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Phylogenetic Analysis of the Olenellina Walcott, 1890 (Trilobita, Cambrian) Bruce S
2 j&o I J. Paleont., 75(1), 2001, pp. 96-115 Copyright © 2001, The Paleontological Society 0022-3360/01 /0075-96$03.00 PHYLOGENETIC ANALYSIS OF THE OLENELLINA WALCOTT, 1890 (TRILOBITA, CAMBRIAN) BRUCE S. LIEBERMAN Departments of Geology and Ecology and Evolutionary Biology, University of Kansas, Lindley Hall, Lawrence 66045, <[email protected]> ABSTRACT—Phylogenetic analysis was used to evaluate evolutionary relationships within the Cambrian suborder Olenellina Walcott, 1890; special emphasis was placed on those taxa outside of the Olenelloidea. Fifty-seven exoskeletal characters were coded for 24 taxa within the Olenellina and two outgroups referable to the "fallotaspidoid" grade. The Olenelloidea, along with the genus Gabriellus Fritz, 1992, are the sister group of the Judomioidea Repina, 1979. The "Nevadioidea" Hupe, 1953 are a paraphyletic grade group. Four new genera are recognized, Plesionevadia, Cambroinyoella, Callavalonia, and Sdzuyomia, and three new species are described, Nevadia fritzi, Cirquella nelsoni, and Cambroinyoella wallacei. Phylogenetic parsimony analysis is also used to make predictions about the ancestral morphology of the Olenellina. This morphology most resembles the morphology found in Plesionevadia and Pseudoju- domia Egorova in Goryanskii and Egorova, 1964. INTRODUCTION group including the "fallotaspidoids" plus the Redlichiina, and HE ANALYSIS of evolutionary patterns during the Early Cam- potentially all other trilobites. Where the Agnostida fit within this T brian has relevance to paleontologists and evolutionary bi- evolutionary topology depends on whether or not one accepts the ologists for several reasons. Chief among these are expanding our arguments of either Fortey and Whittington (1989), Fortey (1990), knowledge of evolutionary mechanisms and topologies. Regard- and Fortey and Theron (1994) or Ramskold and Edgecombe ing evolutionary mechanisms, because the Cambrian radiation (1991). -
New Information on Olenelline Trilobites from the Early Cambrian Sekwi Formation, Northwestern Canada
KU ScholarWorks | http://kuscholarworks.ku.edu Please share your stories about how Open Access to this article benefits you. New information on olenelline trilobites from the Early Cambrian Sekwi Formation, Northwestern Canada by Bruce S. Lieberman 2010 This is the author’s accepted manuscript version of the article, made available with the permission of the publisher. The original published version can be found at the link below. Lieberman, Bruce S. 2010. “New information on olenelline trilobites from the Early Cambrian Sekwi Formation, northwestern Canada.” Canadian Journal of Earth Sciences (47):1445-1449. Published version: http://www.nrcresearchpress.com/doi/ abs/10.1139/E10-073#.U21JO4FdXUL Terms of Use: http://www2.ku.edu/~scholar/docs/license.shtml KU ScholarWorks is a service provided by the KU Libraries’ Office of Scholarly Communication & Copyright. For Review Purposes Only/Aux fins d'examen seulement 1 1 New information on olenelline trilobites from the Early Cambrian Sekwi Formation, 2 Northwestern Canada 3 4 Francine R. Abe 5 Department of Ecology and Evolutionary Biology, University of Kansas, 1345 Jayhawk 6 Boulevard, Lawrence, Kansas 66045, USA, Tel: +1 (785) 864-3369, Fax: +1 (785) 864-5335 7 <[email protected]> 8 Bruce S. Lieberman 9 Department of Geology and Natural History Museum and Biodiversity Research Center, 10 University of Kansas, 1475 Jayhawk Boulevard, 120 Lindley Hall, Lawrence, Kansas 66045, 11 USA, <[email protected]> 12 Michael C. Pope 13 Department of Geology and Geophysics, Texas A&M University, College Station, Texas 77843, 14 USA, <[email protected]> 15 Kelly Dilliard 16 Department of Physical Sciences and Mathematics, Wayne State College, 1111 Main Street, 17 Wayne, Nebraska 68787, USA, <[email protected]> 18 For Review Purposes Only/Aux fins d'examen seulement 2 19 Abstract 20 A new species of olenelline trilobite, Nevadella keelensis, is described from the lower Cambrian Sekwi 21 Formation, Mackenzie Mountains, Canada. -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
