DOCSLIB.ORG

Comparative Study of Larvae of Tenebrionoidea (Coleoptera: Cucujiformia)

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
Comparative Study of Larvae of Tenebrionoidea (Coleoptera: Cucujiformia) Eur. J. Entomol. 102: 241–264, 2005 ISSN 1210-5759 Comparative study of larvae of Tenebrionoidea (Coleoptera: Cucujiformia) ROLF GEORG BEUTEL and FRANK FRIEDRICH Institut für Spezielle Zoologie und Evolutionsbiologie, FSU Jena, 07743 Jena, Germany; e-mails: [email protected], [email protected] Key words. Tenebrionoidea, larvae, morphology, character evolution, phylogeny Abstract. External and internal head structures and external structures of the thorax and abdomen of larval representatives of Melan- dryidae (Orchesia), Ulodidae (Meryx), Oedemeridae (Pseudolycus) and Pythidae (Pytho) are described. The obtained data were compared to characters of other tenebrionoid larvae and to larval characters of other representatives of Cucujiformia. Characters potentially relevant for phylogenetic reconstruction are listed and were analysed cladistically. The data set is characterised by a high degree of homoplasy and the resolution of the strict consensus trees of 2650 or 815 (second analysis) minimal length trees is low. The monophyly of Tenebrionoidea is supported by several larval autapomorphies, e.g. posteriorly diverging gula, anteriorly shifted posterior tentorial arms, asymmetric mandibles and the origin of several bundles of M. tentoriopharyngalis from the well-developed gular ridges. Several features of the larval head are plesiomorphic compared to the cleroid-cucujoid lineage. The interrelationships of most tenebrionoid families not belonging to the pythid-salpingid and anthicid-scraptiid groups were not resolved. Synchroidae were placed as sister group of a clade comprising these two lineages and Prostomidae. A sistergroup relationship between Trictenotomidae and Pythidae seems to be well supported and the monophyly of the anthicid-scraptiid lineage was also confirmed. Another potential clade comprises Prostomidae, Mycteridae and Boridae, and possibly Pyrochroidae (s.str.) and Inopeplinae. The monophyly of Salpingidae (incl. Othniinae and Inopelinae) and Pyrochroidae (incl. Pedilinae) was not supported. Many features such as the shape of the head and body, sutures and ridges of the head capsule, the endocarina, the mandibles, the maxillary apex, and also characters of the terminal abdominal apex are highly variable, even within families. Especially the families Tetratomidae, Melandryidae, Coly- diidae and Zopheridae show a high degree of variation in the larval stages. Several taxa appear isolated in terms of larval mor- phology within the families they are assigned to, e.g. Orchesia within Melandryidae, Sphindocis (Sphindocinae) within Ciidae, Calopus (Calopinae) within Oedemeridae and Penthe (Penthinae) within Tetratomidae. A broader spectrum of characters and a step- wise approach will be needed for a reliable clarification of the relationships within a very complex group like Tenebrionoidea. INTRODUCTION for most families were taken from the literature (e.g., With approximately 30,000 described species and 30 Lawrence, 1977, 1991; Lawrence & Spilman, 1991; Pol- currently recognised families (Lawrence & Newton, lock, 1995; Pollock & Lawrence, 1995; Pollock et al., 1995), Tenebrionoidea