THE PRINTING WOOD BLOCK COLLECTION of the GEOLOGICAL SURVEY of IRELAND by M.A
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Proposed Global Standard Stratotype-Section And
PROPOSED GLOBAL STANDARD STRATOTYPE-SECTION AND POINT FOR THE DRUMIAN STAGE (CAMBRIAN) Prepared for the International Subcommission on Cambrian Stratigraphy by: Loren E. Babcock School of Earth Sciences The Ohio State University, 125 South Oval Mall Columbus, OH 43210, USA [email protected] Richard A. Robison Department of Geology, University of Kansas Lawrence, KS 66045, USA [email protected] Margaret N. Rees Department of Geoscience University of Nevada, Las Vegas Las Vegas, NV 89145, USA [email protected] Peng Shanchi Nanjing Institute of Geology and Palaeontology Chinese Academy of Sciences 39 East Beijing Road, Nanjing 210008, China [email protected] Matthew R. Saltzman School of Earth Sciences The Ohio State University, 125 South Oval Mall Columbus, OH 43210, USA [email protected] 31 July 2006 CONTENTS Introduction …………………………………………………………………………. 2 Proposal: Stratotype Ridge, Drum Mountains (Millard County, Utah, USA) as the GSSP for the base of the Drumian Stage ……………………………………………………. 3 1. Stratigraphic rank of the boundary …………………..…………………………… 3 2. Proposed GSSP – geography and physical geology ……………………………… 3 2.1. Geographic location …………………………………………………...….. 3 2.2. Geological location ………………..……………………...…………...….. 4 2.3. Location of level and specific point ……………………..……………...… 4 2.4. Stratigraphic completeness ………………...……………………………… 5 2.5. Thickness and stratigraphic extent …………………...……………...……. 5 2.6. Provisions for conservation, protection, and accessibility ……………..…. 5 3. Motivation for selection of the boundary level and of the potential stratotype section ………………………………………………………………………………. 6 3.1. Principal correlation event (marker) at GSSP level ………...…………..… 6 3.2. Potential stratotype section …………………………………………....….. 7 3.3. Demonstration of regional and global correlation ………………………... 8 3.3.1. Agnostoid trilobite biostratigraphy ………………………………… 9 3.3.2. Polymerid trilobite biostratigraphy …………....…………….……. -
The Weeks Formation Konservat-Lagerstätte and the Evolutionary Transition of Cambrian Marine Life
Downloaded from http://jgs.lyellcollection.org/ by guest on October 1, 2021 Review focus Journal of the Geological Society Published Online First https://doi.org/10.1144/jgs2018-042 The Weeks Formation Konservat-Lagerstätte and the evolutionary transition of Cambrian marine life Rudy Lerosey-Aubril1*, Robert R. Gaines2, Thomas A. Hegna3, Javier Ortega-Hernández4,5, Peter Van Roy6, Carlo Kier7 & Enrico Bonino7 1 Palaeoscience Research Centre, School of Environmental and Rural Science, University of New England, Armidale, NSW 2351, Australia 2 Geology Department, Pomona College, Claremont, CA 91711, USA 3 Department of Geology, Western Illinois University, 113 Tillman Hall, 1 University Circle, Macomb, IL 61455, USA 4 Department of Zoology, University of Cambridge, Downing Street, Cambridge CB2 3EJ, UK 5 Museum of Comparative Zoology and Department of Organismic and Evolutionary Biology, Harvard University, 26 Oxford Street, Cambridge, MA 02138, USA 6 Department of Geology, Ghent University, Krijgslaan 281/S8, B-9000 Ghent, Belgium 7 Back to the Past Museum, Carretera Cancún, Puerto Morelos, Quintana Roo 77580, Mexico R.L.-A., 0000-0003-2256-1872; R.R.G., 0000-0002-3713-5764; T.A.H., 0000-0001-9067-8787; J.O.-H., 0000-0002- 6801-7373 * Correspondence: [email protected] Abstract: The Weeks Formation in Utah is the youngest (c. 499 Ma) and least studied Cambrian Lagerstätte of the western USA. It preserves a diverse, exceptionally preserved fauna that inhabited a relatively deep water environment at the offshore margin of a carbonate platform, resembling the setting of the underlying Wheeler and Marjum formations. However, the Weeks fauna differs significantly in composition from the other remarkable biotas of the Cambrian Series 3 of Utah, suggesting a significant Guzhangian faunal restructuring. -
AND MICROFOSSIL RECORD of the CAMBRIAN PRIAPULID OTTOIA by MARTIN R