Cambrian Transition in the Southern Great Basin
The Sedimentary Record 2000; Shen and Schidlowski, 2000). Due to The Precambrian- endemic biotas and facies control, it is diffi- cult to correlate directly between siliciclas- Cambrian Transition in the tic- and carbonate-dominated successions. This is particularly true for the PC-C boundary interval because lowermost Southern Great Basin, USA Cambrian biotas are highly endemic and Frank A. Corsetti James W.Hagadorn individual, globally distributed guide fossils Department of Earth Science Department of Geology are lacking (Landing, 1988; Geyer and University of Southern California Amherst College Shergold, 2000). Los Angeles, CA 90089-0740 Amherst, MA 01002 Determination of a stratigraphic bound- [email protected] [email protected] ary generates a large amount of interest because it provides scientists with an oppor- ABSTRACT:The Precambrian-Cambrian boundary presents an interesting tunity to address a variety of related issues, stratigraphic conundrum: the trace fossil used to mark and correlate the base of the including whether the proposed boundary Cambrian, Treptichnus pedum, is restricted to siliciclastic facies, whereas position marks a major event in Earth histo- biomineralized fossils and chemostratigraphic signals are most commonly obtained ry. Sometimes the larger-scale meaning of from carbonate-dominated sections.Thus, it is difficult to correlate directly between the particular boundary can be lost during many of the Precambrian-Cambrian boundary sections, and to assess details of the the process of characterization. This is timing of evolutionary events that transpired during this interval of time.Thick demonstrated in a plot of PC-C boundary sections in the White-Inyo region of eastern California and western Nevada, USA, papers through time (Fig. -
Arthropod Pattern Theory and Cambrian Trilobites
Bijdragen tot de Dierkunde, 64 (4) 193-213 (1995) SPB Academie Publishing bv, The Hague Arthropod pattern theory and Cambrian trilobites Frederick A. Sundberg Research Associate, Invertebrate Paleontology Section, Los Angeles County Museum of Natural History, 900 Exposition Boulevard, Los Angeles, California 90007, USA Keywords: Arthropod pattern theory, Cambrian, trilobites, segment distributions 4 Abstract ou 6). La limite thorax/pygidium se trouve généralementau niveau du node 2 (duplomères 11—13) et du node 3 (duplomères les les 18—20) pour Corynexochides et respectivement pour Pty- An analysis of duplomere (= segment) distribution within the chopariides.Cette limite se trouve dans le champ 4 (duplomères cephalon,thorax, and pygidium of Cambrian trilobites was un- 21—n) dans le cas des Olenellides et des Redlichiides. L’extrémité dertaken to determine if the Arthropod Pattern Theory (APT) du corps se trouve généralementau niveau du node 3 chez les proposed by Schram & Emerson (1991) applies to Cambrian Corynexochides, et au niveau du champ 4 chez les Olenellides, trilobites. The boundary of the cephalon/thorax occurs within les Redlichiides et les Ptychopariides. D’autre part, les épines 1 4 the predicted duplomerenode (duplomeres or 6). The bound- macropleurales, qui pourraient indiquer l’emplacement des ary between the thorax and pygidium generally occurs within gonopores ou de l’anus, sont généralementsituées au niveau des node 2 (duplomeres 11—13) and node 3 (duplomeres 18—20) for duplomères pronostiqués. La limite prothorax/opisthothorax corynexochids and ptychopariids, respectively. This boundary des Olenellides est située dans le node 3 ou près de celui-ci. Ces occurs within field 4 (duplomeres21—n) for olenellids and red- résultats indiquent que nombre et distribution des duplomères lichiids. -
New Olenelloid Trilobites from the Northwest Territories, Canada
