Catalog of Type Specimens of Recent Mammals: Orders Carnivora, Perissodactyla, Artiodactyla, and Cetacea in the National Museum
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Smithsonian Institution Scholarly Press smithsonian contributions to zoology • number 646 Smithsonian Institution Scholarly Press CatalogA Chronology of Type Specimens of Middleof Recent Missouri Mammals: Plains Orders Carnivora, Perissodactyla, Artiodactyla,Village and CetaceaSites in the National Museum of Natural History, Smithsonian Institution By Craig M. Johnson Robert D.with Fisher contributions and Craig A. by Ludwig Stanley A. Ahler, Herbert Haas, and Georges Bonani SERIES PUBLICATIONS OF THE SMITHSONIAN INSTITUTION Emphasis upon publication as a means of “diffusing knowledge” was expressed by the first Secretary of the Smithsonian. In his formal plan for the Institution, Joseph Henry outlined a program that included the following statement: “It is proposed to publish a series of reports, giving an account of the new discoveries in science, and of the changes made from year to year in all branches of knowledge.” This theme of basic research has been adhered to through the years by thousands of titles issued in series publications under the Smithsonian imprint, commencing with Smithsonian Contributions to Knowledge in 1848 and continuing with the following active series: Smithsonian Contributions to Anthropology Smithsonian Contributions to Botany Smithsonian Contributions in History and Technology Smithsonian Contributions to the Marine Sciences Smithsonian Contributions to Museum Conservation Smithsonian Contributions to Paleobiology Smithsonian Contributions to Zoology In these series, the Institution publishes small papers and full-scale monographs that report on the research and collections of its various museums and bureaus. The Smithsonian Contributions Series are distributed via mailing lists to libraries, universities, and similar institutions throughout the world. Manuscripts intended for publication in the Contributions Series undergo substantive peer review and evaluation by SISP’s Editorial Board, as well as evaluation by SISP for compliance with manuscript preparation guidelines (available on SISP’s “Author Resources” page at www.scholarlypress.si.edu). For open access, fully searchable PDFs of volumes in the Smithsonian Contributions Series, visit Open SI (http:// opensi.si.edu). smithsonian contributions to zoology • numb e r 6 4 6 Catalog of Type Specimens of Recent Mammals: Orders Carnivora, Perissodactyla, Artiodactyla, and Cetacea in the National Museum of Natural History, Smithsonian Institution Robert D. Fisher and Craig A. Ludwig WASHINGTON D.C. 2016 ABSTRACT Fisher, Robert D. and Craig A. Ludwig. Catalog of Type Specimens of Recent Mammals: Orders Carnivora, Peris- sodactyla, Artiodactyla, and Cetacea in the National Museum of Natural History, Smithsonian Institution. Smith- sonian Contributions to Zoology, number 646, vi + 112 pages, 2016.—The type collection of Recent mammals in the Division of Mammals, National Museum of Natural History, Smithsonian Institution, contains 612 specimens bearing names of 604 species-group taxa of Carnivora, Perissodactyla, Artiodactyla, and Cetacea as of May 2016. This catalog presents an annotated list of these holdings comprising 582 holotypes; 16 lectotypes, two of which are newly designated herein; 7 syntypes (15 specimens); and 1 neotype. Included are several specimens that should be in the collection but cannot be found or are now known to be in other collections and therefore are not in the database. Thirty-seven of the names are new since the last type catalog covering these orders, Arthur J. Poole and Viola S. Schantz’s 1942 “Catalog of the Type Specimens of Mammals in the United States National Museum, Including the Biological Surveys Collection” (Bulletin of the United States National Museum, 178). One of these, Lutra iowa Goldman, 1941, was transferred to the National Museum’s Paleobiology Department collection and is mentioned only briefly in this work. Orders and families are arranged systematically following D. E. Wilson and D. M. Reeder’s 2005 Mammal Species of the World: A Taxonomic and Geographic Reference, third edition, volume 1; within families, currently recognized genera are arranged alphabetically, and within each currently recognized genus, spe- cies and subspecies accounts are arranged alphabetically by original published name. Information in each account includes original name and abbreviated citation thereto, current name if other than original, citation for first use of current name combination for the taxon, type designation, U.S. National Museum catalog number(s), preparation, age and sex, date of collection and collector, original collector number, type locality, and remarks as appropriate. Digital photographs of each specimen