Zoologische Mededelingen

September 2011

;e naturalis 85

Contents

De Grave, S. & C.HJ.M. Fransen Carideorum catalogus: the recent species of the dendrobranchiate, stenopodidean, procarididean and caridean (Crustácea: ) 195-

Cover: Periclimenneus schnitti Holthuis, 1951. Photo by Arthur Anker. Carideorum Catalogas: The Recent Species of the Dendrobranchiate, Stenopodidean, Procarididean and Caridean Shrimps (Crustácea: Decapoda)

S. De Grave & C.H.J.M. Fransen*

De Grave, S. & C.H.J.M. Fransen. Carideorum catalogus: the recent species of the dendrobranchíate, stenopodídean, procarididean and caridean shrimps (Crustácea: Decapoda). Zool. Med. Leiden 85(9): 30.ix.2011:195-589 figs 1-59.- ISSN 0024-0672, ISBN 978-90-6519-200-4. S. De Grave, Oxford University Mtiseum of Natural Hisíory, Parks Road, Oxford, OX1 3PW, United Kingdom ([email protected]). C.H.J.M. Fransen, Department of Marine Zoology, Netherlands Centre for Biodiversity Naturalis, P.O. Box 9517, 2300 RA Leiden, The Netherlands ([email protected]). * The order of authorship is purely alphabetical, with both authors having equally contributed to this work.

Key words: Crustácea; Decapoda; ; ; ; ; checklist. A checklist of recent species of dendrobranchiate, stenopodidean, procarididean and caridean shrimps including synoiiyms and type localities. Also listed are unavailable ñames, larval ñames, nomina dubia and nomina nuda. A complete list of references to original descriptions of taxa usted is províded.

Con ten ts

Introducción 196 The higher classiñcation of 196 Fossiltaxa 197 How many shrimp species are there? 197 Structure of the list 197 Notes on type locality 199 Notes on dating of some papers 200 Notes on author ñames 201 Notes on authorship of taxa 202 Notes on some papers of nomenclatorial significance 203 Notes on the spelling of some ñames 204 Notes on the gender of generic ñames and gender agreement in species ñames 204 Acknowledgements 205 Checklist 206 Suborder Dendrobranchiata 206 Suborder 248 Infraorder Procarididea 248 Infraorder Stenopodidea 249 Infraorder Caridea 254 Unavailable ñames 471 Penaeoidea Larvata 471 Penaeoidea Nomina dubia 471 196 De Grave & Fransen. Carideorum catalogáis (Crustácea: Decapoda). Zool. Med. Leiden 85 (2011)

Penaeoidea Nomina nuda 472 Sergestoidea Larvata 472 Sergestoidea Nomina dubia 472 Sergestoidea Nomina nuda 473 Candea Larvata 473 Caridea Nomina dubia 475 Caridea Nomina nada 476 References 478 Systematic Índex to currently valid famili.es and genera 585

Introduction

The higher classification of shrimp

Over the last decade or so, nuich has been written on the classification of Decapoda, fuelled by a surge in molecular phylogenetic studies, as well as cióse scrutiny of inter- nal and external morphological characteristics. As discussed by Fransen & De Grave (2009), such studies on shrimps are still somewhat "thin on the ground", at least com- pared to the more extensive work done on the Brachyura and . At a higher level in decapod classiñcation it has long been recognised that three distinct lineages of shrimps can be distinguished: Dendrobranchiata, Stenopodidea and Caridea, a system which has not been seriously challenged by recent studies. The internal classification of Dendrobr anchi ata and Stenopodidea alike has been stable for some time, with the only major addition being the family Macromaxilloca- rididae Alvarez, Iliffe & Villalobos (2006) to the Stenopodidea in recent years. A different picture has emerged for Caridea very recently with Bracken et al. (2009) and Chan et al. (2010), both drawing attention to the non-monophyletic status of certain superfainilies and families. Further, we are aware of work currently in progress (some by the authors of this compilation) corroborating the hypothesis that the current clas- sification of Caridea is unnatural, lines of study which vvill lead to the resurrection of certain family ñames as well as further refinemeiit to other families. As one of our objec- tives for the current effort was to link this compilation of species level informatíon with the earlier work by Chace (1992) for families and Holthuis (1993a) for genera, we have elected to largely follow the classification outlined by De Grave et al. (2009) which builds upon this earlier work. As such, it was deemed advisable to include the recently resurrected family Acanthephyridae Spence Bate, 1888 in the superfamily Oplopho- roidea, rather than in this catalogue to créate a new superfamily, which would perhaps be more congruent with the results in Chan et al. (2010). Although we follow herein the classification scheme of De Grave et al. (2009), two recent changes have been implemented. The clarification of the status of Galatheacaris abyssalis Vereshchaka, 1997a, as the megalopal stage of Eugonatonotus chacei Chan & Yu, 1991a, by De Grave et al. (2010) resulted in the removal of the family Galatheacarididae and superfamily Galatheacaridoidea in the current listing. Bracken et al. (2010) clarified the status of the family Procarididae, resulting in the recognición, of a fourth group of shrimp, Infraorder Procarididea.