[Palaeontology, 2015, pp. 1–17] THE MACRO- AND MICROFOSSIL RECORD OF THE CAMBRIAN PRIAPULID OTTOIA by MARTIN R. SMITH1,THOMASH.P.HARVEY2 and NICHOLAS J. BUTTERFIELD1 1Department of Earth Sciences, University of Cambridge, Cambridge, UK; e-mails: [email protected], [email protected] 2Department of Geology, University of Leicester, Leicester, UK ; e-mail: [email protected] Typescript received 11 December 2014; accepted in revised form 31 March 2015 Abstract: The stem-group priapulid Ottoia Walcott, 1911, Ottoiid priapulids represented an important component of is the most abundant worm in the mid-Cambrian Burgess Cambrian ecosystems: they occur in a range of lithologies Shale, but has not been unambiguously demonstrated else- and thrived in shallow water as well as in the deep-water where. High-resolution electron and optical microscopy of setting of the Burgess Shale. A wider survey of Burgess macroscopic Burgess Shale specimens reveals the detailed Shale macrofossils reveals specific characters that diagnose anatomy of its robust hooks, spines and pharyngeal teeth, priapulid sclerites more generally, establishing the affinity establishing the presence of two species: Ottoia prolifica of a wide range of Small Carbonaceous Fossils and demon- Walcott, 1911, and Ottoia tricuspida sp. nov. Direct com- strating the prominent role of priapulids in Cambrian parison of these sclerotized elements with a suite of shale- seas. hosted mid-to-late Cambrian microfossils extends the range of ottoiid priapulids throughout the middle to upper Key words: Burgess Shale, Small Carbonaceous Fossils, pri- Cambrian strata of the Western Canada Sedimentary Basin. apulid diversity, Selkirkia. S TEM-group priapulid worms were a conspicuous compo- in analyses of its ecological and evolutionary significance nent of level-bottom Cambrian faunas (Conway Morris (Wills 1998; Bruton 2001; Vannier 2012; Wills et al. -
GSSP) of the Drumian Stage (Cambrian) in the Drum Mountains, Utah, USA
Articles 8585 by Loren E. Babcock1, Richard A. Robison2, Margaret N. Rees3, Shanchi Peng4, and Matthew R. Saltzman1 The Global boundary Stratotype Section and Point (GSSP) of the Drumian Stage (Cambrian) in the Drum Mountains, Utah, USA 1 School of Earth Sciences, The Ohio State University, 125 South Oval Mall, Columbus, OH 43210, USA. Email: [email protected] and [email protected] 2 Department of Geology, University of Kansas, Lawrence, KS 66045, USA. Email: [email protected] 3 Department of Geoscience, University of Nevada, Las Vegas, Las Vegas, NV 89145, USA. Email: [email protected] 4 State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China. Email: [email protected] The Global boundary Stratotype Section and Point correlated with precision through all major Cambrian regions. (GSSP) for the base of the Drumian Stage (Cambrian Among the methods that should be considered in the selection of a GSSP (Remane et al., 1996), biostratigraphic, chemostratigraphic, Series 3) is defined at the base of a limestone (cal- paleogeographic, facies-relationship, and sequence-stratigraphic cisiltite) layer 62 m above the base of the Wheeler For- information is available (e.g., Randolph, 1973; White, 1973; McGee, mation in the Stratotype Ridge section, Drum Moun- 1978; Dommer, 1980; Grannis, 1982; Robison, 1982, 1999; Rowell et al. 1982; Rees 1986; Langenburg et al., 2002a, 2002b; Babcock et tains, Utah, USA. The GSSP level contains the lowest al., 2004; Zhu et al., 2006); that information is summarized here. occurrence of the cosmopolitan agnostoid trilobite Pty- Voting members of the International Subcommission on Cam- chagnostus atavus (base of the P. -
Early and Middle Cambrian Trilobites from Antarctica
Early and Middle Cambrian Trilobites From Antarctica GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Early and Middle Cambrian Trilobites From Antarctica By ALLISON R. PALMER and COLIN G. GATEHOUSE CONTRIBUTIONS TO THE GEOLOGY OF ANTARCTICA GEOLOGICAL SURVEY PROFESSIONAL PAPER 456-D Bio stratigraphy and regional significance of nine trilobite faunules from Antarctic outcrops and moraines; 28 species representing 21 genera are described UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1972 UNITED STATES DEPARTMENT OF THE INTERIOR ROGERS C. B. MORTON, Secretary GEOLOGICAL SURVEY V. E. McKelvey, Director Library of Congress catalog-card No. 73-190734 For sale by the Superintendent of Documents, U.S. Government Printing Office Washington, D.C. 20402 - Price 70 cents (paper cover) Stock Number 2401-2071 CONTENTS Page Page Abstract_ _ ________________________ Dl Physical stratigraphy______________________________ D6 I&troduction. _______________________ 1 Regional correlation within Antarctica ________________ 7 Biostratigraphy _____________________ 3 Systematic paleontology._____-_______-____-_-_-----_ 9 Early Cambrian faunules.________ 4 Summary of classification of Antarctic Early and Australaspis magnus faunule_ 4 Chorbusulina wilkesi faunule _ _ 5 Middle Cambrian trilobites. ___________________ 9 Chorbusulina subdita faunule _ _ 5 Agnostida__ _ _________-____-_--____-----__---_ 9 Early Middle Cambrian f aunules __ 5 Redlichiida. __-_--------------------------_---- 12 Xystridura mutilinia faunule- _ 5 Corynexochida._________--________-_-_---_----_ -
