The Pausandra Radlk.

by

J. Lanjouw

Radlk. Tribe The genus Pausandra belongs to the Cluytieae of the . It was described by Radlkofer in 1870 in Flora The is based Thouinia Mori- LIII pp. 79—95. genus on siana of Casaretto. In his Radlkofer discussed at paper this the length that species does not belong to Sapindaceous genus but of the Thouinia, represents a new genus Euphorbiaceae. As at that time female flowers were unknown Radlkofer stated that the of the still systematic position new genus was doubtful, but that subtribe of the most probably it should belong to a new Two described the in Jatropheae. new species were in genus 1873 by Baillon, P. Trianae Baill. based on Trianae

and P. Martinii Müll. Arg. which was published in 1864, Baill. based on material and described Bail- very young erroneously by will be discussed below. the lon as being 3-merous, as He placed

in the of which is not genus affinity Argithamnia Sw., certainly this is different both in habit and in flower- right as genus quite characters. A fourth species was added by Müller Arg. in in Flora Brasiliensis where he 1874 XI. II., inserted the genus in the Radlkofer. same group as was suggested by No more had been described when Pax in his species published 1911 monograph of the Tribe Cluytieae Pax in Engler , Das Pflanzen- reich IV. III. He inserted the Pausandra Radlk, with 147. genus the Griff, and Ricinodendron Müll. genera Givotia Arg. in a new subtribe Ricinodendrinae Pax. I think that this is the right for the it could be in position genus, though placed a separate subtribe for its leaves and the penninerved, glanduliferous cap- sular fruits. It was a pity that Pax published this monograph without studying the original material. He now copied Bail- lon’s bad descriptions and the lack of a thorough study on the caused the of several in genus publication superfluous species recent P. Pax et K. Hoffm. and P. years. quadriglandulosa ex- described in and the P.torris Standley 1919 1929 are same as Trianae (Müll. Arg.) Baill. P. flagellorhachis Lanj. is identic with 759

P. Martinii Baill., while it was proved that the latter species is

not trimerous. P. integrifolia Lanj. could not be maintained in the the Ducke in genus. Only two new species published by 1925

were truly new ones. Moreover three new species were recognized made Krukoff Brazil. for in the recent collections by in It is all these that seemed desirable reasons it to me highly to give a of this several the old and new treatment genus. Perhaps of new be find often small differen- species can united, as one can only

but for the I think advisable them ces, present it to keep separate.

Pausandra Radlk, has been described to be dioecious, but re-

cently it has been proved in some species that they are monoe-

cious, so it is probable that most of them are under special cicumstances.

Tlhe is restricted South and Central America. genus to tropical West No specimens were collected in the Indian Islands. I have studied material of the following herbaria: Brussel

(B.), Chicago, Field Museum (C.), Berlin-Dahlem (D.), Geneve (G.), Kew (K.), Leiden (L.), Paris (P.), and U t r e c h t( U.). For the hospitality and the loan of the

material I am indebted to the Directors of these Institutes.

Pausandra Radik, in Flora LIII (1870), p. 92, t. 2; Mull. Arg.

in FI. Bras. XL t. Bentham Hook. Gen. 2. (1874) p. 503, 99; et III. Pax in Prantl, Pflanzenfamilien pi. (1880) p. 298; Engler u. 2nd ed. in III. J. (1890) p. 87; id. vol. 19c (1931) p. 173; Pax Pflanzenreich Engler, Das IV. 147. III. (1911) p. 41, fig. 13, 14;

Diet, Phan. V. — Pausandra Radlk. Lemee, plantes (1934) p. 84;

sect. in Kew Bull. Eupausandra Lanj. 1932 p. 184.