is a large and complex super- 2000; Selander, 1991; Young, 1991). A cladistic analysis family of cucujiform beetles. The composition of the was carried out even though it was apparent that the use group is largely agreed upon (Lawrence & Newton, of the larval characters alone would not be sufficient for a 1995). However, the monophyly is not well founded and good resolution within the complex superfamily. It is the interrelationships of the families are largely unre- likely that the inclusion of adult characters would have solved despite of a considerable number of phylogenetic improved the results of the phylogenetic analyses. How- investigations (e.g. Crowson, 1966; Abdullah, 1973, ever, this would have been clearly beyond the scope of 1974; Lawrence, 1977; Doyen & Lawrence, 1979; Pol- this study. lock, 1995; Pollock & Lawrence, 1995). A large number MATERIAL AND METHODS of studies on tenebrionoid larvae have been published. List of taxa examined (fixed in 70% ethanol unless otherwise However, most of them are not very detailed, and a thor- noted) ough description of internal structures is only available for a single species, Tenebrio molitor (Jösting, 1942). Tenebrionoidea, Melandryidae: Orchesia micans (Panzer, 1794) (Dubosq Brazil), Mycetoma sp., Xylita livida Sahlberg, Therefore, the main goal of this study is to improve the 1883 (determined material provided by S.A. ĝlipiĔski) knowledge of the larval morphology of Tenebrionoidea. Mordellidae: Curtimorda sp., Tomoxia bucephala Costa, 1854 For the detailed descriptions, four larvae representing dif- (prepupa) (determined material provided by S.A. ĝlipiĔski) ferent families and morphological types were chosen: Oedemeridae: Pseudolycus haemorrhoidalis (Fabricius, 1801) Orchesia micans (Panzer, 1794) (Melandryidae), Pseu- (determined by S.A. ĝlipiĔski) dolycus haemorrhoidalis (Fabricius, 1801) (Oedeme- Pyrochroidae: Pyrochroa sp. (FAE = formaldehyde + ethanol ridae), Meryx sp. (Ulodidae) and Pytho niger Kirby, 1837 + acetic acid) (identified by R.G. Beutel) (Pythidae). The findings are compared to structural fea- Pythidae: Pytho niger Kirby, 1837 (Kahle’s fluid) (deter- tures of other tenebrionoid larvae, and a list of characters mined material provided by D.A. Pollock) potentially useful for phylogenetic reconstruction and a Ulodidae: Meryx rugosa Latreille, 1807 (determined material provided by S.A. ĝlipiĔski) character state matrix are presented (Table 1). The data 241 242 TABLE 1. Character state matrix. 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 46 47 48 49 50 51 52 53 Derodontidae 0 0 0 0 0 0 0 1 0 0 0 0 0 1 0&1 0 0 0 0 0 0 0 0&1 0 0 0 0 – – 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Protocucujidae 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 1 1 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Mycetophagidae 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 1 ? ? 1 0 1 0 1 0 0 1 0 0 0 0 ? ? 0 0 0 0 0 0 0 0&1 0 0 0 0 0 0 0 0 0 0&1 0&1 0 0 Archeocrypticidae 0 0 0 0 0 0 1 1 0 0 0 0 0 0 0 1 ? ? 1 0 1 0 1 0 0 1 0 1 0 1 ? ? 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Pterogeniidae 0 0 0 0 0 0 0 0 0 1 0 0 1 0 0 1 ? ? 1 0 0 1 1 0 0 1 0 1 0 1 ? ? 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Ciidae 0 0 1 0&1 0 0 1 0 0 0 1 0 0 1 1 1 ? ? 0&1 0&1 0&1 0&1 1 0&1 0 0 0 – – 0 ? ? 