B 5J3, Canada 6 Department of Geology and Mineral Resources Engineering, Norwegian University of Sciences and Technology, NO-7491 Trondheim, Norway Keywords Abstract Ammonoid biostratigraphy; Boreal; conodonts; magnetostratigraphy; Middle An integrated biomagnetostratigraphic study of the latest Early Triassic to Triassic. the upper parts of the Middle Triassic, at Milne Edwardsfjellet in central Spitsbergen, Svalbard, allows a detailed correlation of Boreal and Tethyan Correspondence biostratigraphies. The biostratigraphy consists of ammonoid and palynomorph Mark W. Hounslow, Centre for Environmental zonations, supported by conodonts, through some 234 m of succession in two Magnetism and Palaeomagnetism, adjacent sections. The magnetostratigraphy consists of 10 substantive normal— Geography, Lancaster Environment Centre, Lancaster University, Bailrigg, Lancaster, LA1 reverse polarity chrons, defined by sampling at 150 stratigraphic levels. The 4YQ, UK. E-mail: [email protected] magnetization is carried by magnetite and an unidentified magnetic sulphide, and is difficult to fully separate from a strong present-day-like magnetization. doi:10.1111/j.1751-8369.2008.00074.x The biomagnetostratigraphy from the late Olenekian (Vendomdalen Member) is supplemented by data from nearby Vikinghøgda.
    [Show full text]
  • Download Download
    I I Rivista Italiana di Paleon roìor,iaeSrrltrqrriia I rolur ;;- -, | f;-' "oo, l UPPER OLENEKIAN (SPATHIAN) AMMONOIDS FROM CHIOS (LO\T/ER TRIASSIC, GREECE): TAXONOMY AND STRATIGRAPHIC POSITION. DOROTHEE MERTMANN' & VOLKER TACOBSHAGEN' Receh:ed July 1st,2002; accepted June 25,20A3 Key-raords: Ammonoidea, Triassic, Olenekian, Chios, Greece. There, the fossils are included in red limestones of rhe Hallstatt facies (Marmarorrapeza. Formation). Renz & Abstact. On the Greek island of Chios, Lower Triassic sections Renz (1948) have delivered rich ammonoid faunas from red limestones of the Hall- published a palaeontological description of statt facies. Here, we present a palaeontological description of Spatl-rian its ammonoids in a monograph. Bender (1970) was the ammonoids, which have been collected during two field campaigns by first to collect fossils from measured sections (CM i, kalian-German teams (Assereto et al. 1980; Gaetani et al. 1992). The II, III) in order to establish coordinared ammonoid and ammonoid assocìations are composed of Ceratitida and Phyllocerati- conodont stratigraphies. This author recognized, more- da, each order containing species of the Noritaceae, Hedenstroemia- ceae, Xenodiscaceae, Dinaritaceae, Pinacocerataceae, Phyllocerataceae, over, that the Spathian beds are overlain by lowermost Ptychitaceae, lJssuritaceae and Megaphyllitaceae. Long ranging gen- Anisian ones s/ith ammonoids comparable with faunas era, e.g. Procarnites and Leioplrylliter, are present as well as endemic of the so-called Hydaspian substage from the Himalayas ones, e.g. Cbiotites, which occurs exclusively in the Lower Trìassic of and Timor. Furthermore, Jacobshagen & Tietze (1974) Chios. The ammonoìd association is indicative of the Probungarites- described an associarion of both Spathian and Lower Subcolwmbites zone sensu Kumn.rel (1973a,).
    [Show full text]
  • Exceptional Vertebrate Biotas from the Triassic of China, and the Expansion of Marine Ecosystems After the Permo-Triassic Mass Extinction
    Earth-Science Reviews 125 (2013) 199–243 Contents lists available at ScienceDirect Earth-Science Reviews journal homepage: www.elsevier.com/locate/earscirev Exceptional vertebrate biotas from the Triassic of China, and the expansion of marine ecosystems after the Permo-Triassic mass extinction Michael J. Benton a,⁎, Qiyue Zhang b, Shixue Hu b, Zhong-Qiang Chen c, Wen Wen b, Jun Liu b, Jinyuan Huang b, Changyong Zhou b, Tao Xie b, Jinnan Tong c, Brian Choo d a School of Earth Sciences, University of Bristol, Bristol BS8 1RJ, UK b Chengdu Center of China Geological Survey, Chengdu 610081, China c State Key Laboratory of Biogeology and Environmental Geology, China University of Geosciences (Wuhan), Wuhan 430074, China d Key Laboratory of Evolutionary Systematics of Vertebrates, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, Beijing 100044, China article info abstract Article history: The Triassic was a time of turmoil, as life recovered from the most devastating of all mass extinctions, the Received 11 February 2013 Permo-Triassic event 252 million years ago. The Triassic marine rock succession of southwest China provides Accepted 31 May 2013 unique documentation of the recovery of marine life through a series of well dated, exceptionally preserved Available online 20 June 2013 fossil assemblages in the Daye, Guanling, Zhuganpo, and Xiaowa formations. New work shows the richness of the faunas of fishes and reptiles, and that recovery of vertebrate faunas was delayed by harsh environmental Keywords: conditions and then occurred rapidly in the Anisian. The key faunas of fishes and reptiles come from a limited Triassic Recovery area in eastern Yunnan and western Guizhou provinces, and these may be dated relative to shared strati- Reptile graphic units, and their palaeoenvironments reconstructed.