Flowers dioecious or monoecious. Male flower: Calyx unequal imbricate. Corolla less 5- rarely 3-Iobed, more or gamopetalous, inside barbate at the throat, j-lobed, lobes imbricate. Disk extra- sometimes Stamens staminal, urceolate, deeply 5-lobed. 3 —7, and anthers usually 6, 5 epipetalous 1 central; oblong, longitudi- dehiscent. Female flower: nally Rudimentary ovary o. Calyx j- lobed. Petals free inside the middle. j, (or connate?), barbate at Disk urceolate or j-lobed. Ovary hairy, 3-celled; 1 ovule in each cell. 2-lobed. Styles 3, Capsule 3-coccous, loculicidally dehiscent;

seeds caruncled. — oblong, more or less marmorate, Small trees,

laticiferous. Leaves petiolate, bistipulate, penninerved, often cu-

neate-spathulate, glanduliferous serrate-dentate, biglandular at the base. All of the when often parts , especially young, more less with dibrachiate with hairs. or clothed hairs, rarely simple of both less Inflorescences sexes spicate sometimes more or pani- 760

culate; flowers in dichasial glomerules, bibracteolate, in the axils of biglandular bracts. Distribution Central and South : Species 9. Tropical

America, not in the West Indian Islands.

Key to the species.

Side-nerves fewer than Limb much shorter than 1 a. 23. usually

30 cm 2 b. Side-nerves than Limb than usually more 23. usually longer

30 cm 7 male flowers 2 a. Only present 3 female flowers 6 b. Only present the base. 3 a. Leaves not long cuneate to Branches, petioles and midrib hirsute hirsuta patently 9. P. Lanj.

b. Leaves cuneate the base. Plants not long to patently hirsute 4

Petioles — Limb at both 4a. very short, 1 i.j cm long. glabrous sides. Disk deeply j-lobed 8. P. macropetala Ducke b. Petioles much than Limb usually longer i.j cm. sparsely clothed with dibrachiate hairs beneath. Disk urceolate, undulate slightly 5 much than. 5a. Spikes flagelliform, gracile, usually longer 10 cm; internodes between the of flowers glomerules usually 2—4

cm long 6. P. Martini! Baill. and than b. Spikes not flagelliform gracile, usually not longer between of flowers 10 cm. Internodes the glomerules usually

much shorter 1. P. Morisiana (Casar.) Radik.

Leaves at both shorter than side- 6a. glabrous sides, 30 cm; c. 20 Ducke nerves. Disk deeply j-lobed 8. P. .... macropetala b. Leaves sparsely clothed with appressed hairs beneath, longer than side-nerves. Disk undu- 30 cm; c. 30 urceolate, slightly late P. sericea 7. Lanj.

of the leaves serrate with subhelici- 7a. Margin upwards curved, form teeth. Basal (See fig. ta) glands usually 4 P. Trianae 3. (Mull. Arg.) Baill.

b. Teeth of the margin not upwards curved. (See fig. ib) Basal

glands 2 8

8a. cm Glomerules Inflorescences $, 7—11 long. 3-flowered ....

2. P. megalophylla Mull. Arg b. Inflorescences than Glomerules $, longer 20 cm. 6—40-

flowered 9 6—12-flowered. Internodes 9a. Glomerules c. short P. Ducke 4. macrostachya b. Glomerules than very densely flowered, more 20, usually c. 761

Internodes rather densiflora 30—4°- long j. P. Lanj.

Pausandra Radik, i. Morisiana (Casaretto) in Flora LIII (1870)

Mull. in Mart. XI. p. 92, t. II; Arg. Fl. Bras. 2 (1874) p. 503,

t. Pax in Das Pflanzenreich IV. Ill 707, 99 > Engler, 147. (1911)

— Thouinia Morisiana p. 42, no, fig. 13 A-B, 14; Casaretto, Novar. Brasil. Deced. IX Stirp. (1845) p. 75; Walpers Repert.

Bot. V (1845/46) p. 365. small branches clothed with dibrachiate A tree. Young appres- sed hairs, glabrescent; bark dark reddish brown, lenticellate. Pe- tioles —6 the (1) 2.5 cm olng, longitudinally striate, at top trans- versely rugulose and slightly thickened, sparsely clothed with Limb appressed hairs, glabrescent. (10) 16—30 cm long, (3) 6 —10

cm broad, chartaceous or subcoriaceous, oblanceolate or oblanceo- the late-oblong, shortly acuminate at apex, long cuneate to the dark-brown and much and base, glabrous above, paler very spar- clothed with hairs basal sely appressed beneath; glands two,

small, shortly stipitate, subpatelliform, margin glanduliferous ser- side-nerves midrib in the rate-dentate; 15 —21, slightly impressed basal and in the of the limb part slightly prominent upper part

above, strongly prominent beneath, side nerves prominent at both sides. Inflorescence S spicate, usually 1 rarely 2 in the axils of the rhachis clothed leaves, 5 —10 (26) cm long, densely with appressed dibrachiate hairs, flowers in dichasial, few-flowe-

red glomerules at short distances along the rhachis in the axils of small, hardly 1 mm long, triangular bracts, which are hairy outside; flowerbuds nearly sessile, subglobose, bibracteolate, brac-

teoles lanceolate, hairy outside. Calyx thick, j-lobed, lobes une-

qual, triangular and subacute or oblong and obtuse, glabrous.