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900
An Appraisal of the Great Basin Middle Cambrian Trilobites Described Before 1900 By ALLISON R. PALMER A SHORTER CONTRIBUTION TO GENERAL GEOLOGY GEOLOGICAL SURVEY PROFESSIONAL PAPER 264-D Of the 2ty species described prior to I(?OO, 2/ are redescribed and 2C} refigured, and a new name is proposedfor I species UNITED STATES GOVERNMENT PRINTING OFFICE, WASHINGTON : 1954 UNITED STATES DEPARTMENT OF THE INTERIOR Douglas McKay, Secretary GEOLOGICAL SURVEY W. E. Wrather, Director For sale by the Superintendent of Documents, U. S. Government Printing Office Washington 25, D. C. - Price $1 (paper cover) CONTENTS Page Abstract..__________________________________ 55 Introduction ________________________________ 55 Original and present taxonomic names of species. 57 Stratigraphic distribution of species ____________ 57 Collection localities._________________________ 58 Systematic descriptions.______________________ 59 Literature cited____________________________ 82 Index __-_-__-__---_--______________________ 85 ILLUSTRATIONS [Plates 13-17 follow page 86] PLATE 13. Agnostidae and Dolichometopidae 14. Dorypygidae 15. Oryctocephalidae, Dorypygidae, Zacanthoididae, and Ptychoparioidea 16. Ptychoparioidea 17. Ptychoparioidea FIGUBE 3. Index map showing collecting localities____________________________ . Page 56 in A SHORTER CONTRIBUTION TO GENERAL GEOLOGY AN APPRAISAL OF THE GREAT BASIN MIDDLE CAMBRIAN TRILOBITES DESCRIBED BEFORE 1900 By ALLISON R. PALMER ABSTRACT the species and changes in their generic assignments All 29 species of Middle Cambrian trilobites -
The Anatomy, Affinity, and Phylogenetic Significance of Markuelia
EVOLUTION & DEVELOPMENT 7:5, 468–482 (2005) The anatomy, affinity, and phylogenetic significance of Markuelia Xi-ping Dong,a,Ã Philip C. J. Donoghue,b,Ã John A. Cunningham,b,1 Jian-bo Liu,a andHongChengc aDepartment of Earth and Space Sciences, Peking University, Beijing 100871, China bDepartment of Earth Sciences, University of Bristol, Wills Memorial Building, Queen’s Road, Bristol BS8 1RJ, UK cCollege of Life Sciences, Peking University, Beijing 100871, China ÃAuthors for correspondence (email: [email protected], [email protected]) 1Present address: Department of Earth and Ocean Sciences, University of Liverpool, 4 Brownlow Street, Liverpool L69 3GP, UK. SUMMARY The fossil record provides a paucity of data on analyses have hitherto suggested assignment to stem- the development of extinct organisms, particularly for their Scalidophora (phyla Kinorhyncha, Loricifera, Priapulida). We embryology. The recovery of fossilized embryos heralds new test this assumption with additional data and through the insight into the evolution of development but advances are inclusion of additional taxa. The available evidence supports limited by an almost complete absence of phylogenetic stem-Scalidophora affinity, leading to the conclusion that sca- constraint. Markuelia is an exception to this, known from lidophorans, cyclonerualians, and ecdysozoans are primitive cleavage and pre-hatchling stages as a vermiform and direct developers, and the likelihood that scalidophorans are profusely annulated direct-developing bilaterian with terminal primitively metameric. circumoral and posterior radial arrays of spines. Phylogenetic INTRODUCTION et al. 2004b). Very early cleavage-stage embryos of presumed metazoans and, possibly, bilaterian metazoans, have been re- The fossil record is largely a record of adult life and, thus, covered from the late Neoproterozoic (Xiao et al. -
Terreneuvian Orthothecid (Hyolitha) Digestive Tracts from Northern Montagne Noire, France; Taphonomic, Ontogenetic and Phylogenetic Implications
