Chapter 36 Transport in Vascular Plants Lecture Outline
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Transpiration
TRANSPIRATION BY: Dr. Madhu Gupta (Guest Faculty) SOS in Botany Jiwaji University Gwalior What is it? The loss of water in the vapour form from the exposed parts of a plant is called transpiration. The loss of water due to transpiration is quite high. Rather 98-99% of the water absorbed by a plant is lost in transpiration. Hardly 0.2% is used in photosynthesis while the remaining is retained in the plant during growth. Most of the transpiration occurs through foliar surface or surface of the leaves. It is known as foliar transpiration. Foliar transpiration accounts for over 90% of the total transpiration. Transpiration occurs through young or mature stem is called as Cauline transpiration. Depending upon the plant surface, transpiration is classified into three types: Stomatal • Water vapour diffuses out through minute pore (stomata) present in soft aerial part of plant is known Transpiration as Stomatal Transpiration • Sometimes water may evaporate through certain Lenticular other openings present on the older stems. These openings are called Lenticels and the transpiration Transpiration that takes place through term is known as Lenticular Transpiration. • Loss of water may also take place through cuticle, but Cuticular the amount so lost is relatively small • This type of transpiration depends upon the thickness Transpiration of the cuticle and presence or absence of wax coating on the surface of the leaves. Stomatal Transpiration Lenticular Transpiration Cuticular Transpiration Factors Affecting Transpiration: Water Stress: Whenever the rate of transpiration exceeds the rate of absorption, a water deficit is created in the plants and results in the incipient wilting of leaves. -
Photosynthesis Respiration and Transpiration.Notebook February 13, 2017
Photosynthesis Respiration and Transpiration.notebook February 13, 2017 Essential Question: What processes are required for plant survival? Key Concept: All living things need energy to carry out their basic functions. Living things break down food to get their energy.One thing that makes plants different from most others organisms is how they get their food. 1 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 There are 3 processes that take place in the leaves that are mandatory for plant to survive. Photosynthesis Respiration Transpiration 2 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 ALL PLANTS ARE AUTOTROPHS which means they make their own food for energy. Food for plants is SUGAR Remember: Chloroplast: Attract sunlight with chlorophyll Mitochondria: Energy Factory 3 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 Observe the diagrams below. Can you infer which one is photosynthesis and respiration? 4 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 What is needed: 1st Photosynthesis 1. Sunlight 2. Carbon Dioxide 3. Water What it Makes: Sugar (keeps) Oxygen (released for humans) Where it happens: Chloroplasts Sunlight Water H20 Carbon Dioxide CO2 Equation: Sunlight + 6H2O + 6CO2 = 6O2 + C6H12O6 5 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 6 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 Water Photosynthesis Sugar Carbon Oxygen Dioxide Excess Solar Energy Water 7 Photosynthesis Respiration and Transpiration.notebook February 13, 2017 Photosynthesis makes sugar, but in order to use the food the plant must break it down into usable energy through a process called Respiration. Food Respiration is the process of how plants break down the sugar so the plant can use it for energy. -
Plant Water Relations: Absorption, Transport and Control Mechanisms
