Revision of the Snail Genus <I>Austropyrgus</I> (Gastropoda
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www.amonline.net.au/publications/ ISBN 0 7347 2313 X RECORDS OF THE AUSTRALIAN MUSEUM 2003 Supplement 28 RECORDS OF THE AUSTRALIAN MUSEUM Revision of the Snail Genus Austropyrgus (Gastropoda: Hydrobiidae) A Morphostatic Radiation of Freshwater Gastropods in Southeastern Australia by A M Stephanie A. Clark, AUSTRALIAN Alison C. Miller and Winston F. Ponder MUSEUM Publications may be purchased at the Australian Museum Shop or online at: www.amonline.net.au/shop/ SUPPLEMENT 28 25 June 2003 Revision of the Snail Genus Austropyrgus (Gastropoda: Hydrobiidae) A Morphostatic Radiation of Freshwater Gastropods in Southeastern Australia by Stephanie A. Clark, Alison C. Miller and Winston F. Ponder A M AUSTRALIAN MUSEUM Records of the Australian Museum, Supplement 28 (2003) ISBN 0 7347 2313 X © Copyright Australian Museum, 2003 Records of the Australian Museum, Supplement 28 (2003): 1–109 ISBN 0-7347-2313-X Revision of the Snail Genus Austropyrgus (Gastropoda: Hydrobiidae): A Morphostatic Radiation of Freshwater Gastropods in Southeastern Australia STEPHANIE A. CLARK1, ALISON C. MILLER AND WINSTON F. PONDER Australian Museum, 6 College Street, Sydney NSW 2010, Australia 1 current address: Department of Biological Sciences, Biodiversity & Systematics, University of Alabama, Box 870345, Tuscaloosa, AL 35487, Unites States of America [email protected] · [email protected] · [email protected] ABSTRACT. The species that comprise a morphostatic radiation in southeastern Australia of the hydrobiid genus Austropyrgus Cotton are described using shell, opercular, anatomical and radular characters, and the genus is redefined. The previously described taxa have been included under several generic names, including Fluvidona (now a separate genus), Rivisessor, Pupiphryx and Angrobia, which are now treated as synonyms of Austropyrgus. Seventy-four species are described from southeastern Australia, fifty- seven of them new. These are grouped into six informal morphological groups to aid identification. Two recently described outlying species, A. bunyaensis from Mt Bunya, southern Queensland and A. centralia from Dalhousie Springs, northern South Australia, are not included in this revision. Members of the genus are typically found in streams, although a few species live in springs, rivers or lakes. A large number of the species have restricted distributions and several are known only from single locations. In southern New South Wales, Victoria and South Australia Austropyrgus is the dominant native freshwater hydrobiid genus whereas in Tasmania it is often sympatric with the native freshwater hydrobiid genera Beddomeia and/or Phrantela. CLARK, STEPHANIE A., ALISON C. MILLER & WINSTON F. PONDER, 2003. Revision of the snail genus Austropyrgus (Gastropoda: Hydrobiidae): a morphostatic radiation of freshwater gastropods in southeastern Australia. Records of the Australian Museum, Supplement 28: 1–109. Contents Introduction ............................................................................................................................... 3 Materials and methods............................................................................................................... 4 Abbreviations............................................................................................................................. 5 Taxonomy .................................................................................................................................. 5 Family Hydrobiidae Troschel, 1857.......................................................................................... 5 Genus Austropyrgus Cotton, 1942 ............................................................................................ 5 Group 1. Austropyrgus cooma group ........................................................................................ 6 Mainland taxa ..................................................................................................................... 6 Austropyrgus cooma (Iredale, 1943) .......................................................................... 6 Austropyrgus monaroensis n.sp.................................................................................. 9 Austropyrgus viridarium n.sp. .................................................................................. 13 www.amonline.net.au/pdf/publications/1377_complete.pdf 2 Records of the Australian Museum, Supplement 28 (2003) Austropyrgus lippus n.sp. ................................................................................... 14 Austropyrgus avius n.sp. .................................................................................... 16 Austropyrgus bullerensis n.sp. ........................................................................... 17 Austropyrgus grampianensis (Gabriel, 1939).................................................... 17 Austropyrgus daylesfordensis n.sp. .................................................................... 18 Austropyrgus otwayensis n.sp. ........................................................................... 19 Austropyrgus macedonensis n.sp. ...................................................................... 22 Austropyrgus colensis n.sp. ................................................................................ 22 Tasmanian taxa .......................................................................................................... 23 Austropyrgus nanus n.sp. ................................................................................... 23 Austropyrgus mersus n.sp. ................................................................................. 26 Austropyrgus smithii (Petterd, 1889) ................................................................. 27 Austropyrgus pisinnus n.sp. ............................................................................... 28 Austropyrgus parvus n.sp. .................................................................................. 28 Austropyrgus solitarius n.sp. ............................................................................. 29 Austropyrgus conicus n.sp. ................................................................................ 30 Group 2. Austropyrgus petterdianus group ....................................................................... 31 Austropyrgus petterdianus (Brazier, 1875) ........................................................ 31 Austropyrgus juliae n.sp..................................................................................... 33 Austropyrgus nanoacuminatus n.sp. .................................................................. 35 Austropyrgus stevensmithi n.sp. ......................................................................... 36 Austropyrgus privus n.sp. ................................................................................... 36 Austropyrgus lochi n.sp. ..................................................................................... 37 Group 3. Austropyrgus sparsus group .............................................................................. 38 Southern New South Wales taxa ............................................................................... 38 Austropyrgus sparsus (Iredale, 1944) ................................................................ 38 Austropyrgus bungoniensis n.sp. ....................................................................... 41 Austropyrgus synoria n.sp. ................................................................................. 43 Austropyrgus tathraensis n.sp. ........................................................................... 45 Austropyrgus wombeyanensis n.sp. ................................................................... 47 Austropyrgus nepeanensis n.sp. ......................................................................... 49 Austropyrgus abercrombiensis n.sp. .................................................................. 50 Eastern Victorian taxa ................................................................................................ 51 Austropyrgus dekeyzeri n.sp. ............................................................................. 51 Austropyrgus buchanensis n.sp. ......................................................................... 52 Austropyrgus gippslandicus (Ponder et al., 1994) ............................................ 53 Austropyrgus turbatus (Ponder et al., 1994) ..................................................... 54 Austropyrgus macaulayi n.sp. ............................................................................ 56 Austropyrgus ora n.sp. ....................................................................................... 57 Western Victorian taxa ............................................................................................... 59 Austropyrgus pusillus n.sp. ................................................................................ 59 Austropyrgus exiguus n.sp. ................................................................................ 59 Austropyrgus spectus n.sp. ................................................................................. 62 Austropyrgus tumidus n.sp. ................................................................................ 63 Austropyrgus glenelgensis n.sp. ......................................................................... 63 Austropyrgus aslini n.sp. .................................................................................... 64 Austropyrgus angasi (Smith, 1882) ..................................................................