Petals connate the rotundate 5, whitish, oblong, at base, or slightly the ciliolate the emarginate at apex, minutely along margin,

glabrous outside, and at about 1/4 from the base rufous-barbate

inside. Disk urceolate, slightly undulate. Stamens 6, slightly shor- than the anthers the ter petals; ovate, subcordate at base, introrse, with rather broad connective. Inflorescence a $ 7—8 cm long, outside appressed pubescent. Sepals ovate, subacute, nairy, 4 mm Petals obtuse. long. 5 mm long, free, rhombeoid-lanceolate, Ovary Fruit seeds 3-celled, pubescent. c. 22 mm high; brownish, obscurely

marmorate, c. 16 mm long and n mm broad, caruncled.

Distribution: Southern Brazil.

Rio

[D.], 3 fl.; Sello 599! [D.], 3 fl.); Rio de Janeiro, Serra da Estrella fl. (Casaretto 695! [G.], 3 Type of the genus!); Buger Creek near Blumenau, Santa Catharina (Ule 1190! [D.], 9 fl.); Brazil without precise locality (Luschnath s.n.l [D.K.P.], 3 fl.; Riedel s.n.! [P.], 3 fl.; Burohell 3825! [B.P.], mouth of 3 fl.); Amazonas, near Rio Embira, tributary of Rio Tarauaca fl. A with abnormal (Krukoff 4797! [U.], very young 3 specimen which large leaves, up to 40 cm long, probably belongs to this species).

2. Pausandra megalophylla Mull. Arg. in Mart. Fl, Bras. XI 2 Pax in Das Pflanzenreich IV. (1874) p. 504; Engler, 147. Ill

(1911) p. 42, no.

A small tree. Young branches lenticellate, clothed with more less or patent hairs, glabrescent. Stipules 8 —10 mm long, lanceolate,

—6 slightly hairy outside, deciduous. Petioles 2 cm long, rather clothed with hairs in the thick, sparsely especially upper part, the articulate but glabrescent, longitudinally striate, at apex not

conspicuously rugulose. Limb lanceolate, shortly acuminate at the the with apex, long cuneate to base, 2 small, 1— i.j mm long basal glands, chartaceous, margin glanduliferous serrate-dentate, sparsely

clothed with dibrachiate hairs beneath, nerves prominent at both side- sides, especially beneath, 32—80 cm long, 10—24.5 cm broad; Inflorescences in the nerves 25—38. 3, spicate, often branched lower flowers three-flowered part, in dichasia in the axils of c

2 mm long, triangular bracts, which are sparsely hairy and glab-

rescent rhachis clothed with less outside, densely more or patent

hairs. c. lobes Calyx 4 mm long, unequally 5-lobed, 2 small, and and triangular subacute, 3 larger, oblong or ovate and obtuse Petals or emarginate, sparsely hairy, soon glabrescent. c. 8 mm

long, connate in the lower part, just below the middle, lobes outside oblong, obtuse, glabrous, inside barbate at the throat.

Stamens 6, about as long as the petals, introrse, connective rather broad. Disk lobes deeply j-lobed, subovate, c. 1 mm long. In- florescences bracts denser clothed 9, 7 —11 cm long, not branched, with hairs. Flowers in the subpatent specimens seen very young, and disk calyx petals j-lobed, annular, ovary 3-celled.

Distribution : Southern Brazil, Rio de Janeiro.

Rio dc Mandioca fl. Janeiro near (Riedel s.n.! [G. P.], 3 Type of the Serra 9 species!); dos Orgaos (Glaziou 7434! [P.], young fl.; Glaziou

17211a! [P.], 3 fl.); Jacaripago (Glaziou 2524! [B.K.P.], young 9 fl.); without Brazil, precise locality (Burohell 257A! [K.], young 3 fl.).