5 Plant Water Relations: Absorption, Transport and Control Mechanisms Geraldo Chavarria1 and Henrique Pessoa dos Santos2 1The University of Passo Fundo 2Embrapa Grape & Wine Brazil 1. Introduction Although water is abundant on Earth - covering 71% of the total surface - its distribution is not uniform and can easily cause restrictions in availability to vegetal production. At global scale, these restrictions are easily observed in dry climates and can appear in other regions which do not currently experience drought, as provided by the future backdrop of climate change (IPCC, 2007). The influences of water restriction on losses in the production and distribution of vegetation on the terrestrial surface are significantly larger than all other losses combined which are caused by biotic and abiotic factors (Boyer, 1985). This striking effect of water on plants emerges from its physiological importance, being an essential factor for successful plant growth, involving photosynthesis and several other biochemical processes such as the synthesis of energetic composites and new tissue. Therefore, in order to characterise the growth and productive behaviour of plant species it is essential to have an understanding of plant water relations, as well as the consequences of an inadequate water supply. Broadly, the water state of a plant is controlled by relative rates of loss and absorption, moreover it depends on the ability to adjust and keep an adequate water status. This will be considered throughout this chapter. 2. Absorption and water flow through plants Independent of the species, plants require from the soil a water volume that overcomes its metabolic necessities. Through the transpiration process plants transmit to the atmosphere the majority of the water absorbed from soil (generally around 90%). -
Anatomy of Leaf Apical Hydathodes in Four Monocotyledon Plants of Economic and Academic Relevance Alain Jauneau, Aude Cerutti, Marie-Christine Auriac, Laurent D
Anatomy of leaf apical hydathodes in four monocotyledon plants of economic and academic relevance Alain Jauneau, Aude Cerutti, Marie-Christine Auriac, Laurent D. Noël To cite this version: Alain Jauneau, Aude Cerutti, Marie-Christine Auriac, Laurent D. Noël. Anatomy of leaf apical hydathodes in four monocotyledon plants of economic and academic relevance. PLoS ONE, Public Library of Science, 2020, 15 (9), pp.e0232566. 10.1371/journal.pone.0232566. hal-02972304 HAL Id: hal-02972304 https://hal.inrae.fr/hal-02972304 Submitted on 20 Oct 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. Distributed under a Creative Commons Attribution| 4.0 International License PLOS ONE RESEARCH ARTICLE Anatomy of leaf apical hydathodes in four monocotyledon plants of economic and academic relevance 1☯ 2☯ 1,2 2 Alain Jauneau *, Aude Cerutti , Marie-Christine Auriac , Laurent D. NoeÈlID * 1 FeÂdeÂration de Recherche 3450, Universite de Toulouse, CNRS, Universite Paul Sabatier, Castanet- Tolosan, France, 2 LIPM, Universite de Toulouse, INRAE, CNRS, Universite Paul Sabatier, Castanet- Tolosan, France ☯ These authors contributed equally to this work. a1111111111 * [email protected] (AJ); [email protected] (LN) a1111111111 a1111111111 a1111111111 a1111111111 Abstract Hydathode is a plant organ responsible for guttation in vascular plants, i.e. -
3115 SAPS Operating Instructions
OPERATING INSTRUCTIONS FIELD PLANT WATER STATUS CONSOLE March 2017 Fig. 1 - Model 3115 Portable Plant Water Console shown with 22 c/f Compressed Gas Cylinder, 3072V22 SOILMOISTURE EQUIPMENT CORP. P.O. Box 30025, Santa Barbara, CA 93105 U.S.A. Telephone 805-964-3525 - Fax No. 805-683-2189 Email: [email protected] - Website: http://www.soilmoisture.com Table of Contents Chapter - Page 1 Description ............................................................................................................................... 1 - 4 2 Technical Specification ............................................................................................................ 2 - 5 2.1 Weight .......................................................................................................................... 2 - 5 2.2 Dimensions ................................................................................................................... 2 - 5 2.3 Pressure Vessel ............................................................................................................. 2 - 5 2.4 Gauges .......................................................................................................................... 2 - 5 2.5 Valves ........................................................................................................................... 2 - 5 2.6 Connecting Hose .......................................................................................................... 2 - 5 2.7 Pressure Tank .............................................................................................................. -
