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Bonner Zoologische Beiträge ZOBODAT - www.zobodat.at Zoologisch-Botanische Datenbank/Zoological-Botanical Database Digitale Literatur/Digital Literature Zeitschrift/Journal: Bonn zoological Bulletin - früher Bonner Zoologische Beiträge. Jahr/Year: 2005/2006 Band/Volume: 54 Autor(en)/Author(s): Assing Volker Artikel/Article: A Revision of Western Palaearctic Medon: the Species of the Atlantic Islands, the Western Mediterranean, and Europe, Except for the Southeast (Insecta: Coleoptera: Staphylinidae: Paederinae) 25-95 © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.zoologicalbulletin.de; www.biologiezentrum.at Bonner zoologische Beiträge Band 54 (2005) Heft 1 Seiten 25-95 Bonn, April 2006 A Revision of Western Palaearctic Medom the Species of the Atlantic Islands, the Western Mediterranean, and Europe, Except for the Southeast (Insecta: Coleóptera: Staphylinidae: Paederinae) Volker ASSING Hannover, Germany Abstract. Types and additional material oí Medon Stephens are revised. 29 valid species and subspecies, including three doubtful species, are recognized in the region comprising the Western Mediterranean (including Italy and Tunisia), the Atlantic Islands, as well as Western, Central, and Northern Europe. Medon oromii sp. n. (Canary Islands: Gran Ca- naria), M. cmtricola sp. n. (Canary Islands: El Hierro), and M. kabylicus sp. n. (Algeria) are described. Twenty-three synonymies, two of them pertaining to the previously revised species of the Eastern Mediterranean region, are estab- lished: Medon apicalis (Kraatz, 1857) = Medon orgianus Bordoni, 1980 syn. n.; Medon pemiger Coiffait, 1978 = Me- don perniger fraudulentas Coiffait, 1978 syn. n.; Medon dilutus (Erichson, 1839) = M. dilutus meridional is Bordoni, 1980 syn. n.; M. dilutus cephalus Koch, 1938 = M. dilutus boeticus Jeannel & Jarrige, 1949 syn. n.; Medon dilutus quadriceps (Wollaston. 1864) = Medon vitalei Bernhauer. 1936 syn. n. = M. dilutus eremicus Koch, 1939 syn. n. = M. marocanus Coiffait, 1970 syn. n. = M. mateui Coiffait, 1973 syn. n.; Medon ruflventris (Nordmann, 1837) = M. burdi- galensis Coiffait, 1970 syn. n. = M. aquitanicus Coiffait, 1970 syn. n. = M. siculus Coiffait, 1970 syn. n. = M. sicilianas Coiffait, 1970 syn. n.; Medon africanas (Fauvel. 1872) = M. bodemeyeri Bernhauer, 1915 syn. n.; Medon piceus (Kraatz, 1858) = M. murciensis Coiffait, 1970 syn. n.; Medon cauchoisi Jarrige, 1949 = M. oculatior Peyerimhoff, 1949 syn. n. = M. lusitanicus Coiffait, 1970 syn. n. = M. pan'iphallus Coiffait, 1973 syn. n. = M mazices Coiffait. 1973 syn. n. = M. fongondi Coiffait, 1980 syn. n. = M. perraulti Coiffait, 1980 syn. n.; Medon mirei Coiffait, 1980 = M. giachinoi Bordoni. 1988 syn. n.; Medon semiobscurus (Fauvel, 1875) = M. ruber Sahlberg, 1908 syn. n. = M. sahlbergi Scheer- peltz, 1933 syn. n. Medon procerus (Perez Areas, 1874) and M. sericellus (Fainnaire, 1860), previously regarded as synonyms of M. castaneus (Gravenhorst. 1802) and M. apicalis, respectively, are revalidated. A neotype is designated for Lithocharis sericella Fairmaire, 1860. Lectotypes are designated for Lithocharis oppidana Kraatz, 1857, L. picea Kraatz, 1858, Paeclerus castaneus Gravenhorst, 1802, Lithocharis africanas Fauvel, 1872, Medon bodemeyeri Bern- hauer, 1915, M. bucharicus Bernhauer, 1902, and M. ruber Sahlberg, 1908. Medon pythonissa (Saulcy, 1864) is re- garded as a subspecies of M. dilutus. All the species not treated in the first part of the revision are described and illus- trated. Bionomic and Zoogeographie data are compiled, and the distributions of most species are mapped. A key to the Medon species of the Western Mediterranean region and the Atlantic Islands is provided. Medon bucharicus Bernhauer, 1902 (Middle Asia) is redescribed and moved to Medon again (from Sunius Curtis). Some additional records of species from other regions are presented. The previously published revision of Eastern Mediterranean Medon contains several errors, which are corrected in a separate section. These corrigenda include the following synonymies: M. lydicus Bor- doni, 1980 = M. mimulus Fagel, 1970, syn. n. = M. rhodicus Franz, 1987, syn. n. = M. choparti Coiffait, 1987, syn. n. Key words. Taxonomy, revision, new species, new synonyms, lectotype designations, neotype designation 1. INTRODUCTION The species had never been thoroughly revised and nu- merous species have been described especially from In the recently published first part of the revision of France, the Iberian Peninsula, and from Northwest Af- Western Palaearctic Medon and a subsequent supple- rica. Coiffait (1984) lists 36 species and subspecies for ment (ASSING 2004a, 2004b), the species of the Eastern the Western Mediterranean (including Italy, the Atlantic Mediterranean and adjacent regions were