This species is closely related to P. Trianae (Mull. Arg.) Baill.

from which it differs in basal having 2 glands only, in the teeth the along margin which are not curved, in the lobed disk and in the shorter inflorescences. From P. macrostachya Ducke it is 763

of the leaves of P. Trianae Baill. and P. Fig. 1. Margin a. b. megalophylla Müll. Arg.

only to distinguish by the small leaf-glands and the short inflores- cences with only 3-flowered glomerules.

Pausandra Trianae in Adansonia XI 3. (Mull. Ang.) Baill. Pax in Das Pflanzenreich IV. Ill (1873) p. 92; Engler, 147. (1911)

— Mull. p. 42; Pogonophora Trianae Arg. in Flora XLVII (1864)

Pax l.c. — Pax p. 434; p. 109; Pausandra quadriglandulosa et K. Hoffm. in Das Pflanzenreich IV. XIV Engler, 147. (1919)

— P. extorris in Pub!. Field Mus. Bot. Ser. IV P- 43; Standley

(1929) p. 219. branches A small tree, j—15 m high (Ule). Young lenticellate, clothed with Petioles densely appressed hairs, glabrescent. i.j—- thickened and 5.5 mm long, longitudinally striate, rugulose at the Limb apex. 30—60 cm long, 8.5 —ij.j cm broad, oblanceolate, acuminate the shortly and rounded at apex, long cuneate to the base, usually 4-glandular at the base, rarely 2-glandular, coriaceous, subcoriaceous or chartaceous, margin serrate-dentate, with upwards curved c. 1 mm broad glanduliferous teeth, brownish above, paler beneath, densely clothed with yellowish brown hairs on both surfaces when both young, soon glabrescent; nerves prominent at the side-nerves Inflorescences sides, especially midrib, 25—30.

S 10—20 cm long, in the lower part sometimes with c. 764

2 cm long side-branches, with small few-flowered glome- rules of flowers distance from each other at c. i cm along the rhachis; rhachis densely clothed with yellowish brown, appressed

hairs; flowers shortly pedicellate, c. 8 mm long. Calyx shortly j-lobed, outside covered with short hairs, lobes obtusely triangular. Petals of their lobes connate up to 1/3 length, oblong or ovate-

oblong, rounded, outside glabrous, Inside at the throat with a ring of woolly hairs. Disk long cupulate, with a slightly undulate Stamens about the with margin. 5 —7, as long as corolla, long

filiform filaments and small oblong or rarely subtriangular anthers.

From Distribution : Nicaragua to Brazil.

Nicaragua: Region of Braggman’c Bluff (F.C. Englesing 216! [C.], P. extorris young $ fl., Type of Standley); Choutales, alt. 600 m (Levy 484! [K.P.], $ fl. June). Colombia : New Granada, Prov. Begota, plains of Sn. Martin, Meta R., alt. 300 m (Triana 2597! [G.P.K.], young $ fl., Type of Pogonophora Trianae Mull. Arg.). Rio S. Francisco Brazil: , Seringal (Ule 9538! [D.G.L.U.], young

$ fl. April, Type of P. quadriglandulosa Pax et K. Hofftn.).

Pausandra Ducke in Arch. Rio de IV 4. macrostachya Janeiro

(1925) p. 114.

A small tree. Young branches densely hairy, glabrescent, lenti-

cellate. Stipules c. 1 cm long, lanceolate, acuminate, outside clothed with short deciduous. Petioles hairs, c. 7 cm long, sparsely hairy, the glabrescent, at apex transversely rugulose and sulcate above.

Limb — 33 50 cm long, 12 —16 cm broad, oblanceolate, shortly

acuminate the cuneate the at apex, to base, subcoriaceous, margin basal rather glanduliferous serrate-dentate, glands large c. 2 —3 and covered with hairs and mm long i.j mm broad, patent, above clothed with hairs glabrous sparsely beneath; nerves pro-

minent at both sides Inflores- especially beneath, side-nerves c. 23. flowers cence S 20—40 cm long, solitary or geminate, spicate, in of dichasia in the axils of small glomerules c. 1—1.5 mm long, triangular densely hairy bracts; rhachis densely clothed with hairs. lobes appressed Calyx c. 3 mm long, shortly j-lobed, obtuse, densely hairy outside. Corolla j-lobed, lobes short, obtuse, outside inside barbate glabrous, at the throat. Stamens 3—7, slightly longer than the corolla. Disk urceolate, slightly undulate. Female flowers unknown.