0 0 0 0 0 0 0 0&1 0&1 0 0 0 0 0 0 0&1 0 0&1 0&1 0 0 Tetratomidae 0 0 0 0&1 0 0 0&1 0 0 0 0 0&1 0 0&1 0 1 ? ? 1 0&1 0 0&1 1 0 0 1 0&1 0 0 0&1 ? ? 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Melandryidae 0 0 0 0&1 0 0&1 0&1 0&1 0 0 0&1 0 0 0&1 0 1 1 0 0&1 0&1 0 0 1 0 0 1 0 0&1 0&1 0 1 1 0 0 0 0 0 0 0 0&1 0 0 0&1 0 0&1 0 0 0 0 0&1 0&1 0 0 Mordellidae 0 0 1 1 0 0 1 0 1 0 – – 0 0 0 1 0 0 0 1 – – 1 0 0 1 0 0 0 0 1 ? 0 0 1 0 0 0 0 0&1 0&1 0 0 0 0 0 0 0 0 0 1 0 0 Rhipiphoridae 0&1 0&1 0&1 0 0 0 0 0 0 0 1 0 0 ? 0 1 ? ? 0 1 – – 1 1 – – – – – 0 ? ? 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 1 Colydiidae 0 0 0 0&1 0&1 0&1 0 0&1 0 0 0&1 0 0 0&1 0 1 1 0 0 0&1 0 0&1 1 0 0 1 0&1 1 0 ? 1 ? 0 0 0 0 0 0 0 0&1 0&1 0 0&1 0 0 0 0 0&1 0 0 0 0 0 Monommatidae 0 0 0 0 1 0 0 1 0 0 0 0 0 0 0 1 ? ? 0 1 – – 1 0 0 1 1 1 1 0 ? ? 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 Prostomidae 1 1 0 0 0 0 0 1 0 0 0 0 0 1 0 1 1 1 1 0 1 1 1 0 1 1 0 0 1 0 1 1 0 1 0 0 1 1 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 Synchroidae 0 1 0 0 0 0 0 0 0 0 0 0 0 1 0 1 ? ? 1 0 0 1 1 0 1 1 0 0 1 1 ? ? 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 Zopheridae 0 0 0 0&1 0 0&1 0&1 0&1 0 0 0&1 0 0 0&1 0 1 ? ? 0&1 0 0 0&1 1 0 0 1 0&1 1 1 0&1 ? ? 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Ulodidae 0 0 0 1 0 0 1 1 0 0 0 0 0 0 0 1 1 0 1 1 – – 1 0 0 1 1 1 1 1 1 1 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 Perimylopidae 0 0 0 ? ? ? ? 1 0 0 0 0 0 1 0 ? ? ? 1 1 – – 1 0 0 1 0 0 1 ? ? ? 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 0 1 0 0 Chalcodryidae 0 0 0 0 0 0 1 0 0 0 1 0 1 1 0 1 ? ? ? 0 0 ? 1 0 0 1 0 0 0 ? ? ? ? 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0&1 0&1 0&1 0 Tenebrionidae 0 0 0 0 0 0 1 0 0 0 0&1 0 0 1 0&1 1 1 0 0&1 0 0&1 0&1 1 0 0 1 0&1 0&1 0 0 1 ? 0 0 0 0 0 0 0 0&1 0&1 0 0 0 0 0 0 0 0 0 1 0 0 Stenotrachelidae 0 0 0 0 0 0&1 0&1 0&1 0 0 0&1 0 0 0&1 0 1 ? ? 1 0 1 1 1 0 0 1 0&1 0&1 0 1 ? ? 0 0 0 0 0 0 0 0&1 0 0 0 0 0 0 0 0 0 0 0 0 0 Meloidae 0 0 0&1 0 0 0 0 0&1 0 0 0&1 0 0 0 0 1 ? ? 0 1 – – 1 1 – 1 0 0&1 0 0 ? ? 0 0 0 0 0 0 0 1 0 0 0 0 0 0 0 0 0 0 1 0 1 Oedemeridae 0 0 0 0 0 1 0&1 0 0&1 0 0&1 0 0 0&1 0 1 1 0 1 0 1 1 1 0 0 1 1 0 0 1 1 1 0 0 0 0 0 0 0 0&1 0 0 0&1 0 0 0&1 0 0 0 0 0&1 0&1 0 Mycteridae 1 1 0 0&1 0 0 0 1 0 0 0 0 0 0 0 1 ? ? 0&1 0&1 0 1 1 0 1 1 0&1 0 0 0 ? ? 0 0 0 1 1 1 0 0 0 0&1 0 1 0&1 0&1 0 0 0 0 0 1 0 Boridae 1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 1 ? ? 1 0 1 1 1 0 1 1 0 0 0 0 ? ? 1 0 0 1 1 1 0 0 0 0 0 1 0&1 0 0 0 0&1 0 0&1 0 0 Trictenotomidae 0 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 ? ? 1 0 1 1 1 0 ? 1 1 0&1 1 1 ? ? 0 0 0 0 1 0 1 0 0 0 0 0 0 0 1 0 0 0 1 0 0 Pythidae 0&1 1 0 0 0 0 0 1 0 0 0 0 0 1 1 1 1 0 1 0 1 1 1 0 1 1 1 0 1 1 1 1 1 0 0 0 1 0 1 0 0 0 0&1 0 0 0 1 0 0 0 0&1 0 0 Pyrochroidae 0&1 1 0 0 0 0 0 1 0 0 0 0 0 0 0 1 1 0 1 0 1 1 1 0 1 1 0&1 0&1 0 0&1 1 1 0&1 0 0 0&1 1 0&1 0 0 0 1 0 0&1 0 0&1 0&1 0 0 0 1 0 0 /Pyrochroidae* /1 /1 /1 /1 /1 /1 /1 /1 /0 /1 Pedilinae 0 1 0 0 0 0 0 1 0 0 0 0 0 ? ? 1 ? ? 1 0 1 1 1 0 1 1 0 0 0 0 ? ? 0 0 0 0 1 0 0 0 0 1 0 0 0 1 0 0 0 0 1 0 0 Othniinae 0 1 0 0 0 0 0 1 0 0 0 0 0 0 0 1 ? ? 1 0 1 1 1 0 1 1 1 1 1 1 ? ? 1 0 0 0 1 0 0 0 0 0 0 0 0 0 0&1 0 0 0 1 0 0 Salpingidae 0 1 0 0 0&1 0 0 1 0 0 0 0 0 0 0 1 ? ? 0&1 0&1
Load more

Recommended publications
  • Insect Cold Tolerance: How Many Kinds of Frozen?