197 Section 9 Sunflower (Helianthus
SECTION 9 SUNFLOWER (HELIANTHUS ANNUUS L.) 1. Taxonomy of the Genus Helianthus, Natural Habitat and Origins of the Cultivated Sunflower A. Taxonomy of the genus Helianthus The sunflower belongs to the genus Helianthus in the Composite family (Asterales order), which includes species with very diverse morphologies (herbs, shrubs, lianas, etc.). The genus Helianthus belongs to the Heliantheae tribe. This includes approximately 50 species originating in North and Central America. The basis for the botanical classification of the genus Helianthus was proposed by Heiser et al. (1969) and refined subsequently using new phenological, cladistic and biosystematic methods, (Robinson, 1979; Anashchenko, 1974, 1979; Schilling and Heiser, 1981) or molecular markers (Sossey-Alaoui et al., 1998). This approach splits Helianthus into four sections: Helianthus, Agrestes, Ciliares and Atrorubens. This classification is set out in Table 1.18. Section Helianthus This section comprises 12 species, including H. annuus, the cultivated sunflower. These species, which are diploid (2n = 34), are interfertile and annual in almost all cases. For the majority, the natural distribution is central and western North America. They are generally well adapted to dry or even arid areas and sandy soils. The widespread H. annuus L. species includes (Heiser et al., 1969) plants cultivated for seed or fodder referred to as H. annuus var. macrocarpus (D.C), or cultivated for ornament (H. annuus subsp. annuus), and uncultivated wild and weedy plants (H. annuus subsp. lenticularis, H. annuus subsp. Texanus, etc.). Leaves of these species are usually alternate, ovoid and with a long petiole. Flower heads, or capitula, consist of tubular and ligulate florets, which may be deep purple, red or yellow. -
Summary a Plant Is an Integrated System Which: 1
Summary A plant is an integrated system which: 1. Obtains water and nutrients from the soil. 2. Transports them 3. Combines the H2O with CO2 to make sugar. 4. Exports sugar to where it’s needed Today, we’ll start to go over how this occurs Transport in Plants – Outline I.I. PlantPlant waterwater needsneeds II.II. TransportTransport ofof waterwater andand mineralsminerals A.A. FromFrom SoilSoil intointo RootsRoots B.B. FromFrom RootsRoots toto leavesleaves C.C. StomataStomata andand transpirationtranspiration WhyWhy dodo plantsplants needneed soso muchmuch water?water? TheThe importanceimportance ofof waterwater potential,potential, pressure,pressure, solutessolutes andand osmosisosmosis inin movingmoving water…water… Transport in Plants 1.1. AnimalsAnimals havehave circulatorycirculatory systems.systems. 2.2. VascularVascular plantsplants havehave oneone wayway systems.systems. Transport in Plants •• OneOne wayway systems:systems: plantsplants needneed aa lotlot moremore waterwater thanthan samesame sizedsized animals.animals. •• AA sunflowersunflower plantplant “drinks”“drinks” andand “perspires”“perspires” 1717 timestimes asas muchmuch asas aa human,human, perper unitunit ofof mass.mass. Transport of water and minerals in Plants WaterWater isis goodgood forfor plants:plants: 1.1. UsedUsed withwith CO2CO2 inin photosynthesisphotosynthesis toto makemake “food”.“food”. 2.2. TheThe “blood”“blood” ofof plantsplants –– circulationcirculation (used(used toto movemove stuffstuff around).around). 3.3. EvaporativeEvaporative coolingcooling. -
Co-Ordination of Signalling Elements in Guard Cell Ion Channel Control
Journal of Experimental Botany, Vol. 49, Special Issue, pp. 351–360, March 1998 Co-ordination of signalling elements in guard cell ion channel control A. Grabov1 and M.R. Blatt Laboratory of Plant Physiology and Biophysics, Wye College, University of London, Wye, Ashford, Kent TN25 5AH, UK Received 3 November 1997; Accepted 10 November 1997 Abstract ably important for providing a plasticity of cellular response to external and environmental stimuli. Fine regulation of solutes transport across the guard Understanding the interdependence and hierarchy of cell plasma membrane for osmotic modulation is signalling elements now presents a major challenge essential for the maintenance of the proper stomatal for research in plant biology. aperture in response to environmental stimuli. The major osmotica, K+,Cl− and malate are transported Key words: Ion channels, stomatal