Distribution : Brazil (Para). forest Para, primitive near Francez, Tapajoz R. (Ducke, H.J.B.R. 17897! [D.U.], S fl. Oct., Type).

This species is closely related to P. megalophylla Mull. Arg. 765

Pausandra densiflora Fig. 2. Lanj. 766

From and to P. Morisiana Radik. the latter it can be distinguished by the larger leaves, the number of side-nerves, the extremely long inflorescences and the long-connate petals.

Perhaps belongs to this species also a specimen collected by San Gabriel with Spruce near (Herb. Kew) ripe c. 13 mm high fruits. The leaves of this specimen and especially the basal

glands, the texture of the leaves and the number of side-nerves

are similar to D u c k e’s very specimen.

Pausandra densiflora j. Lanj. nov. spec. fig. 2.

Ab omnibus speciebus inflorescentiis perlongis et densifloris distinguitur.

Arbor c. 10 m alta. Ramuli in statu sicco longitudinaliter sulcati,

juveniles cum stipulis petiolisque pilis brevibus, rigidis, dibrachiatis, demum adpressis vestiti, glabrati. Stipulae lanceolatae, acutae, c. deciduae. Petiolus sulcatus. Limbus 8 mm longae, 2 —3 cm longus,

33—38 cm longus, 11 —13 cm latus, spathulato-oblanceolatus, basin versus longe cuneato-angustatus, apice acute acuminatus, acumine fere membranaceus, serrato- 5 mm longo, rigide margine

ima basi et dentato, glandulis binis, c. 1.5 mm longis 1 mm latis, fere subtus praeditus, supra glaber, praesertim secus nervos pilis nervis brevibus, dibrachiatis, adpressis sparse vestitus, utrinque, secundariae Flores praesertim subtus, prominentibus; costae 26—29. inflorescentiis rhachis dibrachiatis $ in c. 40 —55 cm longis; pilis dense vestita. Flores in glomerulis densifloris in axillis bractearum

extus brevibus intus late triangularium, sparse pilis vestitarum,

glabrarum, in dichasiis sessiles. Sepala 5 extus sparse vestita, intus Petala intus ad faucem barbata. Discus floribus glabra. 5 in mihi nondum evolutus. Stamina juvenilibus qui tantum suppetunt plerumque 6, turn unum centrale. Flores $ ignoti.

Brasilia: Territ. ostium fluminis in firma Acre, pr. Macauban terra

fl. S in Herb. U.; Duplicates arc {Krukoff 5464! Aug. 13, 1933, Typus!

distributed by the New York Botanical Garden to several Herbaria).

Pausandra Martinii Baill. in Adanlsonia XI 6. (1873) p. 92;

Pax in Das Pflanzenreich IV. Ill — Engler, 147. (1911) p. 43; of P. flagellorhachis Lanj., The Euphorbiac. Surinam (1931) p. 30, Flora of fig. 6; Lanjouw in Pulle, Suriname II (1932) p. 71.

A small 8 red bran- tree, —10 m high, with milky juice; young ches clothed with short rigid dibrachiate hairs, glabrescent. Stipules broad the acumi- triangular-lanceolate, j mm long, 2 mm at base, with and hairs nate, glabrous above, covered short, rigid appressed beneath. Petioles 1 —6 cm long, obscurely longitudinally striate, sulcate Limb above, sparsely hairy, glabrescent. (8.5) 12—35 cm 767

long, 3-j —ii.y cm broad, oblong-oblanceolate, cuneate-angustate sometimes and subrounded the acuminate or angustate to base, at the apex, chartaceous, glanduliferous serrate-dentate, biglandular

at the base, dark-brown above, paler beneath, nerves prominent both side-nerves at sides; (12) 17—22, tertiary nerves parallel and on the side-nerves. Inflorescences S 10— subperpendicular , with few-flowered and 30 cm long, flagelliform spikes, glomerules rhachis flowers su'bsessile. 1 —4 cm long internodes; sparsely hairy; Calyx thick, unequally y-lobed, outside clothed with rigid hairs; lobes rotundate. Petals triangular, 5, thick, broadly ovate, obtuse, outside sparsely covered with small rigid hairs. Disk shortly

urceolate. Stamens 6, with short filaments. Flowers 2 only seen fruit Pedicels about in an early stage. thick, short, 2 mm long. outside Sepals y, broadly triangular, acute, sparsely pilose. Ovary about 3-celled, 7 mm long, 5 mm broad, oblong, sparsely pilose. Disk annular, small. Seeds smooth caruncled.

