Contributions ro a MAI{TIAL OF PALAEARCTIC DIPTERA (with special reference to flies of economic importance) Volume 3 Higher Brachycera Låszl6 Prpp and B6la Darvas (Editors) Published by Science Herald, Budapest l.ggg Issued November 30, 1,998 Original illustrations by Albert Szappanos (Kecskemdt) Jånos Pål (Budapest) and the authors Cover design: Jånos Lengyel (Budapest) Illustrations editor: Zsuzsanm E. Prpp (Budapest) Authorised distributor: E. §ø. Classey Ltd Oxford House l\'larlborough Street Faringdon, Oxfordshire SN7 TP, U.K. Price subject to change without notice ISBN 963 04 8835 1, (Series) ISBN 963 04 8 83 8 8 (Volume 3) O Foundation for the Publicity of the Hungarian Science Baross u. 13, Budapest, PO Box 137, H-1431, Hungary Åll rights reserved. No part of this book mav be reproduced in any form or by any means' particilarly involving elictronic processing, rvithout written permission from the publisher. Educarional use is permirted for universiry and high school courses. Type-setting: PARS Ltd, Budapest, Hungary Printing: Dabas-Jegyzer Kft, Dirbas, Hungary Cover figures: Chrysotoxum elegans Loew and Neomyia cornicina (Fabricius), del. Jånos Pål Contents 3.0. Introduction - L. Papp and B. Darvas 7 3.L. Family Lonchopteridae - M. Bartåk 13 3.2. Family Opetiidae - P. J. Chandler 17 3.3. Family Platypezidae - P. J. Chandler and A. I. Shatalkin 27 3.4. Family Phoridae - R. H. L. Disney 51 3.5. Family Syrphidae - F. C. Thompson and G. Rotheray 81 3.6. Family Pipunculidae - M. Kozånek, M. De Meyer and A. Albrecht 1,4L 3.7. Family Cypselosomatidae - D. K. McAlpine 151 3.g. Family Pseudopomyzidae - D. K. McAlpine and A. I. shatalkin 155 3.9. Family Tanypezidae - J. RohåEek 165 3.10. Family Strongylophthalmyiidae - M. Iwasa 173 3.1,1. Family Psilidae - M. Iwasa 177 3.72. Family Otitidae - L. Greve 185 3.13. Family Platystomatidae - D. K. McAlpine L93 3.74. Family Pallopteridae - B. Merz 201. 3.15. Family Carnidae - L. PaPP 211, 3.16. Family Clusiidae - M. Sasakawa 21,9 3.17. Family Acartophthalmidae - L. Papp and A. L. Ozerov 227 3.18. Family Odiniidae - L. PaPP 233 3.19. Family Tethinidae - L. Munari 243 3.20. Family Canacidae - §f. N. Mathis 251, 3.21. Family Opomyzidae - E. Brunel 259 3.22. Family Anthomyzidae -J. Rohådek 267 3.23. Family Aulacigastridae - L. Papp 279 3.24. Family Periscelididae - §0. N. Mathis and L. Papp 285 3.25. Family Asteiidae - L. Papp 295 3.26. Family Xenasteiidae - L. Papp 305 3.27. Family Nannodastiidae - L. Papp and'W. N. Mathis 309 3.28. Family Milichiidae - L. Papp and T. A. §Theeler 315 3.29. Family Braulidae - L. Papp 325 3.30. Family Teratomyzidae - D. K. McAlpine 331 3.31. Family Coelopidae - D. K. McAlpine 335 3.32. Family Helcomyzidae - D. K. McAlpine 341 3.33. Family Heterocheilidae - D. K. McAlpine 345 3.34. Family Dryomyzidae - A. L. Ozerov 349 3.35. Family Sciomyzidae - R. Rozko§nyi 357 3.36. Family Phaeomyiidae - R. Rozko§nf 377 3.37. Family Lauxaniidae - L. Papp and A. t. Shatalkin 383 3.38. Family Celyphidae - L. Papp 401 3.39. Family Cremifaniidae - L. Papp 409 3.40. Family Chamaemyiidae - I. F. G. Mclean 41,5 3.41. Families of Heleomyzoidea - L. Papp 425 3.42. Family Chyromyidae - T. A. 'Wheeler 457 3.43. Family Sphaeroceridae - J. Rohååek 453 3.44. Family Curtonotidae - L. Papp 497 3.45. Family Drosophilidae - G. Båchli 503 3.46. Family Campichoetidae - P. J. Chandler 515 3.47. Family Diastatidae - P. J. Chandler 523 3.48. Family Camillidae - L. Prpp 531 3.49. Family Ephydridae - '§(/'. N. Mathis and T. Zattwarnicki 537 3.50. Family Anthomyiidae - M. Suwa and B. Darvas 571, 3.5 1.. Family Calliphoridae - K. Rognes 61,7 3.52. Family Sarcophagidae - T. Pape 649 3.53. Family Rhinophoridae - T. Pape 679 3.54. Family Tachinidae - H.-P. Tschorsnig and V. A. Richter 691, 3.55. Family Nycteribiidae - K. Hfirka 829 3.56. Family Streblidae - K. Hfrrka 839 Index - L. Peregovits and L. Prpp 849 3.51 . Family CALLIPHORIDAE Knut ROCNES Small to very large robust flies. (Fig. 1) of terior ocellus, but occasionally sexes more or less black or yellow ground-colour, with mostly sil- similar in this respect. Length 4-1'5 mm. very, yellowish or golden microtomentum in a variable paftern, though sometimes undusted; Adult. Head. usually much shorter than high non-metallic or metallic blue, bluish-black, blu- or deepr ås wide as thorax. Frons sloping, anten- ish-green, brassy or green. Some species all yel- nae usually inserted at middle of eye as seen in low. Male frons usually much narrower than in profile. Eyes bare, occasionally hairy. Facets oc- the female , at narrowest even narrower than an- casionally enlarged towards frons and in upper Fig. 51.1 . Lucilia sericata (Meigen), male. # 11, u u { å 618 Knut ROGNES f half. Lunula exposed, more or less shinin g, rarely groups siruated much higher than lower facial setose. Frontal vitta of varying width, withour margin (e.g. , Pollenia). small vibrissae above cruciate setae (except in some Mesembrinellinae large one ascending for varying distances ro- - a Neotropical group). Male frons usually only wards lunula. These ascendirrg setae very strong with inner and outer vertical setae, but occasion- in Catapicephala. Scape short, sometimes more ally with a lateroclinate (prevertical) seta, and or less erect. First flagellomere of varying length, even with proclinate orbital setae in some spe- from as long as pedicel (Fig. 2) to 7-B times as cies. Female frons usually with inner and outer long. fuista long- (Fig. 9) or short-plumose (Fig. verricals, a single lateroclinate orbital (preverti- 8), Lare (Figs i-l) or pectinate [is. I).. t# | f cal) and 2 proclinate orbitals, but in Villeneu- with or without a carina. Gena with genal dila- ' uiella with only inner or inner and outer vertical tion below eye. Gena very narrow (e.g., in Ben- setae retained (Fig. 3). Frontal setae reaching galia, Fig. 7) to very broad (Fig. 9), depending pedicel, these setae reduced in Villeneuuiella on size of eye. Occiput concave or convex, more (Figs 24). Fronto-orbital plate usually with nu- or less microtomentose with numerous black or merous setulae beside stronger frontal setae. white setulae, also in upper part, but in Rhiniinae Paraf.acial setose for longer or shorter distances the occiput lacks setulae in upper parr and is al- or bare; in one genus very strong setae in lower most always shining black. Paravertical setae pre- part (Eurychaeta) (Fig. 5). Lower facial margin sent, absent in Rhiniinae. Intrapostocular cilia vertical in profile or more or less projecting as often present benveen posterior eye margin and seen in profile; in Rhiniinae it is often strongly the row of postocular cilia. Proboscis short, but protruding (Fig. 4). Vibrissae srrong, in some lengthened e.g., in some Bellardia and Rhiniinae, I I J Figs 512-9. Hcads of Calliphoridae.2-3: Villeæutiella stackelberyf Grunin. anterior view: 2: male, 3: female. 4-9: male heads in profile: 4z Stomortina lunata (Fabriøts);5: Phormia regina (Meigen);6: Eurychaeta pølpalis (Robineau-Dewoidy); 7z Bengølia lyneborgt James (not Pdaearctic, paratype); 8: An§oneurø acerba (Meigen);9: Calliphorø uralensis Villeneuve (2-3 afær Grunin 1957,4-9 after Rognes l99ta). 3.51. Family CALLTPHORTDAE 619 and very stout palpus in Bengalia (Fig. 7). cla- Polleniinae (Neso dexia, Morinia, and also the vate, moderate to long, sornetimes flattened; Nearctic Melanodexia) and many Rhiniinae the often yellow. anterior lappet is a narrow fringe, whereas the Thorax. Humeral (- posrpronotal) chaeto- posterior one is a larger almost circular flap (Fig. taxy variable: usually 3 serae in a gently curved 15). In other groups the merarhoracic spiracle is line, or in the corner of a right-angled triangle. much larger, and particularly the anterior lapper often a single or several additional setae in front is greatly enlarged, sometimes much larger than of these. The number of humeral setae reduced the posterior lappet (Figs 1s, 18). In some to 2 in e.g. Dexopollenia , and Bengalia. usually Melanomyinae the spiracle is very much re- 34+34 acrostichal serae and 34 + 34 dor- duced. In some Polleniinae and in Catapicephala socentral setae, but often these series are re- there are a few or several yellow or black setae duced. Presutural intraalar setae (- posterior on the anrerior lappet. In Protophormia aticeps presutural intraalar seta) absent or present. No- (Zeæerstedt) the lappets of the rnerarhoracic spir- topleuron with 2 setae, occasionally with supple- acle (and the fringes bordering all other spir"ll.t mental notopleural setae in Protocalliphora and on body) are greatly thickened. In Chrysornyinae Trypocalliphora. Proepisrernal and proepimeral there is a row of long serulae along the an- setae usually present. Meron with a row of teroventral edge of the metathoracic spiracle pro- strong setae below metathoracic spiracle. (Fig. 15). subscutellum somerimes somewhat sternum, proepisternal depression, postalar wall swollen, but never as much as in the Tachinidae. and metasternal '§7ings. area setose or bare in various srem vein with a row of slender setae combinations in the various subfamilies. In- on dorsal surface of wing in Rhiniinae and Chry- frasquamal hairs always present. Coxopleural somyinae (also in Lucilia sinensis Aubertin and streak absent or present (Figs 15-18). Katatergite Pollenia atramentaria (Meigen), Figs zz-23). (supraspiracular convexiry) usually bare, but Costa setulose below ar least ro iunction with hu- with long thin erect hairs in Hemipyrellia. Sup- meral crossvein (Fig. 2{), but for varying dis- rasquamal ridge bare or serose, in Luciliinae a tances beyond this poini accordirrg ro subgroup [ø separate green metallic sclerite is present towards (Figs 23-25). Subcostal sclerite usually bare, but the hind end of the ridge which has a bundle of with setae in some Luciliinae and Polleniinae.