    POINT OF VIEW Eur. J. Entomol. 96:157—164, 1999 ISSN 1210-5759 Insect cold tolerance: How many kinds of frozen? B rent J. SINCLAIR Department o f Zoology, University o f Otago, PO Box 56, Dunedin, New Zealand; e-mail: [email protected] Key words. Insect, cold hardiness, strategies, Freezing tolerance, Freeze intolerance Abstract. Insect cold tolerance mechanisms are often divided into freezing tolerance and freeze intolerance. This division has been criticised in recent years; Bale (1996) established five categories of cold tolerance. In Bale’s view, freezing tolerance is at the ex­ treme end of the spectrum o f cold tolerance, and represents insects which are most able to survive low temperatures. Data in the lit­ erature from 53 species o f freezing tolerant insects suggest that the freezing tolerance strategies o f these species are divisible into four groups according to supercooling point (SCP) and lower lethal temperature (LLT): (1) Partially Freezing Tolerant-species that survive a small proportion o f their body water converted into ice, (2) Moderately Freezing Tolerant-species die less than ten degrees below their SCP, (3) Strongly Freezing Tolerant-insects with LLTs 20 degrees or more below their SCP, and (4) Freezing Tolerant Species with Low Supercooling Points which freeze at very low temperatures, and can survive a few degrees below their SCP. The last 3 groups can survive the conversion of body water into ice to an equilibrium at sub-lethal environmental temperatures. Statistical analyses o f these groups are presented in this paper. However, the data set is small and biased, and there are many other aspects o f freezing tolerance, for example proportion o f body water frozen, and site o f ice nucleation, so these categories may have to be re­ vised in the future.
    [Show full text]
  • Thorne Moors :A Palaeoecological Study of A
    T...o"..e MO<J "S " "",Ae Oe COlOOIC'" S T<.OY OF A e"ONZE AGE slTE - .. "c euc~ , A"O a • n ,• THORNE MOORS :A PALAEOECOLOGICAL STUDY OF A BRONZE AGE SITE A contribution to the history of the British Insect fauna P.c. Buckland, Department of Geography, University of Birmingham. © Authors Copyright ISBN ~o. 0 7044 0359 5 List of Contents Page Introduction 3 Previous research 6 The archaeological evidence 10 The geological sequence 19 The samples 22 Table 1 : Insect remains from Thorne Moors 25 Environmental interpretation 41 Table 2 : Thorne Moors : Trackway site - pollen and spores from sediments beneath peat and from basal peat sample 42 Table 3 Tho~ne Moors Plants indicated by the insect record 51 Table 4 Thorne Moors pollen from upper four samples in Sphagnum peat (to current cutting surface) 64 Discussion : the flooding mechanism 65 The insect fauna : notes on particular species 73 Discussion : man, climate and the British insect fauna 134 Acknowledgements 156 Bibliography 157 List of Figures Frontispiece Pelta grossum from pupal chamber in small birch, Thorne Moors (1972). Age of specimen c. 2,500 B.P. 1. The Humberhead Levels, showing Thorne and Hatfield Moors and the principal rivers. 2 2. Thorne Moors the surface before peat extraction (1975). 5 3. Thorne Moors the same locality after peat cutting (1975). 5 4. Thorne Moors location of sites examined. 9 5. Thorne Moors plan of trackway (1972). 12 6. Thorne Moors trackway timbers exposed in new dyke section (1972) • 15 7. Thorne Moors the trackway and peat succession (1977).
    [Show full text]
  • Elytra Reduction May Affect the Evolution of Beetle Hind Wings
    Zoomorphology https://doi.org/10.1007/s00435-017-0388-1 ORIGINAL PAPER Elytra reduction may affect the evolution of beetle hind wings Jakub Goczał1 · Robert Rossa1 · Adam Tofilski2 Received: 21 July 2017 / Revised: 31 October 2017 / Accepted: 14 November 2017 © The Author(s) 2017. This article is an open access publication Abstract Beetles are one of the largest and most diverse groups of animals in the world. Conversion of forewings into hardened shields is perceived as a key adaptation that has greatly supported the evolutionary success of this taxa. Beetle elytra play an essential role: they minimize the influence of unfavorable external factors and protect insects against predators. Therefore, it is particularly interesting why some beetles have reduced their shields. This rare phenomenon is called brachelytry and its evolution and implications remain largely unexplored. In this paper, we focused on rare group of brachelytrous beetles with exposed hind wings. We have investigated whether the elytra loss in different beetle taxa is accompanied with the hind wing shape modification, and whether these changes are similar among unrelated beetle taxa. We found that hind wings shape differ markedly between related brachelytrous and macroelytrous beetles. Moreover, we revealed that modifications of hind wings have followed similar patterns and resulted in homoplasy in this trait among some unrelated groups of wing-exposed brachelytrous beetles. Our results suggest that elytra reduction may affect the evolution of beetle hind wings. Keywords Beetle · Elytra · Evolution · Wings · Homoplasy · Brachelytry Introduction same mechanism determines wing modification in all other insects, including beetles. However, recent studies have The Coleoptera order encompasses almost the quarter of all provided evidence that formation of elytra in beetles is less currently known animal species (Grimaldi and Engel 2005; affected by Hox gene than previously expected (Tomoyasu Hunt et al.
