Early Carnivorous Dinosaurs Crossed Continents 10 December 2009
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Tyrannosaurus Rex by Guy Belleranti
Name: ______________________________ Tyrannosaurus Rex By Guy Belleranti One of the most dangerous dinosaurs was the Tyrannosaurus rex. It looked like a huge lizard with sharp teeth. It lived over 60 million years ago. From nose to tail, T-rex was as long as a school bus. It was taller than a house. It weighed more than an airplane. T-rex’s head was as long as a kitchen table. T-rex was the biggest meat-eating dinosaur. It could eat hundreds of pounds of meat in one bite. Animals that eat meat have sharp teeth. T-rex had 60 of them! Some of the teeth were as big as bananas. When T-rex lost a tooth, it grew a new one. T-rex stood on two powerful legs. It also had two small arms. Its strong tail helped keep it from falling over. It might be fun to see a live Tyrannosaurus rex, but I wouldn’t want to meet one. Would you? Super Teacher Worksheets - www.superteacherworksheets.com Name: ______________________________ Tyrannosaurus Rex By Guy Belleranti 1. How many teeth did a Tyrannosaurus rex have? a. thirty b. sixteen c. sixty d. seventy 2. How long ago did Tyrannosaurus rex live? ________________________________________________________________ 3. What did Tyrannosaurus rex eat? a. leaves from tall trees b. other dinosaurs c. small insects d. people 4. A T-rex was as long as a ______________________________________. 5. A T-rex weighed as much as an _______________________________. 6. Which dinosaurs had sharp teeth? a. all dinosaurs b. dinosaurs that had tails c. dinosaurs that were big d. -
Fused and Vaulted Nasals of Tyrannosaurid Dinosaurs: Implications for Cranial Strength and Feeding Mechanics
Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics ERIC SNIVELY, DONALD M. HENDERSON, and DOUG S. PHILLIPS Snively, E., Henderson, D.M., and Phillips, D.S. 2006. Fused and vaulted nasals of tyrannosaurid dinosaurs: Implications for cranial strength and feeding mechanics. Acta Palaeontologica Polonica 51 (3): 435–454. Tyrannosaurid theropods display several unusual adaptations of the skulls and teeth. Their nasals are fused and vaulted, suggesting that these elements braced the cranium against high feeding forces. Exceptionally high strengths of maxillary teeth in Tyrannosaurus rex indicate that it could exert relatively greater feeding forces than other tyrannosaurids. Areas and second moments of area of the nasals, calculated from CT cross−sections, show higher nasal strengths for large tyrannosaurids than for Allosaurus fragilis. Cross−sectional geometry of theropod crania reveals high second moments of area in tyrannosaurids, with resulting high strengths in bending and torsion, when compared with the crania of similarly sized theropods. In tyrannosaurids trends of strength increase are positively allomeric and have similar allometric expo− nents, indicating correlated progression towards unusually high strengths of the feeding apparatus. Fused, arched nasals and broad crania of tyrannosaurids are consistent with deep bites that impacted bone and powerful lateral movements of the head for dismembering prey. Key words: Theropoda, Carnosauria, Tyrannosauridae, biomechanics, feeding mechanics, computer modeling, com− puted tomography. Eric Snively [[email protected]], Department of Biological Sciences, University of Calgary, 2500 University Drive NW, Calgary, Alberta T2N 1N4, Canada; Donald M. Henderson [[email protected]], Royal Tyrrell Museum of Palaeontology, Box 7500, Drumheller, Alberta T0J 0Y0, Canada; Doug S. -
Anomalously High Variation in Postnatal Development Is Ancestral for Dinosaurs but Lost in Birds