Terreneuvian Orthothecid (Hyolitha) Digestive Tracts from Northern Montagne Noire, France; Taphonomic, Ontogenetic and Phylogenetic Implications Le´a Devaere1*,Se´bastien Clausen1, J. Javier A´ lvaro2, John S. Peel3, Daniel Vachard1 1 UMR 8217 Ge´osyste`mes CNRS – Universite´ Lille 1Villeneuve d’Ascq, France, 2 Centro de Astrobiologı´a, Instituto Nacional de Te´cnica Aeroespacial, Consejo Superior de Investigaciones Cientı´ficas, Torrejo´n de Ardoz, Spain, 3 Department of Earth Sciences (Palaeobiology), Uppsala University, Uppsala, Sweden Abstract More than 285 specimens of Conotheca subcurvata with three-dimensionally preserved digestive tracts were recovered from the Terreneuvian (early Cambrian) Heraultia Limestone of the northern Montagne Noire, southern France. They represent one of the oldest occurrences of such preserved guts. The newly discovered operculum of some complete specimens provides additional data allowing emendation of the species diagnosis. Infestation of the U-shaped digestive tracts by smooth uniseriate, branching to anastomosing filaments along with isolated botryoidal coccoids attests to their early, microbially mediated phosphatisation. Apart from taphonomic deformation, C. subcurvata exhibits three different configurations of the digestive tract: (1) anal tube and gut parallel, straight to slightly undulating; (2) anal tube straight and loosely folded gut; and (3) anal tube straight and gut straight with local zigzag folds. The arrangement of the digestive tracts and its correlation with the mean apertural diameter of the specimens are interpreted as ontogenetically dependent. The simple U-shaped gut, usually considered as characteristic of the Hyolithida, developed in earlier stages of C. subcurvata, whereas the more complex orthothecid type-3 only appears in largest specimens. This growth pattern suggests a distinct phylogenetic relationship between these two hyolith orders through heterochronic processes. -
Paleontological Contributions
THE UNIVERSITY OF KANSAS PALEONTOLOGICAL CONTRIBUTIONS July 24, 1984 Paper 111 EXCEPTIONALLY PRESERVED NONTRILOBITE ARTHROPODS AND ANOMALOCARIS FROM THE MIDDLE CAMBRIAN OF UTAH' D. E. G. BRIGGS and R. A. ROBISON Department of Geology, Goldsmiths' College, University of London, Creek Road, London SE8 3BU, and Department of Geology, University of Kansas, Lawrence, Kansas 66045 Abstract—For the first time arthropods with preserved soft parts and appendages are recorded from Middle Cambrian strata in Utah. Occurrences of four nontrilobite taxa are described, including Branchiocaris pretiosa (Resser) and Emeraldella? sp. from the Marjum Formation, Sidneyia? sp. from the Wheeler Formation, and Leanchoilia? hanceyi, n. sp., from the Spence Shale. A small specimen of the giant predator Anomalocaris nathorsti (Walcott) also is described from the Marjum Formation. These occurrences extend upward the observed stratigraphie ranges of Anomalocaris, Branchiocaris, and questionably Emeraldella and Sidneyia. Emeraldella, Leanchoilia, and Sidneyia hitherto have been recorded from only the Stephen Formation in British Columbia. Further evaluation indicates that Dicerocaris opisthoeces Robison and Rich- ards, 1981, is a junior synonym of Pseudoarctolepis sharpi Brooks and Caster, 1956. DURING RECENT years, intensive collecting has 1983). Although providing little new morpho- produced rare but diverse, soft-bodied or scler- logic data, the Utah specimens are important otized Middle Cambrian fossils from several because of new information they provide about -
10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea
No. 1.] 43 10. Brief Notes on the Eodiscids 11, Phylogeny of the Dawsonidea. By Teiichi KoBAYASHI. (Comm. by H. YABE,M.I.A., Jan. 12, 1943.) Microdiscus has long been in common use for eodiscids, but as pointed out by Barrande in 1881, Microdiscus quadricostatus Emmons which is the type species of the genus, was a larval form of Trinucleus which was 'found in association with Ordovician graptolites and pele- cypods. Because Microdiscus Emmons, 1885, and the Microdiscidae Koken or Coquin, 196, lost their standing, Raymond adopted Eodiscus Hartt in place of Microdiscus and established Eodiscidae to include, beside the genus, two new ones, Goniodiscus and Weymouthia. While the three genera were all blind, Delgado described eodiscids with eyes in 1904 and Vogdes founded Delgadoia on Microdiscus caudatus Delgado in 1917. One year prior to this, Walcott had described proparian Pagetia