aperture, signalling through selective ion channels in the plasma mem- pathway, second messengers, guard cells. brane and tonoplast of guard cells. To date, a number of ion channels have been shown to operate in the guard cell plasma membrane: outwardly- and inwardly- Introduction + rectifying K channels (IK,in and IK,out), slowly- and Guard cell control of stomatal aperture is crucial to rapid-activating anion channels, and stretch-activated balancing transpiration stream and gas exchange for non-selective channels. Slow and fast vacuolar chan- photosynthesis. As transpiration and CO2 exchange can nels (SV and FV) and voltage-independent K+-selective not be regulated independently, the demand of photosyn- (VK) channels have been found at the guard cell tono- thetic machinery can lead to excessive water evaporation plast. On the molecular level, the regulation of the under adverse environmental conditions. -
The Relationship Between Plant Growth and Water Consumption : a History from Greek Philosophers to Early 20Th Century Scientists
The relationship between plant growth and water consumption : A history from Greek philosophers to early 20th century scientists. Oliver Brendel Université de Lorraine, AgroParisTech, INRA, UMR SILVA Nancy, France email : [email protected] Tel : 00 33 383394100 postal address: Oliver Brendel, INRAE, UMR Silva, F-54280 CHAMPENOUX, France Abstract The relationship between plant growth and water consumption has for a long time occupied the minds of philosophers and natural scientists. The ratio between biomass accumulation and water consumption is known as water use efficiency and is widely relevant today in fields as diverse as crop improvement, forest ecology and climate change. Defined at scales varying from single leaf physiology to whole plants, it shows how botanical investigations changed through time, generally in tandem with developing disciplines and improving methods. The history started as a purely philosophical question by Greek philosophers of how plants grow, progressed through thought and actual experiments, towards an interest in plant functioning and their relationship to the environment. This article retraces this history by elucidating the progression of scientific questions posed through the centuries, presents the main methodological and conceptual developments. Keywords Transpiration efficiency; water use efficiency; plant physiology; botany Introduction The ratio of biomass accumulation per unit water consumption is known today as water use efficiency (WUE) and is widely relevant to agriculture ( e.g. Vadez et al.2014; Tallec et al.; Blum 2009), to forest ecology (e.g. Linares and Camarero 2012; Lévesque et al. 2014), and in the context of global climate change (e.g., Cernusak et al. 2019). This ratio can be defined at various levels, from the physiological functioning of a leaf to the whole plant and at the ecosystem level. -
Differences Between Transpiration and Guttation
BIOLOGY TRANSPIRATION Significance and Factors affecting Transpiration Factors Affecting the Rate of Transpiration •On a bright sunny day, stomata open fully, so transpiration is increased. •On a cloudy day, stomata open partially, so Intensity of sunlight transpiration is reduced. •At night, stomata close; hence, transpiration is greatly reduced or negligible. •Increase in temperature of the air increases the rate Temperature of transpiration. •Transpiration increases with rapid or active air Velocity of wind movement. Humidity •If the air is humid, the rate of transpiration is reduced. •Increase in the CO2 level in the atmosphere over Carbon dioxide normal 0.03% causes stomatal closure. Hence, it decreases the rate of transpiration. •With decrease in atmospheric pressure, the rate of Atmospheric pressure transpiration increases. www.topperlearning.com 2 BIOLOGY TRANSPIRATION Adaptation in Plants to Control Excessive Transpiration Plants which grow in dry climate have evolved a variety of adaptations to curtail transpiration. Morphological Leaves may be modified into spines as in cactus or into needles as in pines. Adaptations Spines Needles Leaves may be folded or rolled up. Rolled up Leaves Leaves may be shed. Example: deciduous trees. Deciduous Trees Anatomical The number of stomata is reduced, and they may be sunken in pits. Adaptations Sunken Stoma www.topperlearning.com 3 BIOLOGY TRANSPIRATION Structure of Sunken Stomata A thick waxy cuticle develops on the leaves. Example: Banyan tree, evergreen trees. Banyan Tree Shrubs and grass develop a waterproof covering of cork or bark. A multiple epidermis may develop in some leaves. www.topperlearning.com 4 BIOLOGY TRANSPIRATION Significance of Transpiration Cooling Effect Suction Force Distribution of Water and Mineral Salts Evaporation reduces the As water evaporates from leaves, Higher the rate of transpiration, temperature of leaf a suction force is created. -