Anomalously high variation in postnatal development is ancestral for dinosaurs but lost in birds Christopher T. Griffina,1 and Sterling J. Nesbitta aDepartment of Geosciences, Virginia Polytechnic Institute and State University, Blacksburg, VA 24061 Edited by Neil H. Shubin, The University of Chicago, Chicago, IL, and approved November 3, 2016 (received for review August 19, 2016) Compared with all other living reptiles, birds grow extremely fast sequence analysis (OSA) (32) to reconstruct growth sequences of and possess unusually low levels of intraspecific variation during these early dinosaurs, two avian species (Branta canadensis and postnatal development. It is now clear that birds inherited their high Meleagris gallopavo), and a single crocodylian species (Alligator rates of growth from their dinosaurian ancestors, but the origin of mississippiensis), and demonstrate that the earliest dinosaurs the avian condition of low variation during development is poorly developed differently than living archosaurs. constrained. The most well-understood growth trajectories of later Mesozoic theropods (e.g., Tyrannosaurus, Allosaurus)showsimilarly Results low variation to birds, contrasting with higher variation in extant Our OSAs indicate that both C. bauri and M. rhodesiensis pos- crocodylians. Here, we show that deep within Dinosauria, among sessed a high level of intraspecific variation, both in sequence the earliest-diverging dinosaurs, anomalously high intraspecific var- polymorphism and in body size at different levels of morpho- iation is widespread but then is lost in more derived theropods. This logical maturity (Figs. 1 and 2). Analysis of the 27 ontogenetic style of development is ancestral for dinosaurs and their closest characters for C. bauri reconstructed 136 equally parsimonious relatives, and, surprisingly, this level of variation is far higher than developmental sequences (Fig. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
Rule Booklet
Dig for fossils, build skeletons, and attract the most visitors to your museum! TM SCAN FOR VIDEO RULES AND MORE! FOSSILCANYON.COM Dinosaurs of North America edimentary rock formations of western North America are famous for the fossilized remains of dinosaurs The rules are simple enough for young players, but and other animals from the Triassic, Jurassic, and serious players can benefit Cretaceous periods of the Mesozoic Era. Your objective from keeping track of the cards that is to dig up fossils, build complete skeletons, and display have appeared, reasoning about them in your museum to attract as many visitors as possible. probabilities and expected returns, and choosing between aggressive Watch your museum’s popularity grow using jigsaw-puzzle and conservative plays. scoring that turns the competition into a race! GAME CONTENTS TM 200,000300,000 160,000 VISITORS VISITORS PER YEAR 140,000 VISITORS PER YEAR 180,000 VISITORS PER YEAR 400,000 VISITORS PER YEAR Dig for fossils, build skeletons, and 340,000 VISITORS PER YEAR RD COLOR ELETONS CA GENUS PERIODDIET SK FOSSIL VISITORSPARTS 360,000 VISITORS PER YEAR PER YEAR attract the most visitors to your museum! VISITORS PER YEAR PER YEAR Tyrannosaurus K C 1 4 500,000 Brachiosaurus J H 1 3 400,000 ON YOUR TURN: TM SCAN FOR VIDEO Triceratops K H 1 3 380,000 RULES AND MORE! Allosaurus J C 2 Dig3 a first360,000 card. If it is a fossil, keep it hidden. FOSSILCANYON.COM Ankylosaurus K H 2 If it3 is an340,000 action card, perform the action. -
University of Birmingham a New Dinosaur With
University of Birmingham A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil Marsola, Julio; Bittencourt, Jonathas; Butler, Richard; Da Rosa, Atila; Sayao, Juliana; Langer, Max DOI: 10.1080/02724634.2018.1531878 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Marsola, J, Bittencourt, J, Butler, R, Da Rosa, A, Sayao, J & Langer, M 2019, 'A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil', Journal of Vertebrate Paleontology, vol. 38, no. 5, e1531878. https://doi.org/10.1080/02724634.2018.1531878 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 25/07/2018 This is the accepted manuscript for a forthcoming publication in Journal of Vertebrate Paleontology. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. -
A Re-Evaluation of the Enigmatic Dinosauriform Caseosaurus Crosbyensis from the Late Triassic of Texas, USA and Its Implications for Early Dinosaur Evolution