Distribution : Suriname and French Guiana.

French Guiana: Cayenne (Martin s.n.! [P.], Type of P. Martinii Bail!.). Suriname: Brownsberg (B.W. 6725! [U.K.R.], $ fl. July, Type of P. flagellorhachis Lanj.; B.W. 724! [U.], S fl. Sept.; B.W. 3290! [U.L.J, B.W. young fr. and $ fl. Sept.; 6484! [U.P.], $ fl. June). Brazil: Velho Para, Juruty (Duoke H.J.B.R. 20790! [D.U.], young

S fl. and 2 fl. without corolla. Leaves sometimes 4-glandular at the base.

This specimen I have inserted in this species though the determination is

not quite certain. It seems to be a transition to P. Trianae.).

Pausandra sericea 7. Lanj. nov. spec. P. macropetalae Ducke valde affinis, differt foliis longioribus, disco urceolato. sparse pilosis, Arbor Ramuli in sicco parva. statu longitudinaliter sulcati, Folia —8 glabri. alterna, petiolata; petiolus 4.5 cm longus, longi- leviter tudinaliter striatus, in parte superiore canaliculatus, apice incrassulatus, sericeis, dibrachiatis vestitus, pilis sparse glabrescens, limbus 33—yo cm longus, 8.y —10.y cm latus, spathulato-oblan- acumine fere ceolatus, basi cuneatus, apice acute acuminatus, membranaceus, 2 —3 cm longo, rigide margine serrato-dentato, parte inferiore Integro, ima basi glandulis binis c. 1 mm longis praedita, subtus supra glaber, praesertim secus nervos pilis adpressis sparse vestitus, nervis utrinque praesertim subtus prominentibus; costae secundariae femineae in axillis foliorum c. 30. Spicae terminales,

12—18 cm longae, rhachi sericea, pilis non arete adpressis, dibra- in chiatis; floribus solitariis axillis bractearum parvarum, glandulis

binis e basi rotundata breviter cilindricis, apice leviter impressis 768

remotis. Flores praeditarum, j—10 mm c. 1 mm pedicellati;

late et obtuse pedicellus cum calyce sericeus; sepala j, triangularia vel intus ovata, imbricata, glabra, 1 —i.j mm longa; petala j, vel libera, fere j mm longa, oblonga obovato-oblonga, extus glabra, intus circa medium barbata; discus brevis, cupulatus, obscure ovarium dense crassiusculis. 5-lobatus; sericeum, stylis 3 trigonis, apice breviter bilobatis. Fructus et flores $ ignoti.

Brasilia; Matto fluv. Machado prov. Grosso, prope Tabajara, sup. (Krukoff fl. Nov. Dec. in Herb. distributed 1337! 1931, Typus Utrecht; duplicates the York Card, several by New Bot. to Herbaria). Upper Rio Negro (Ducke

H.J.B.R. 23326! [D.], young $ fl. This specimen probably belongs to this Inflorescences and species. up to 45 cm long with long internodes few-flowe- red and corolla glomerules; calyx 5-merous, stamens c. 7).

8. Pausandra macropetala Ducke in Arch. Rio de Janeiro IV

(1925) p. 114.

A small tree, monoecious. Young branches and petioles sparsely clothed with dibrachiate appressed hairs, glabrescent. Stipules c.

outside. Petioles — obs- 4 mm long, hairy very short 1 i.j cm long, curely longitudinally striate, slightly thickened and subrugulose the Limb at apex. 17 —30 cm long, 6—9 cm broad, oblanceolate, acuminate the the chartaceous shortly at apex, long cuneate to base, both to subcoriaceous, glaucous green, paler beneath, glabrous at

sides, margin serrate-dentate, nerves prominent at both sides, side- Inflorescences with few- nerves c. 20. $ c. 12—ij cm long, flowered the axils of small glomerules in c. 1 mm long, triangular, rhachis clothed with hairy, conspicuously biglandular bracts, ap- hairs. Flowers pressed c. 5 mm long. Calyx c. 3—3.5 mm long.