as a member of the Eodiscidae, which became a subject of debate among trilobite students. Poulsen mentioned in 1927 that the sole reason for the resemblance of Pagetia to hypoparian eodiscids is adaptative convergence. This opinion is intenable because it is now known in Pagetia, Delgadoia or Al emtejoia that the two forms of each genus which are almost identical except for the presence or the absence of eyes and facial sutures, are found at the same locality. Supporting Beecher's view on the migration of the facial suture, Raymond emphasized in 1917, that Pagetia shows the stage in which the suture just came up to the dorsal side from the ventral in the agnostidian stage, notwithstanding the fact that Dindstrom and others have proven that the agnostid has no suture on the ventral side of its cephalon. -
The Early Paleozoic World: Cambrian Period (544-459 MY)
The Early Paleozoic World: Cambrian Period (544-459 MY) Jarðsaga 1 -Saga Lífs og Lands – Ólafur Ingólfsson The Cambrian Period: Precambrian Super- continent has broken up and life explodes... • Sea level rose throughout the Cambrian – very widespread deposition of marine sediments • Rodina Supercontinent broken up • Skeletal animals appear in the fossil record • Predation becomes a way-of-life • Trilobates appear on the scene, and develop rapidly • A major diversification of large animals • Trilobate mass extinctions towards the end of the period Cambrian key-sites Cambrian sediments occur on all Earths continents Cambrian was named by the 19th-century English geologist Adam Sedgwick, who first studied the great sequence of rocks characteristic of the period near Cambria, Wales. Early Cambrian continental configuration Rodina had broken up by early Cambrian, and Laurentia, Siberia and Baltica were separated from “Proto-Gondwana” Early Cambrian climate Early Cambrian climate was warm, but not too hot, and there is no evidence of any glacial ice at the poles. Transgression of sea onto continental cratons! http://www.scotese.com/ What caused the Cambrian transgression? What can cause globally rising sea-levels? 1. A decrease in ocean basin volume due to: • Formation of new mid-ocean ridges • Increased area of continents (rifting) • Accumulation of sediments on the sea-floor 2. An increase in water volume by melting of glaciers on land What can cause regionally/locally rising sea-levels? 1. Subsidence of land along a passive continental margin 2. Local/regional glacio-isostatic loading The vidence for a global Cambrian transgression 1. Wide-spread marine sediments on all cratons 2. -
New Palaeoscolecidan Worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale
New palaeoscolecidan worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale SIMON CONWAY MORRIS and JOHN S. PEEL Conway Morris, S. and Peel, J.S. 2010. New palaeoscolecidan worms from the Lower Cambrian: Sirius Passet, Latham Shale and Kinzers Shale. Acta Palaeontologica Polonica 55 (1): 141–156. Palaeoscolecidan worms are an important component of many Lower Palaeozoic marine assemblages, with notable oc− currences in a number of Burgess Shale−type Fossil−Lagerstätten. In addition to material from the lower Cambrian Kinzers Formation and Latham Shale, we also describe two new palaeoscolecidan taxa from the lower Cambrian Sirius Passet Fossil−Lagerstätte of North Greenland: Chalazoscolex pharkus gen. et sp. nov and Xystoscolex boreogyrus gen. et sp. nov. These palaeoscolecidans appear to be the oldest known (Cambrian Series 2, Stage 3) soft−bodied examples, being somewhat older than the diverse assemblages from the Chengjiang Fossil−Lagerstätte of China. In the Sirius Passet taxa the body is composed of a spinose introvert (or proboscis), trunk with ornamentation that includes regions bearing cuticu− lar ridges and sclerites, and a caudal zone with prominent circles of sclerites. The taxa are evidently quite closely related; generic differentiation is based on degree of trunk ornamentation, details of introvert structure and nature of the caudal re− gion. The worms were probably infaunal or semi−epifaunal; gut contents suggest that at least X. boreogyrus may have preyed on the arthropod Isoxys. Comparison with other palaeoscolecidans is relatively straightforward in terms of compa− rable examples in other Burgess Shale−type occurrences, but is much more tenuous with respect to the important record of isolated sclerites.