treated. This Islands, and Northwest Africa eastwards to Tunisia); project had been initiated especially by the difficulties three additional species were later described by Serrano encountered when trying to identify specimens from this (1993), BORDONI (1988), and ASSING (1998). However, area; attributing names to them was often little more a preliminary examination of abundant material, which than a good guess. This observation gave rise to the had been accumulated during various field trips in the suspicion that many names were synonymous and/or past decades, suggested a significantly lower species di- misinterpreted, and, as was expected, one of the main versity in this region. Many Medon species are inhabi- results of this study was a significant reduction of the tants of the leaf litter and occur at relatively high densi- number of valid names. ties, so that it seemed rather unlikely that a considerable For several reasons, the situation was hypothesized to number of species had just not been found. Moreover, be similar for the species of the Western Mediterranean. the revision of the Eastern Mediterranean representa- © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.zoologicalbulletin.de; www.biologiezentrum.at 26 Bonner zoologische Beiträge 54 (2005) tives of the genus had shown that the extent of the dis- could not be put down to extremely restricted distribu- tributions and the range of intraspecific variation had tions. These considerations, as well as the difficulty en- generally been underestimated by previous authors. countered when trying to identify Medon material from Also, some species, like Medon burdigalensis Coiffait the study area suggested that the previous taxonomic and M aquitanicus Coiffait, had never been recorded concept of the Medon fauna of the Western Mediterra- again after their description, although they had been de- nean was at least in part inadequate and eventually initi- scribed from regions where the presence of local en- ated the present revision. demics is unlikely; consequently, the absence of records > a W\ Map 1: The study region. 1 1 1 © Biodiversity Heritage Library, http://www.biodiversitylibrary.org/; www.zoologicalbulletin.de; www.biologiezentrum.at Volker AssiNG: Western Palaearctic Medon 27 The revision focuses on the species of the Atlantic is- CVJ 1 priVd.Lc COI IL C II UM Kj , vJ 1 1 1 C I 1 0 1 S , V di UCi LJ lands and the western parts of the Mediterranean east- firntl nnviifp r*ol it^ci inn A ("innt^rpnk'n OHcsi'sM ^ VJ \J 1 IJ1 1 V CI \.\, CA M 1 V, V. 1 1 \-t 1 1 t \ . VJ VJ1 1 ICl 1 C 1 I I\ VJ, WU^OOCl wards to Tunisia and Italy. However, since the ranges of several widespread species also extend into Western, cRap LMlValtT CUllv.LLli.MI r\ . rvaUU, I\ cl 1 1 1\ W L 1 Central, Northern, and even Eastern Europe, these re- Lrk'nhrvui l UIlVclLC LUIICLL1UM r. IVUI11L1, DU1I1I1C1II1 gions are included, too (Map 1 ). nri viitp 1 Ipcti í í z\ I n ci i iiíi cOro L/llVLllv- vUllvven 1 1 VJon1 1 1P . VOrom7 1 Willi, 1 < CI L-iCl ii LI 1 1 CI The recently published revision of Eastern Mediterra- nean Medon is based on an old manuscript version and cOwe private collection J. A. Owen, Epsom contains some nomenclatural errors. I use the opportu- cRou private collection G. de Rougemont, Londinieres nity to correct them in a separate section at the end of this paper. cSch private collection M. Schülke, Berlin cTro private collection M. Tronquet, Molitg-les-Bains 2. MATERIAL AND MEASUREMENTS cWun private collection P. Wunderle, Mönchen- Types and additional material from the following insti- gladbach tutions and private collections were examined: cZan private collection A. Zanetti, Verona DEI Deutsches Entomologisches Institut, Eberswalde (L. Zerche) Head length was measured from the anterior margin of the clypeus to the posterior margin of the head, elytral DZUL Departamento de Biología Animal - Zoología, length along suture from the apex of the scutellum to the Universidad de La Laguna (P. Oromí) elytral hind margin. IRSNB Institut Royal des Sciences Naturelles de Bel- gique, Bruxelles (D. Drugmand) 3. THE MEDON FAUNA OF THE WESTERN FMNH Field Museum of Natural History, Chicago MEDITERRANEAN (via L. H. Herman) Based on an examination of 5600 specimens of Medon, MCNT Museo de Ciencias Naturales de Tenerife including the accessible types, 26 valid species and sub- species are recognized, among them two revalidated and MHNG Muséum d'Histoire Naturelle, Geneve (G. three previously undescribed species. The identities Cuccodoro) (and validity) of three additional species, M. augur Fau- vel, M. subterraneus Coiffait, and M. spelaeus (Scriba), MNHNP Muséum National d'Histoire Naturelle, Paris remain doubtful; augur is represented only a fe- (N. Berti) M. by male type
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