3-lobed, lobes broadly ovate, rounded or subacute, densely hairy outside. Petals till above the connate middle, 5-lobed, lobes broad,

rotundate, inside barbate at the throat. Stamens 6, glabrous. Disk Inflorescence than the deeply j-lobed. 9 5 cm long, more robust male flowers in the flowers one, solitary, as S or larger, ovary thick, Fruit densely sericeous, styles 3, short, bipartite. c. 13 mm Seed caruncled. high. oblong, 9 mm long, 7 mm broad, marmorate,

Distribution: Brazil (Para).

River Tapajoz near Montanha cataract (Ducke, H.J.B.R. 17896! [D.U.], $ fl. The Type in Herb. Rio de Janeiro bears $ and 9 flowers after the de Olivenca description of Ducke); Rio Solimoes, near Sao Paulo (Ducke, H.J.B.R. 20789! [D.], fr. Nov.).

this have male branch in the Utrecht From species I only seen a

Herbarium and a fruiting specimen from Berlin. The species is

closely related to P. sericea Lanj., from which it differs in size and indument of the leaves and in the nearly free disk glands. 769

Pausandra hirsuta 9. Lanj. nov. spec. A P. Martinii cui valde affinis differt ramulis Bail!., est, costis petiolisque pilis simplicibus hirsutis, calyce glabro, foliis baud cuneatis.

Frutex c. 6 m altus, ramulis longitudinaliter striatis, hirsute

patenti-pilosis, foliis alternis, bistipulatis; stipulae lanceolatae, c.

8 mm longae, acutae, extus in parte inferiore margineque pilis vestitae. Petiolus rigidis i.y —3.5 cm longus, longitudinaliter

striatus, apice non vel vix incrassatus, dense hirsuto-pilosus. Limbus

18 basin (12) —25 cm longus, 4—9.5 cm latus, versus acute basi binis subacutis angustatus, glandulis c. 1.5 mm longis, ornatus, subtus in chartaceus, supra glaber, pallidior, costa hirsuto-pilosus, in nervis secundariis tertiariisque pilis sparsissime vestitus, margine serrato-dentatus, dentibus parvis glanduliferis, nervis venisque

nervis lateralibus — utrinque prominentibus, 15 16. Spicae $ gra- florum ciles, 5 —10 cm longae, glomerulis parvis, paucifloris,

remotis, in axillis bractearum biglandularum, lanceolatarum, acu- basi tarum, et margine pilosarum. Flores (juveniles tantum visi) in bractearum axillis triangularium, acutarum, carinatarum, praeter carinam et marginem glabrarum. Calyx 5-lobatus, praeter piles minutos lobis fimbriatis. raros, glaber, subrotundatis, margine Stamina filamentis brevibus. Discus 6, quorum unum centrale, Flos fructus parvus, cupulatus. $ et ignoti.

Casas Brasilia: Amazonas, Humayata prope Tres (Krukoff 6084! fl. Sept.

Oct. in New York 1934, Typus Herb. Utrecht; duplicates distributed by the

Bot. Garden to several Herbaria).

Excluded species: Pausandra in Bull. integrifolia Lanj. Kew 1932, p. 183. Now that I have examined all the material of the I genus am

convinced that this must be excluded from the As species genus. it differs in several characters from had the the genus I placed It species in a new sect. Pausandrella Lanj. seems highly probable chat second of the it represents a species genus: Ducke described from Brazil. The the some years ago petioles, glands and the texture of the leaves, the inflorescence, the hairs, the of the disk the indument and pollen grains suggest its being member of this D k e’s is and a new genus. u c species 4-merous

has more stamens and also the shape of the leaves is different.

did not female material of and As I see good my species Mr. d who S a n w i t h, collected the type-material, thinks it highly

probable that new specimens can be obtained from British Guiana, it wait I think better to with transferring it definitely to the Ducke till material has been collected. genus Dodecastigma new