    [Show full text]
  • Coleoptera: Endomychidae: Leiestinae) with a Checklist and Nomenclatural Notes Regarding Fossil Endomychidae
    Zootaxa 3755 (4): 391–400 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3755.4.5 http://zoobank.org/urn:lsid:zoobank.org:pub:13446D49-76A1-4C12-975E-F59106AF4BD3 Glesirhanis bercioi, a new genus and species from Baltic amber (Coleoptera: Endomychidae: Leiestinae) with a checklist and nomenclatural notes regarding fossil Endomychidae FLOYD W. SHOCKLEY1& VITALY I. ALEKSEEV2 1Department of Entomology, National Museum of Natural History, Smithsonian Institution, P.O. Box 37012, MRC 165, Washington, DC 20013-7012, U.S.A. Email: [email protected] 2Department of Zootechny, Kaliningrad State Technical University, Sovetsky av. 1. 236000, Kaliningrad, Russia. E-mail: [email protected] Abstract A new genus and species of handsome fungus beetle, Glesirhanis bercioi gen. nov., sp. nov. (Coleoptera: Endomychidae: Leiestinae) is described from Baltic amber. The newly described genus is compared with all known extant and extinct genera of the subfamily. A key to the genera of Leiestinae including fossils and a checklist of fossil Endomychidae are provided. The status of two taxa previously placed in Endomychidae, Palaeoendomychus gymnus Zhang and Tetrameropsis mesozoica Kirejtshuk & Azar, is discussed, and a new status for the latter, elevating it to the family-level as Tetrameropseidae status nov., is proposed. Key words: new genus, new species, new status, Coleoptera, Endomychidae, Leiestinae, Baltic amber, Tertiary, Eocene, key, checklist, fossil Introduction Baltic amber (succinite) constitutes the largest known deposit of fossil plant resin and the richest repository of fossil insects of any age. Unfortunately, most references to Coleoptera in Baltic amber are only determined to family or generic levels.
    [Show full text]
  • Keys to Families of Beetles in America North of Mexico
    816 · Key to Families Keys to Families of Beetles in America North of Mexico by Michael A. Ivie hese keys are specifically designed for North American and, where possible, overly long lists of options, but when nec- taxa and may lead to incorrect identifications of many essary, I have erred on the side of directing the user to a correct Ttaxa from outside this region. They are aimed at the suc- identification. cessful family placement of all beetles in North America north of No key will work on all specimens because of abnormalities Mexico, and as such will not always be simple to use. A key to the of development, poor preservation, previously unknown spe- most common 50% of species in North America would be short cies, sexes or variation, or simple errors in characterization. Fur- and simple to use. However, after an initial learning period, most thermore, with more than 30,000 species to be considered, there coleopterists recognize those groups on sight, and never again are undoubtedly rare forms that escaped my notice and even key them out. It is the odd, the rare and the exceptional that make possibly some common and easily collected species with excep- a complex key necessary, and it is in its ability to correctly place tional characters that I overlooked. While this key should work those taxa that a key is eventually judged. Although these keys for at least 95% of specimens collected and 90% of North Ameri- build on many previous successful efforts, especially those of can species, the specialized collector who delves into unique habi- Crowson (1955), Arnett (1973) and Borror et al.