Zootaxa 3866 (4): 479–498 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3866.4.2 http://zoobank.org/urn:lsid:zoobank.org:pub:D06E5477-4D5C-4402-B909-09A2AAFFB556 New olenelloid trilobites from the Northwest Territories, Canada I. WESLEY GAPP1,2 & BRUCE S. LIEBERMAN2 1Chevron U. S. A., Inc., 1400 Smith Street, Houston, TX 77002, USA. E-mail: [email protected] 2Department of Ecology & Evolutionary Biology and Biodiversity Institute, University of Kansas, 1345 Jayhawk Blvd, Dyche Hall, Lawrence, KS 66045, USA. E-mail: [email protected] Abstract The Olenelloidea are a superfamily of early Cambrian trilobites, which have been the subject of several phylogenetic anal- yses and also used to address macroevolutionary questions regarding the nature and timing of the Cambrian radiation. The Sekwi Formation of the Mackenzie Mountains, Northwest Territories, Canada, has yielded numerous species from this clade, and here we present new information that expands on the diversity known from this biogeographically and biostrati- graphically important region. In particular, we describe seven new species, (Olenellus baileyi, Mesonacis wileyi, Ellipto- cephala jaredi, Holmiella taurus, H. domackae, Mummaspis delgadoae, and Bristolia colberti). Also recovered are specimens of Elliptocephala logani, specimens that shared affinities with Olenellus clarki, O. getzi, O. fowleri, and Fri- zolenellus hanseni, and one partial specimen, which appears to be a new species of Bolbolenellus. Key words: Cambrian, Trilobita, Olenelloidea, Northwest Territories Introduction The Olenelloidea Walcott, 1890 is a diverse superfamily of early Cambrian trilobites referable to the suborder Olenellina Walcott, 1890 and have been the focus of much attention in the study of evolutionary tempo and mode during the Cambrian radiation (Fortey et al. -
New Palaeoscolecidan Worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale
New palaeoscolecidan worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale SIMON CONWAY MORRIS and JOHN S. PEEL Conway Morris, S. and Peel, J.S. 2010. New palaeoscolecidan worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale. Acta Palaeontologica Polonica 55 (1): 141–156. Palaeoscolecidan worms are an important component of many Lower Palaeozoic marine assemblages, with notable oc− currences in a number of Burgess Shale−type Fossil−Lagerstätten. In addition to material from the lower Cambrian Kinzers Formation and Latham Shale, we also describe two new palaeoscolecidan taxa from the lower Cambrian Sirius Passet Fossil−Lagerstätte of North Greenland: Chalazoscolex pharkus gen. et sp. nov and Xystoscolex boreogyrus gen. et sp. nov. These palaeoscolecidans appear to be the oldest known (Cambrian Series 2, Stage 3) soft−bodied examples, being somewhat older than the diverse assemblages from the Chengjiang Fossil−Lagerstätte of China. In the Sirius Passet taxa the body is composed of a spinose introvert (or proboscis), trunk with ornamentation that includes regions bearing cuticu− lar ridges and sclerites, and a caudal zone with prominent circles of sclerites. The taxa are evidently quite closely related; generic differentiation is based on degree of trunk ornamentation, details of introvert structure and nature of the caudal re− gion. The worms were probably infaunal or semi−epifaunal; gut contents suggest that at least X. boreogyrus may have preyed on the arthropod Isoxys. Comparison with other palaeoscolecidans is relatively straightforward in terms of compa− rable examples in other Burgess Shale−type occurrences, but is much more tenuous with respect to the important record of isolated sclerites. -
New Olenelline Trilobites from the Northwest Territories, Canada, and the Phylogenetic Placement of Judomia Absita
KU ScholarWorks | http://kuscholarworks.ku.edu Please share your stories about how Open Access to this article benefits you. New olenelline trilobites from the Northwest Territories, Canada, and the phylogenetic placement of Judomia absita by Bruce S. Lieberman 2011 This is the author’s accepted manuscript version of the article, made available with the permission of the publisher. The original published version can be found at the link below. Lieberman, Bruce S. 2011. “New olenelline trilobites from the Northwest Territories, Canada, and the phylogenetic placement of Judomia absita.” Zootaxa (2918):15-28 Published version: http://www.mapress.com/zootaxa/2011/2/ zt02918p028.pdf Terms of Use: http://www2.ku.edu/~scholar/docs/license.shtml