Historical Review
1 Historical Review INTRODUCTION This chapter presents a brief historical review of progress in the field of plant water relations because the authors feel that it is impossible to fully understand the present without some knowledge of the past. As the Danish philosopher Kierkegaarde wrote, "Life can only be understood backward, but it can only be lived forward," and this also is true of science. The present generation needs to be reminded that some generally accepted concepts have their origin in ideas of 17th or 18th century writers and although others were suggested many decades ago, they were neglected until recently. As might be expected, the importance of water to plant growth was recog- nized by prehistoric farmers because irrigation systems already existed in Egypt, Babylonia (modern Iraq), and China at the beginning of recorded history, and the first European explorers found extensive irrigation systems in both North and South America. However, irrigation was not used extensively in agriculture in the United States until after the middle of the 19th century and little research on plant water relations occurred until the 20th century. Early Research Although plant water relations appear to have been the first area of plant physiology to be studied, progress was slow from Aristotle who died in 322 B.C. to the middle of the 19th century. According to Aristotle, plants absorbed their food ready for use from the soil, and plant nutrition was controlled by a soul or vital principle that ailowed plants to absorb only those substances useful in 2 1. Historical Review growth. This idea only began to be questioned in the 17th century by Jung, van Helmont, Mariotte, and others, and it ~ersistedinto the 19th century. -
Plant Adaptation to Fluctuating Environment and Biomass Production Are Strongly Dependent on Guard Cell Potassium Channels
Plant adaptation to fluctuating environment and biomass production are strongly dependent on guard cell potassium channels Anne Lebaudy*, Alain Vavasseur†, Eric Hosy*, Ingo Dreyer*‡, Nathalie Leonhardt†, Jean-Baptiste Thibaud*, Anne-Alie´ nor Ve´ ry*, Thierry Simonneau§, and Herve´ Sentenac*¶ *Biochimie et Physiologie Mole´culaire des Plantes, Unite´Mixte de Recherche 5004, Centre National de la Recherche Scientifique/Institut National de la Recherche Agronomique (U.386)/Montpellier SupAgro/Universite´Montpellier 2, 1 Place Viala, 34060 Montpellier Cedex 1, France; †Laboratoire des Echanges Membranaires et Signalisation, Unite´Mixte de Recherche 6191, Centre National de la Recherche Scientifique/Commissariat a`l’Energie Atomique/Universite´ Aix-Marseille, 13108 St. Paul lez Durance Cedex, France; and §Laboratoire d’Ecophysiologie des Plantes sous Stress Environnementaux, Unite´Mixte de Recherche 759, Institut National de la Recherche Agronomique/Montpellier SupAgro, 1 Place Viala, 34060 Montpellier Cedex 1, France Edited by Maarten J. Chrispeels, University of California at San Diego, La Jolla, CA, and approved February 8, 2008 (received for review October 12, 2007) At least four genes encoding plasma membrane inward K؉ chan- plant physiology by engineering an Arabidopsis mutant totally nels (Kin channels) are expressed in Arabidopsis guard cells. A deprived of this activity. double mutant plant was engineered by disruption of a major Kin channel gene and expression of a dominant negative channel Results construct. Using the patch-clamp technique revealed that this Genetic Engineering of an Arabidopsis Mutant Deprived of GCKin mutant was totally deprived of guard cell Kin channel (GCKin) Activity. In a first step, we screened candidate mutant lines in activity, providing a model to investigate the roles of this activity which expression of inward Kϩ channels is affected, by looking in the plant.