    [Show full text]
  • BR IFIC N° 2505 Index/Indice
    BR IFIC N° 2505 Index/Indice International Frequency Information Circular (Terrestrial Services) ITU - Radiocommunication Bureau Circular Internacional de Información sobre Frecuencias (Servicios Terrenales) UIT - Oficina de Radiocomunicaciones Circulaire Internationale d'Information sur les Fréquences (Services de Terre) UIT - Bureau des Radiocommunications Part 1 / Partie 1 / Parte 1 Date/Fecha: 21.10.2003 Description of Columns Description des colonnes Descripción de columnas No. Sequential number Numéro séquenciel Número sequencial BR Id. BR identification number Numéro d'identification du BR Número de identificación de la BR Adm Notifying Administration Administration notificatrice Administración notificante 1A [MHz] Assigned frequency [MHz] Fréquence assignée [MHz] Frecuencia asignada [MHz] Name of the location of Nom de l'emplacement de Nombre del emplazamiento de 4A/5A transmitting / receiving station la station d'émission / réception estación transmisora / receptora 4B/5B Geographical area Zone géographique Zona geográfica 4C/5C Geographical coordinates Coordonnées géographiques Coordenadas geográficas 6A Class of station Classe de station Clase de estación Purpose of the notification: Objet de la notification: Propósito de la notificación: Intent ADD-addition MOD-modify ADD-additioner MOD-modifier ADD-añadir MOD-modificar SUP-suppress W/D-withdraw SUP-supprimer W/D-retirer SUP-suprimir W/D-retirar No. BR Id Adm 1A [MHz] 4A/5A 4B/5B 4C/5C 6A Part Intent 1 103047968 BEL 1377.2500 HOUTHALEN HELCHTEREN MIJNSITEBEL 5E22'19" 51N1'30"
    [Show full text]
  • Anisian Ammonoids from Qingyan
    1661-8726/08/020546-16 Swiss J. Geosci. 101 (2008) 547–562 DOI 10.1007/s00015-008-1274-0 Birkhäuser Verlag, Basel, 2008 Anisian ammonoids from Qingyan, southwestern China: biostratigraphical implications for the age of the Qingyan Formation FRANK STILLER1, * & HUGO BUCHER 2 Key words: ammonoids, Anisian, Qingyan Formation, southwestern China, taxonomy, biostratigraphy ABSTRACT The Upper Qingyan Formation in the northeastern vicinity of Qingyan, Gui- blage correlates with the Billingsites cordeyi Subzone in northwestern Nevada zhou Province, southwestern China, yields an ammonoid sequence comprising and the Rieppelites cimeganus Zone in Lombardy. The middle/late Anisian a latest middle Anisian assemblage in its lower part followed by an earliest late boundary is situated in the upper middle part of the highly fossiliferous sec- Anisian assemblage in its middle and upper parts. Age-diagnostic genera of tion at Leidapo. Judging from re-study of available and figured specimens, the lower assemblage include Bulogites and Acrochordiceras; additional gen- reports of stratigraphically younger taxa such as Paraceratites trinodosus from era comprise Proarcestes, Sageceras, and Beyrichitinae gen. indet. This assem- the Upper Qingyan Formation of this area are based on misidentifications. blage well correlates with the Bulogites mojsvari Subzone in northwestern Ne- The time interval represented by the Qingyan Formation at its type locality is vada and the Bulogites zoldianus Zone in Lombardy and Hungary. The upper not equal to the Anisian as has long been supposed in Chinese stratigraphy. assemblage comprises the age-diagnostic genera Rieppelites and Judicarites; An earliest late Anisian age for the uppermost strata of the Qingyan Forma- additional characteristic genera include Ptychites and Gosauites.