KU ScholarWorks is a service provided by the KU Libraries’ Office of Scholarly Communication & Copyright. New olenelline trilobites from the Northwest Territories, Canada, and the phylogenetic placement of Judomia absita I. WESLEY GAPP1, BRUCE S. LIEBERMAN1, MICHAEL C. POPE2 & KELLY A. DILLIARD3 1Department of Geology and Biodiversity Institute, University of Kansas, 1475 Jayhawk Blvd, 120 Lindley Hall, Lawrence 66045 Email:[email protected] 2Department of Geology and Geophysics, Texas A&M University, College Station, TX 77843 Email:[email protected] 3Department of Physical Sciences and Mathematics, Wayne State College, 1111 Main Street, Wayne, NE 68787 Email:[email protected] Key words: Trilobita, Olenellina, Cambrian radiation, phylogenetics, paleobiogeography Abstract The Early Cambrian olenelline trilobites are a diverse clade that have been the subject of several phylogenetic analyses. Here, three new species of Bradyfallotaspis Fritz, 1972 (B. coriae, B. nicolascagei, and B. sekwiensis) and one new species of Nevadia Walcott, 1910 (N. -
New Species of Olenellina Trilobites from the Early
THE NATURE OF EVOLUTIONARY RADIATIONS WITH A SPECIAL FOCUS ON DEVONIAN CALMONIID TRILOBITES by Francine R. Abe B.Sc., Bates College, 2003 Submitted to the Department of Ecology and Evolutionary Biology and the Faculty of the Graduate School of the University of Kansas in partial fulfillment of the requirements for the degree of Doctor of Philosophy Bruce S. Lieberman, Co-Chair Edward O. Wiley, Co-Chair Paul Selden Maria Orive Linda Trueb th Date Defended: June 10 , 2010 The Dissertation Committee for Francine R. Abe certifies that this is the approved version of the following dissertation: THE NATURE OF EVOLUTIONARY RADIATIONS WITH A SPECIAL FOCUS ON DEVONIAN CALMONIID TRILOBITES Committee: Bruce S. Lieberman, Co-Chair Edward O. Wiley, Co-Chair Paul Selden Maria Orive Linda Trueb Date Defended: June 10th, 2010 ii ABSTRACT Trilobites were a diverse group of arthropods that left an extensive fossil record which are today used today to study macroevolutionary patterns and processes. A new species and form of olenelloidea trilobites (Nevadella keelensis n. sp. and Esmeraldina rowei) are described from the lower Cambrian Sekwi Formation, Canada. One particular clade of trilobites, the Devonian calmoniid trilobites of the Malvinokaffric Realm, was hailed as a classic paleontological example of an adaptive radiation. Three aspects of the radiation were examined in detail: diversification rates, biogeography, and morphological change. The rates of speciation calculated from stratigraphy were highest when sea level was lowest suggesting causal effect of abiotic factors in speciation. A phylogenetic biogeographic analysis indicated a geographically complex area, and this geographic complexity created various opportunities for speciation via allopatry through sea level changes. -
Coleophora Nepetellae Baldizzone & Nel, a New Species of the C. Lixella Group
A peer-reviewed open-access journal ZooKeys 459: Coleophora119–135 (2014) nepetellae Baldizzone & Nel, a new species of the C. lixella group... 119 doi: 10.3897/zookeys.459.7983 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Coleophora nepetellae Baldizzone & Nel, a new species of the C. lixella group (Lepidoptera, Coleophoridae) from France and Italy Giorgio Baldizzone1,*, Jacques Nel2, Jean-François Landry3 1 Via Manzoni, 24, I-14100 Asti, Italy 2 78, Avenue Fernand Gassion, F-13600 La Ciotat, France 3 Agriculture and Agri-Food Canada, C.E.F., 960 Carling Avenue, Ottawa, Ontario K1A 0C6, Canada Corresponding author: Jean-François Landry ([email protected]) Academic editor: E.van Nieukerken | Received 30 May 2014 | Accepted 24 October 2014 | Published 2 December 2014 http://zoobank.org/993DCA9D-8A21-444C-8D5D-487756B7B07A Citation: Baldizzone G, Nel J, Landry J-F (2014) Coleophora nepetellae Baldizzone & Nel, a new species of the C. lixella group (Lepidoptera, Coleophoridae) from France and Italy. ZooKeys 459: 119–135. doi: 10.3897/zookeys.459.7983 Abstract Coleophora nepetellae Baldizzone & Nel, sp. n. is described from the southern Alps (Italy and France). It belongs to the Coleophora lixella species group. Its host plants are Nepeta nepetella L. (Lamiaceae) and an unidentified Poaceae. The fifth instar larva, its case, the adult habitus, and genitalia are illustrated. The species is compared to C. nevadella Baldizzone, 1985, here newly confirmed from France and whose larvae feed on Nepeta latifolia DC. in the Eastern Pyrénées. DNA barcodes are shown to be distinct and congruent with morphological differences among species of the lixella group. -