    [Show full text]
  • Coleoptera Tenebrionoidea) with Redescription of Falsopseudotomoxia Argyropleura (Franciscolo, 1942) N
    BOLL. SOC. ENTOMOL. ITAL., 145 (3): 103-115, ISSN 0373-3491 15 DICEMBRE 2013 Enrico ruZZiEr Taxonomic and faunistic notes on Italian Mordellidae (Coleoptera Tenebrionoidea) with redescription of Falsopseudotomoxia argyropleura (Franciscolo, 1942) n. comb. Riassunto: Note faunistiche e tassonomiche sui Mordellidi italiani con ridescrizione di Falsopseudotomoxia argyropleura (Franciscolo, 1942) n. comb. Nel presente lavoro sono forniti nuovi dati faunistici sui Mordellidae italiani ed è redatta una nuova checklist. Viene inoltre ridescritta Variimorda argyropleura e fornita una nuova combinazione tassonomica. Abstract: New faunistic records of italian Mordellidae and an updated checklist are given. Variimorda argyropleura is re-described and the species is assigned to the genus Falsopseudotomoxia. Key words: Coleoptera; Tenebrionoidea; Mordellidae; faunistic. iNTroduCTioN species whose status was in doubt. in this paper the Mordellidae is an extremely complex and ho- new status of Falsopseudotomoxia argyropleura mogeneous beetle family where a secure identifica- (Franciscolo, 1942) will be explained and an updated tion at species level is not possible without a check list of italian Mordellidae will be given. combination of genital morphology, external charac- ters (such as ridges on hind tibiae and tarsi, colour CHECK LisT oF iTALiAN MordELLidAE of the hairs on the elytra) and morphometric analysis. (* status not clear; [?] doubtful presence) in particular, genera such as Mordella (Linnaeus, ErPC: Enrico ruzzier Personal Collection, Mirano 1758) and Mordellistena (A. Costa, 1854) require at- (Venezia). tention due to the richness of sibling species (K. Er- CBFV: Centro Nazionale per lo studio e la Conservazione misch, 1954; 1956; 1963; 1965b; 1969; 1977), often della Biodiversità Forestale Bosco Fontana, Verona. sympatric. Therefore, faunistic research requires FAPC: Fernando Angelini Personal Collection, Francavilla careful and precise study of all material available.
    [Show full text]
  • (Coleoptera) Caught in Traps Baited with Pheromones for Dendroctonus Rufi Pennis (Kirby) (Curculionidae: Scolytinae) in Lithuania
    EKOLOGIJA. 2010. Vol. 56. No. 1–2. P. 41–46 DOI: 10.2478/v10055-010-0006-8 © Lietuvos mokslų akademija, 2010 © Lietuvos mokslų akademijos leidykla, 2010 Beetles (Coleoptera) caught in traps baited with pheromones for Dendroctonus rufi pennis (Kirby) (Curculionidae: Scolytinae) in Lithuania Henrikas Ostrauskas1, 2*, Sticky traps baited with pheromones for Dendroctonus rufi pennis were set up in the Klaipėda port and at the Vaidotai railway station alongside temporary stored timbers and Romas Ferenca2, 3 in forests along roads in June–July 2000 (21 localities across the entire Lithuania); 111 bee- tle species and 6 genera were detected. Eight trophic groups of beetles were identifi ed, and 1 State Plant Protection Service, among them the largest number (38.7% of species detected and 28.5% of beetle speci- Sukilėlių 9a, LT-11351 Vilnius, mens) presented a decaying wood and mycetobiont beetle group. Most frequent beetles Lithuania were Dasytes plumbeus (Dasytidae), Sciodrepoides watsoni (Leiodidae) and Polygraphus poligraphus (Curculionidae). Scolytinae were represented by 5 species and 83 beetle speci- 2 Nature Research Centre, mens, No D. rufi pennis was trapped. Rhacopus sahlbergi (Eucnemidae) and Anobium niti- Akademijos 2, dum (Anobiidae) beetles were caught in two localities, and the species were ascertained as LT-08412 Vilnius, Lithuania new for the Lithuanian fauna. Th ere was detected 71 new localities with the occurence of 54 beetle species rare for Lithuania. 3 Kaunas T. Ivanauskas Zoological Museum, Key words: bark beetles, sticky traps, rare Lithuanian species, new fauna species Laisvės al. 106, LT-44253 Kaunas, Lithuania INTRODUCTION risk of introducing the species via international trade.