A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution MATTHEW G. BARON and MEGAN E. WILLIAMS Baron, M.G. and Williams, M.E. 2018. A re-evaluation of the enigmatic dinosauriform Caseosaurus crosbyensis from the Late Triassic of Texas, USA and its implications for early dinosaur evolution. Acta Palaeontologica Polonica 63 (1): 129–145. The holotype specimen of the Late Triassic dinosauriform Caseosaurus crosbyensis is redescribed and evaluated phylogenetically for the first time, providing new anatomical information and data on the earliest dinosaurs and their evolution within the dinosauromorph lineage. Historically, Caseosaurus crosbyensis has been considered to represent an early saurischian dinosaur, and often a herrerasaur. More recent work on Triassic dinosaurs has cast doubt over its supposed dinosaurian affinities and uncertainty about particular features in the holotype and only known specimen has led to the species being regarded as a dinosauriform of indeterminate position. Here, we present a new diagnosis for Caseosaurus crosbyensis and refer additional material to the taxon—a partial right ilium from Snyder Quarry. Our com- parisons and phylogenetic analyses suggest that Caseosaurus crosbyensis belongs in a clade with herrerasaurs and that this clade is the sister taxon of Dinosauria, rather than positioned within it. This result, along with other recent analyses of early dinosaurs, pulls apart what remains of the “traditional” group of dinosaurs collectively termed saurischians into a polyphyletic assemblage and implies that Dinosauria should be regarded as composed exclusively of Ornithoscelida (Ornithischia + Theropoda) and Sauropodomorpha. In addition, our analysis recovers the enigmatic European taxon Saltopus elginensis among herrerasaurs for the first time. -
A Dinosaur Track from New Jersey at the State Museum in Trenton
New Jersey Geological and Water Survey Information Circular What's in a Rock? A Dinosaur Track from New Jersey at the State Museum in Trenton Introduction a large dinosaur track (fig. 2) on the bottom. Most of the rock is sedimentary, sandstone from the 15,000-foot-thick Passaic A large, red rock in front of the New Jersey State Museum Formation. The bottom part is igneous, lava from the 525-foot- (NJSM) in Trenton (fig. 1) is more than just a rock. It has a thick Orange Mountain Basalt, which overspread the Passaic fascinating geological history. This three-ton slab, was excavated Formation. (The overspreading lava was originally at the top of from a construction site in Woodland Park, Passaic County. It the rock, but the rock is displayed upside down to showcase the was brought to Trenton in 2010 and placed upside down to show dinosaur footprint). The rock is about 200 million years old, from the Triassic footprints Period of geologic time. It formed in a rift valley, the Newark Passaic Formation Basin, when Africa, positioned adjacent to the mid-Atlantic states, began to pull eastward and North America began to pull westward contact to open the Atlantic Ocean. The pulling and stretching caused faults to move and the rift valley to subside along border faults including the Ramapo Fault of northeastern New Jersey, about 8 miles west of Woodland Park. Sediments from erosion of higher Collection site Orange Mountain Basalt top N Figure 1. Rock at the New Jersey State Museum. Photo by W. Kuehne Adhesion ripples DESCRIPTION OF MAP UNITS 0 1 2 mi Orange Mountain Basalt L 32 cm Jo (Lower Jurassic) 0 1 2 km W 25.4 cm contour interval 20 feet ^p Passaic Formation (Upper Triassic) Figure 3. -
Tyrannosaurus Rex.Pmd
North Dakota Stratigraphy Tyrannosaurus rex ROCK ROCK UNIT COLUMN PERIOD EPOCH AGES MILLIONS OF YEARS AGO Common Name: Holocene Oahe .01 Tyrant reptile king Coleharbor Pleistocene QUATERNARY Classification: 1.8 Pliocene Unnamed 5 Miocene Class: Reptilia 25 Arikaree Order: Saurischia Family: Tyrannosauridae Brule Oligocene 38 Tyrannosaurus rex shed tooth. Tooth collected in Morton South Heart Chadron Chalky Buttes County. Height of tooth is 64 mm. North Dakota State Fossil Camels Butte Eocene Golden Collection. 