Brigham Young University Geology Studies
GEOLOGY YOUNG STUDIES UNIVERSITY CONTENTS Paleontology and Depositional Environments: Cambrian of Western North America A symposium U. J. Brady University of Kansas D. P. Campbell University of Kansas H. E. Cook U. S. Geological Survey W. H. Fritz Canadian Geological Survey J. C. Kepper University of Nevada, Las Vegas R. B. Koepnidc Williams College V. E. Kurt2 Southwest Missouri State University K. C. Lohmann State University of New Yo&, Stony Brook D. J. McSride University of Kansas J. N. Moore University of California, Los AngeIes A. R. Pher State University of New York, Stony Brook J. K. aigby Brigham Young University R. A. Robison University of Kansas A. J. Bowell University of Kansas James Sprinkle University of Texas, Austin M. E. Taylor u. s. Geological SULV~~ Extended presentations of some papers presented at a Paleontological Society symposium held in Salt Lake City, Utah, on October 20, 1975. Brigham Young University Geology Studies Volume 23, Part 2-July 1976 Contents Trilobites in Utah folklore ................ M. E. Taylor and R. A. Robison Lower Cambrian Stratigraphy, Mackenzie Mountains, Northwestern Canada .................................................... W. H. Fritz Depositional Environments of the Lower Cambrian Poleta Formation and Its Stratigraphic Equivalents, California and Nevada ............................ J. N. Moore Biostratigraphic Implications of Trilobite Biofacies: Albertella Zone, Middle Cambrian, Western United States ............................ A. R. Palmer and D. P. Campbell Some Observations on Occurrences of Cambrian Porifera in Western North America and Their Evolution .......................................................... J. K. Rigby Biostratigraphy and Paleoecology of Cambrian Echinoderms from the Rocky Mountains ................ James Sprinkle Stratigraphic Relationships and Depositional Facies in a Portion of the Middle Cambrian of the Basin and Range Province ............................... -
In Search of Vanished Ages--Field Trips to Fossil Localities in California, Nevada, and Utah
i In Search Of Vanished Ages Field trips to fossil localities in California, Nevada, and Utah By Inyo A view across the middle Miocene Barstow Formation on California’s Mojave Desert. Here, limestone concretions that occur in rocks deposited in a freshwater lake system approximately 17 million year-ago produce exquisitely preserved, fully three-dimensional insects, spiders, water mites, and fairy shrimp that can be dissolved free of their stone encasings with a diluted acid solution—one of only a handful of localities worldwide where fossil insects can be removed successfully from their matrixes without obliterating the specimens. ii Table of Contents Chapter Page 1—Fossil Plants At Aldrich Hill 1 2—A Visit To Ammonite Canyon, Nevada 6 3—Fossil Insects And Vertebrates On The Mojave Desert, California 15 4—Fossil Plants At Buffalo Canyon, Nevada 45 5--Ordovician Fossils At The Great Beatty Mudmound, Nevada 50 6--Fossil Plants And Insects At Bull Run, Nevada 58 7-- Field Trip To The Copper Basin Fossil Flora, Nevada 65 8--Trilobites In The Nopah Range, Inyo County, California 70 9--Field Trip To A Vertebrate Fossil Locality In The Coso Range, California 76 10--Plant Fossils In The Dead Camel Range, Nevada 83 11-- A Visit To The Early Cambrian Waucoba Spring Geologic Section, California 88 12-- Fossils In Millard County, Utah 95 13--A Visit To Fossil Valley, Great Basin Desert, Nevada 107 14--High Inyo Mountains Fossils, California 119 15--Early Cambrian Fossils In Western Nevada 126 16--Field Trip To The Kettleman Hills Fossil District,