    [Show full text]
  • The Beetle Fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and Distribution
    INSECTA MUNDI, Vol. 20, No. 3-4, September-December, 2006 165 The beetle fauna of Dominica, Lesser Antilles (Insecta: Coleoptera): Diversity and distribution Stewart B. Peck Department of Biology, Carleton University, 1125 Colonel By Drive, Ottawa, Ontario K1S 5B6, Canada stewart_peck@carleton. ca Abstract. The beetle fauna of the island of Dominica is summarized. It is presently known to contain 269 genera, and 361 species (in 42 families), of which 347 are named at a species level. Of these, 62 species are endemic to the island. The other naturally occurring species number 262, and another 23 species are of such wide distribution that they have probably been accidentally introduced and distributed, at least in part, by human activities. Undoubtedly, the actual numbers of species on Dominica are many times higher than now reported. This highlights the poor level of knowledge of the beetles of Dominica and the Lesser Antilles in general. Of the species known to occur elsewhere, the largest numbers are shared with neighboring Guadeloupe (201), and then with South America (126), Puerto Rico (113), Cuba (107), and Mexico-Central America (108). The Antillean island chain probably represents the main avenue of natural overwater dispersal via intermediate stepping-stone islands. The distributional patterns of the species shared with Dominica and elsewhere in the Caribbean suggest stages in a dynamic taxon cycle of species origin, range expansion, distribution contraction, and re-speciation. Introduction windward (eastern) side (with an average of 250 mm of rain annually). Rainfall is heavy and varies season- The islands of the West Indies are increasingly ally, with the dry season from mid-January to mid- recognized as a hotspot for species biodiversity June and the rainy season from mid-June to mid- (Myers et al.
    [Show full text]
  • Newsletter Alaska Entomological Society
    Newsletter of the Alaska Entomological Society Volume 12, Issue 1, March 2019 In this issue: Some food items of introduced Alaska blackfish (Dallia pectoralis T. H. Bean, 1880) in Kenai, Alaska8 Announcements . .1 Two new records of mayflies (Ephemeroptera) Arthropods potentially associated with spruce from Alaska . 11 (Picea spp.) in Interior Alaska . .2 Changes in soil fungal communities in response to A second Alaska record for Polix coloradella (Wals- invasion by Lumbricus terrestris Linnaeus, 1758 ingham, 1888) (Lepidoptera: Gelechioidea: Oe- at Stormy Lake, Nikiski, Alaska . 12 cophoridae), the “Skunk Moth” . .5 Review of the twelfth annual meeting . 19 Announcements New research to assess the risk of ticks tat suitability and probabilistic establishment model to dis- cover the climatic limits and probability of tick survival and tick-borne pathogens in Alaska in Alaska. For more information on ticks in Alaska and to learn how you can Submit-A-Tick, please visit: https: The geographic range of many tick species has expanded //dec.alaska.gov/eh/vet/ticks (website is in develop- substantially due to changes in climate, land use, and an- ment) or contact Dr. Micah Hahn ([email protected]). imal and human movement. With Alaska trending to- wards longer summers and milder winters, there is grow- ing concern about ticks surviving further north. Recent th passive surveillance efforts in Alaska have revealed that 69 Western Forest Insect Work Confer- non-native ticks—some with significant medical and vet- ence erinary importance—are present in the state. There is a new collaborative effort between the University of Alaska, The 69th Western Forest Insect Work Conference will the Alaska Department of Fish and Game, and the Of- be held April 22–25 2019 in Anchorage, Alaska at fice of the State Veterinarian to understand the risk of the Anchorage Marriott Downtown.
    [Show full text]
  • The Late Ross H. Arnett, Jr., Ph.D. Michael C. Thomas, Ph.D. Paul E
    VOLUME 2 AMERICAN BEETLES Polyphaga: Scarabaeoidea through Curculionoidea VOLUME 2 AMERICAN BEETLES Polyphaga: Scarabaeoidea through Curculionoidea Edited by the late Ross H. Arnett, Jr., Ph.D. Michael C. Thomas, Ph.D. Paul E. Skelley, Ph.D. and J. Howard Frank, D. Phil. CRC Press Boca Raton London New York Washington, D.C. COVER FIGURES: Center - Coccinellidae, Harmonia axyridus (Palles) [Photo by Fred J. Santana]. Outer rim, clockwise from top: Ripiphoridae, Macrosiagon cruentum (Germar) [by Fred J. Santana]; Meloidae, Lytta magister Horn [by Charles L. Bellamy]; Carabidae, Rhadine exilis (Barr and Lawrence) [by James C. Cokendolpher]; Melyridae, Malachius mirandus (LeConte) [by Max E. Badgley]; Lampyridae, Microphotus angustus LeConte [by Arthur V. Evans]. Library of Congress Cataloging-in-Publication Data American beetles / edited by Ross H. Arnett and Michael C. Thomas. p. cm. Contents: v. 1. Archostemata, Myxophaga, Adephaga, Polyphaga: Staphyliniformia. Includes bibliographical references (p.). ISBN 0-8493-0954-9 (alk. paper : v. 2)) 1. Beetles—North America. I. Arnett, Ross H. II. Thomas, M. C. (Michael Charles). 1948– QL581 .A43 2002 595.76¢097—dc21 00-050809 CIP This book contains information obtained from authentic and highly regarded sources. Reprinted material is quoted with permission, and sources are indicated. A wide variety of references are listed. Reasonable efforts have been made to publish reliable data and information, but the authors and the publisher cannot assume responsibility for the validity of all materials or for the consequences of their use. Neither this book nor any part may be reproduced or transmitted in any form or by any means, electronic or mechanical, including photocopying, microfilming, and recording, or by any information storage or retrieval system, without prior permission in writing from the publisher.