will serve as a condition report and will be attached to each electronic speci- men record. An addendum contains two accounts for holotypes added to the collection subsequent to the publica- tion of the catalog for their taxa. Appendices tabulate summary data for all four of our recent type catalogs (Fisher and Ludwig, 2012, 2014, 2015, and this volume) and include authors of names, collectors of type specimens, coun- tries, islands, and provinces or states in which type specimens were collected, numbers of new names per decade, and summary numbers for holotypes, lectotypes, syntypes, neotypes, and new taxa since the last type catalog (Poole and Schantz, 1942) covering the entire type collection. Cover images: Holotype of Lynchailurus colocolo wolfshoni Garcia-Perea, 1994 (USNM 391853). Left to right: dorsal, lateral, and ventral views of cranium. Photographs by Renee Regan. Published by SMITHSONIAN INSTITUTION SCHOLARLY PRESS P.O. Box 37012, MRC 957 Washington, D.C. 20013-7012 www.scholarlypress.si.edu The text of this publication is in the public domain. The rights to all other text and images in this publication, in- cluding cover and interior designs, are owned either by the Smithsonian Institution, by contributing authors, or by third parties. Fair use of materials is permitted for personal, educational, or noncommercial purposes. Users must cite author and source of content, must not alter or modify copyrighted content, and must comply with all other terms or restrictions that may be applicable. Users are responsible for securing permission from a rights holder for any other use. Library of Congress Cataloging-in-Publication Data Names: National Museum of Natural History (U.S.) | Fisher, Robert D. | Ludwig, Craig A. Title: Catalog of type specimens of recent mammals : orders Carnivora, Perissodactyla, Artiodactyla, and Cetacea in the National Museum of Natural History, Smithsonian Institution / Robert D. Fisher and Craig A. Ludwig. Description: Washington, D.C. : Smithsonian Institution Scholarly Press, 2016. | Series: Smithsonian contributions to zoology ; number 646 Identifiers: LCCN 2016008481 Subjects: LCSH: Carnivora—Type specimens—Catalogs and collections—Washington (D.C.) | Ungulates—Type specimens—Catalogs and collections—Washington (D.C.) | Cetacea—Type specimens—Catalogs and collections— Washington (D.C.) | National Museum of Natural History (U.S.)—Catalogs. Classification: LCC QL737.C2 N37 2016 | DDC 599.7—dc23 LC record available at https://lccn.loc.gov/2016008481 ISSN: 1943-6696 (online); 0081-0282 (print) ZooBank registration: 20 May 2016 (LSID: urn:lsid:zoobank.org:pub:24951B6F-DD78-4BCF-ABF3-604435CE098D) Publication date (online): 8 June 2016 Ó The paper used in this publication meets the minimum requirements of the American National Standard for Perma- nence of Paper for Printed Library Materials Z39.48–1992. Contents FOREWORD 1 Robert P. Reynolds and Kristofer M. Helgen INTRODUCTION 2 Categories of Name-Bearing Types Included 2 Research Approach 3 Classificatory Organization 3 Format of Accounts 3 Acknowledgments 4 ACCOUNTS OF NAME-BEARING TYPE SPECIMENS 4 Order Carnivora Bowdich, 1821 4 Family Felidae Fischer, 1817 4 Genus Acinonyx Brookes, 1828 4 Genus Felis Linnaeus, 1758 4 Genus Herpailurus Severtzov, 1858 5 Genus Leopardus Gray, 1842 5 Genus Lynx Kerr, 1792 6 Genus Panthera Oken, 1816 6 Genus Prionailurus Severtzov, 1858 7 Genus Puma Jardine, 1834 8 Family Viverridae Gray, 1821 9 Genus Arctictis Temminck, 1824 9 Genus Arctogalidia Merriam, 1897 9 Genus Genetta G. Cuvier, 1816 10 Genus Hemigalus Jourdan, 1837 10 Genus Paguma Gray, 1831 10 Genus Paradoxurus F. Cuvier, 1821 10 Genus Viverricula Hodgson, 1838 11 Family Nandiniidae Pocock, 1929 12 Genus Nandinia Gray, 1843 12 Family Herpestidae Bonaparte, 1845 12 Genus Atilax F. Cuvier, 1826 12 Genus Bdeogale Peters, 1850 12 Genus Galerella Gray, 1865 12 Genus Helogale Gray, 1862 12 iv • SMITHSONIAN CONTRIBUTIONS TO ZOOLOGY Genus Ichneumia I. Geoffroy, 1837 12 Genus Mungos E. Geoffroy and G. Cuvier, 1795 12 Family Hyaenidae Gray, 1828 13 Genus Crocuta Kaup, 1828 13 Genus Proteles I. Geoffroy, 1824 13 Family Canidae Fischer, 1817 13 Genus Canis Linnaeus, 1758 13 Genus Otocyon Muller, 1836 16 Genus Speothos Lund, 1839 16 Genus Urocyon Baird, 1857 16 Genus Vulpes Frisch, 1777 17 Family Ursidae Fischer, 1817 19 Genus Ursus Linnaeus, 1758 19 Family Otariidae Gray, 1825 27 Genus Arctocephalus E. Geoffroy and F. Cuvier, 1826 27 Family Phocidae Gray, 1821 27 Genus Mirounga Gray, 1827 27 Genus Phoca Linnaeus, 1758 27 Family Mustelidae Fischer, 1817 28 Genus Aonyx Lesson, 1827 28 Genus Enhydra Fleming, 1822 28 Genus Galictis Bell, 1826 28 Genus Gulo Pallas, 1780 28 Genus Ictonyx Kaup, 1835 28 Genus Lontra Gray, 1843 28 Genus Lutra Brisson, 1762 29 Genus Martes Pinel, 1792 29 Genus Mellivora Storr, 1780 30 Genus Mustela Linnaeus, 1758