55 Valley Bear Den Description: Sentinel Butte Tyrannosaurus rex was one of the largest carnivorous (meat TERTIARY eating) dinosaurs and was one of the largest terrestrial carnivores yet known. The adults grew to lengths of 40 feet from the end of the tail to tip of the nose and weighed about 8 tons. When they Bullion Paleocene Creek stood on their hind legs they were up to about 20 feet tall. They had huge heads, about 5 feet long, and possessed large, Slope approximately 50, dagger-like teeth, some as large as bananas. Cannonball The teeth, which were serrated, could puncture bone and carve Ludlow through flesh of prey. Its back legs were long, heavily built, and 65 powerful with 3 clawed toes on each foot. Each foot was broad Hell Creek with three forward-pointing toes. Each toe ended in a sharply- curved talon. T. rex’s arms were very short and contained hands Fox Hills with only two, clawed fingers on each hand. Its tail was long, heavy, and held off the ground to act as a counterbalance. They ACEOUS could tear off as much as about 500 pounds of flesh at one time Pierre with their powerful jaws. -
Skulls of Tarbosaurus Bataar and Tyrannosaurus Rex Compared
Giant theropod dinosaurs from Asia and North America: Skulls of Tarbosaurus bataar and Tyrannosaurus rex compared Jørn H. Hurum and Karol Sabath Acta Palaeontologica Polonica 48 (2), 2003: 161-190 The skull of a newly prepared Tarbosaurus bataar is described bone by bone and compared with a disarticulated skull of Tyrannosaurus rex. Both Tarbosaurus bataar and Tyrannosaurus rex skulls are deep in lateral view. In dorsal view, the skull of T. rex is extremely broad posteriorly but narrows towards the snout; in Ta. bataar the skull is narrower (especially in its ventral part: the premaxilla, maxilla, jugal, and the quadrate complex), and the expansion of the posterior half of the skull is less abrupt. The slender snout of Ta. bataar is reminiscent of more primitive North American tyrannosaurids. The most obvious difference between T. rex and Ta. bataar is the doming of the nasal in Ta. bataar which is high between the lacrimals and is less attached to the other bones of the skull, than in most tyrannosaurids. This is because of a shift in the handling of the crushing bite in Ta. bataar . We propose a paleogeographically based division of the Tyrannosaurinae into the Asiatic forms (Tarbosaurus and possibly Alioramus) and North American forms (Daspletosaurus and Tyrannosaurus). The division is supported by differences in anatomy of the two groups: in Asiatic forms the nasal is excluded from the major series of bones participating in deflecting the impact in the upper jaw and the dentary-angular interlocking makes a more rigid lower jaw. Key words: Dinosauria, Theropoda, Tyrannosauridae, Tarbosaurus, Tyrannosaurus, skull, anatomy, Mongolia. -
Herrerasaurus
FACT SHEET: herrerasaurus NAME: HERRERASAURUS (huh-RARE-uh-SAW-rus), meaning: "Herrera's lizard" SIZE: 13 feet long (the size of a minivan), 460 pounds TYPE OF DIET: Carnivore (lizards, fish and smaller dinsoaurs) WHEN: Mid Triassic period (230 million years ago) WHERE: Argentina, South America Herrerasaurus is one of the oldest known dinosaurs. It lived about 230 million years ago. It was a theropod dinosaur, meaning that it ate meat and ran on its back legs. It would also walk on all four legs when it was looking for food on the ground. It was probably a fast runner; we know this by studying its trackways, or footprints. Herrerasaurus grew to be 13 feet long, which is about the size of a minivan. It was about four feet high and weighed about 460 pounds. This dinosaur was a carnivore, meaning that it ate meat. It would eat lizards and fish and other smaller dinosaurs. It was probably a deadly hunter. Its teeth were sharp and had jagged edges, like small saws. Herrerasaurus had an unusual joint in its jaw. This allowed it to open its mouth very wide. Herrerasaurus had hands with five fingers and a thumb, like we have. But its fifth finger was probably too small to be seen. Its hands were very useful for grabbing things. It had short arms and sharp, curved claws on its fingers. We believe that it hunted alone, since its bones have not been found together with other Herrerasaurus skeletons. The fossilized bones of this dinosaur were found by a goat farmer named Don Victorino Herrera.