    [Show full text]
  • A03v24n3.Pdf
    Revista peruana de biología 24(3): 243 - 248 (2017) ISSN-L 1561-0837 A New Species of DODECACIUS (Coleoptera: Elateridae) from Madre de Dios, Peru doi: http://dx.doi.org/10.15381/rpb.v24i3.13903 Facultad de Ciencias Biológicas UNMSM TRABAJOS ORIGINALES A New Species of Dodecacius Schwarz (Coleoptera: Elateridae) from Madre de Dios, Peru Una nueva especie de Dodecacius Schwarz (Coleoptera: Elateridae) de Madre de Dios, Perú Paul J. Johnson Insect Biodiversity Lab, Box 2207A, South Dakota State University, Brookings, South Dakota 57007, U.S.A. Email: [email protected] Abstract Dodecacius Schwarz is reviewed, it includes two species known only from the eastern lower slopes of the Andes and adjacent Amazonia in southeastern Peru. Dodecacius paititi new species is described. Dodecacius testaceus Schwarz is treated as a new synonym of D. nigricollis Schwarz. Keywords: taxonomy; endemic; Andes; Amazonia; species discovery. Resumen El género Dodecacius Schwarz es revisado, incluye dos especies conocidas solamente de las laderas orientales bajas de los Andes y la Amazonia adyacente en el sureste de Perú. Se describe la nueva especie Dodecacius paititi y Dodecacius testaceus Schwarz es considerado como un nuevo sinónimo de D. nigricollis Schwarz. Palabras clave: taxonomía; endemismo; Andes; Amazonia; descubrimiento de especies. Publicación registrada en Zoobank/ZooBank article registered: urn:lsid:zoobank.org:pub:CF42CC9C-F496-4B4F-9C1A-FBB413A43E02 Acto nomenclatural/nomenclatural act: urn:lsid:zoobank.org:act:84A545F1-FAF8-42C1-83DA-C9D90CA0CA39 Citation: Johnson P.J. 2017. A New Species of Dodecacius Schwarz (Coleoptera: Elateridae) from Madre de Dios, Peru. Revista peruana de biología 24(3): 243 - 248 (octubre 2017).
    [Show full text]
  • The Evolution and Genomic Basis of Beetle Diversity
    The evolution and genomic basis of beetle diversity Duane D. McKennaa,b,1,2, Seunggwan Shina,b,2, Dirk Ahrensc, Michael Balked, Cristian Beza-Bezaa,b, Dave J. Clarkea,b, Alexander Donathe, Hermes E. Escalonae,f,g, Frank Friedrichh, Harald Letschi, Shanlin Liuj, David Maddisonk, Christoph Mayere, Bernhard Misofe, Peyton J. Murina, Oliver Niehuisg, Ralph S. Petersc, Lars Podsiadlowskie, l m l,n o f l Hans Pohl , Erin D. Scully , Evgeny V. Yan , Xin Zhou , Adam Slipinski , and Rolf G. Beutel aDepartment of Biological Sciences, University of Memphis, Memphis, TN 38152; bCenter for Biodiversity Research, University of Memphis, Memphis, TN 38152; cCenter for Taxonomy and Evolutionary Research, Arthropoda Department, Zoologisches Forschungsmuseum Alexander Koenig, 53113 Bonn, Germany; dBavarian State Collection of Zoology, Bavarian Natural History Collections, 81247 Munich, Germany; eCenter for Molecular Biodiversity Research, Zoological Research Museum Alexander Koenig, 53113 Bonn, Germany; fAustralian National Insect Collection, Commonwealth Scientific and Industrial Research Organisation, Canberra, ACT 2601, Australia; gDepartment of Evolutionary Biology and Ecology, Institute for Biology I (Zoology), University of Freiburg, 79104 Freiburg, Germany; hInstitute of Zoology, University of Hamburg, D-20146 Hamburg, Germany; iDepartment of Botany and Biodiversity Research, University of Wien, Wien 1030, Austria; jChina National GeneBank, BGI-Shenzhen, 518083 Guangdong, People’s Republic of China; kDepartment of Integrative Biology, Oregon State
    [Show full text]