    [Show full text]
  • Paleontological Contributions
    THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS May 15, 1970 Paper 47 SIGNIFICANCE OF SUTURES IN PHYLOGENY OF AMMONOIDEA JURGEN KULLMANN AND JOST WIEDMANN Universinit Tubingen, Germany ABSTRACT Because of their complex structure ammonoid sutures offer best possibilities for the recognition of homologies. Sutures comprise a set of individual elements, which may be changed during the course of ontogeny and phylogeny as a result of heterotopy, hetero- morphy, and heterochrony. By means of a morphogenetic symbol terminology, sutural formulas may be established which show the composition of adult sutures as well as their ontogenetic development. WEDEKIND ' S terminology system is preferred because it is the oldest and morphogenetically the most consequent, whereas RUZHENTSEV ' S system seems to be inadequate because of its usage of different symbols for homologous elements. WEDEKIND ' S system includes only five symbols: E (for external lobe), L (for lateral lobe), I (for internal lobe), A (for adventitious lobe), U (for umbilical lobe). Investigations on ontogenetic development show that all taxonomic groups of the entire superorder Ammonoidea can be compared one with another by means of their sutural development, expressed by their sutural formulas. Most of the higher and many of the lower taxa can be solely characterized and arranged in phylogenetic relationship by use of their sutural formulas. INTRODUCTION Today very few ammonoid workers doubt the (e.g., conch shape, sculpture, growth lines) rep- importance of sutures as indication of ammonoid resent less complicated structures; therefore, phylogeny. The considerable advances in our numerous homeomorphs restrict the usefulness of knowledge of ammonoid evolution during recent these features for phylogenetic investigations.
    [Show full text]
  • Abstracts and Program. – 9Th International Symposium Cephalopods ‒ Present and Past in Combination with the 5Th
    See discussions, stats, and author profiles for this publication at: https://www.researchgate.net/publication/265856753 Abstracts and program. – 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th... Conference Paper · September 2014 CITATIONS READS 0 319 2 authors: Christian Klug Dirk Fuchs University of Zurich 79 PUBLICATIONS 833 CITATIONS 186 PUBLICATIONS 2,148 CITATIONS SEE PROFILE SEE PROFILE Some of the authors of this publication are also working on these related projects: Exceptionally preserved fossil coleoids View project Paleontological and Ecological Changes during the Devonian and Carboniferous in the Anti-Atlas of Morocco View project All content following this page was uploaded by Christian Klug on 22 September 2014. The user has requested enhancement of the downloaded file. in combination with the 5th International Symposium Coleoid Cephalopods through Time Abstracts and program Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum, Universität Zürich Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 2 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ 9th International Symposium Cephalopods ‒ Present and Past in combination with the 5th International Symposium Coleoid Cephalopods through Time Edited by Christian Klug (Zürich) & Dirk Fuchs (Sapporo) Paläontologisches Institut und Museum Universität Zürich, September 2014 3 Cephalopods ‒ Present and Past 9 & Coleoids through Time 5 Zürich 2014 ____________________________________________________________________________ Scientific Committee Prof. Dr. Hugo Bucher (Zürich, Switzerland) Dr. Larisa Doguzhaeva (Moscow, Russia) Dr. Dirk Fuchs (Hokkaido University, Japan) Dr. Christian Klug (Zürich, Switzerland) Dr. Dieter Korn (Berlin, Germany) Dr. Neil Landman (New York, USA) Prof. Pascal Neige (Dijon, France) Dr.
    [Show full text]
  • Chondrichthyan Teeth from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA)
    Zurich Open Repository and Archive University of Zurich Main Library Strickhofstrasse 39 CH-8057 Zurich www.zora.uzh.ch Year: 2019 Chondrichthyan teeth from the Early Triassic Paris Biota (Bear Lake County, Idaho, USA) Romano, Carlo ; Argyriou, Thodoris ; Krumenacker, L J Abstract: A new, diverse and complex Early Triassic assemblage was recently discovered west of the town of Paris, Idaho (Bear Lake County), USA. This assemblage has been coined the Paris Biota. Dated earliest Spathian (i.e., early late Olenekian), the Paris Biota provides further evidence that the biotic recovery from the end-Permian mass extinction was well underway ca. 1.3 million years after the event. This assemblage includes mainly invertebrates, but also vertebrate remains such as ichthyoliths (isolated skeletal remains of fishes). Here we describe first fossils of Chondrichthyes (cartilaginous fishes) from the Paris Biota. The material is composed of isolated teeth (mostly grinding teeth) preserved on two slabs and representing two distinct taxa. Due to incomplete preservation and morphological differences to known taxa, the chondrichthyans from the Paris Biota are provisionally kept in open nomenclature, as Hybodontiformes gen. et sp. indet. A and Hybodontiformes gen. et sp. indet. B, respectively. The present study adds a new occurrence to the chondrichthyan fossil record of the marine Early Triassic western USA Basin, from where other isolated teeth (Omanoselache, other Hybodontiformes) as well as fin spines of Nemacanthus (Neoselachii) and Pyknotylacanthus (Ctenachanthoidea) and denticles have been described previously. DOI: https://doi.org/10.1016/j.geobios.2019.04.001 Posted at the Zurich Open Repository and Archive, University of Zurich ZORA URL: https://doi.org/10.5167/uzh-170655 Journal Article Accepted Version The following work is licensed under a Creative Commons: Attribution-NonCommercial-NoDerivatives 4.0 International (CC BY-NC-ND 4.0) License.
    [Show full text]