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Contributions to the Knowledge of the Malachius Group of Soft-Winged Flower Beetles (Coleoptera, Malachiidae) of the Fauna of Russia and the Adjacent Countries

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Contributions to the Knowledge of the Malachius Group of Soft-Winged Flower Beetles (Coleoptera, Malachiidae) of the Fauna of Russia and the Adjacent Countries BULLETI1 DE L'li STITUT ROYAL DES SCIE 1CES NATURELL ES DE BELG IQUE ENTOi'"IOLOGIE. 70: 199-218.2000 BULLETI! VAN HET KON INKLIJK BELGISCH I STITUUT VOOR NATUU RWETE1 SCI-lAPPEN ENTOMOLOG1E. 70:' 199-21 8.2000 Contributions to the knowledge of the Malachius group of soft-winged flower beetles (Coleoptera, Malachiidae) of the fauna of Russia and the adjacent countries. Pati II by Sergei E. TSHERNYSHEV Abstract In this work two ne>vv species of the genus Clanoptilus are described. A special interest is atteched to a group of 2 new species: Clanoptilus senylia sp. n. (Uzbekistan: Hissar, Kyrgyz­ species with very similar characters. The shape and sculp­ stan, Alaj) and C. pseudo!utescens sp. n. (Talysh, Georgia) are de­ scribed. A rare species, Clanoptilus tedschenensis Evers, 1985 is re­ ture of the pro no tum and elytra, the coloration and shape described and fi gured. A new status, C/anoptilus (1-/ypopti/w) foveal us of antennae and heads, allows one to regard these species (MEDVEDEV, 1964) stat.n. is proposed. A key to a new Clanoptilus as belonging to a new group, Clanoptilus spinosus . .spinosu s group of species is provided. A new genus, C!anomalachius gen.n. is described ~ov ith the review of species: C/anomalachius dolini Amongst them a special male character developed differ­ (WITTMER, 199 1) comb.n., C. insignicomis (P IC, 1909) comb.n., C. ently: from a very small depression without an appendage kiesel/lvetleri (Sols ky, 188 1) comb.n., C. resecatus (WITTMER, 1979) in C. .foveatus to a deep impression with large and com­ comb.n., C. debrosi (WITTMER, 1979) stat. et comb.n., C. kiil111efti (I-IlCKER, 1959) comb.n. A generic key for the Malachius-group is plicated appendage in C. spinosus. given. Illustrations and distributional maps of species are presented. The same situation with the development of special characters occurs in a group of species with very similar Key words: Coleoptera, Ma lachiidae, ivlalachius-group, C /anopti!us, general characters which neu1ge nus, new species, taxonomy. allows this group to be regarded as Malachius or C!anoptilus according to a scheme of EV ERS ( 1985). The very specific shape of the apical tergite in males, being complexly cu rved and emarginate Pe3JOMe shows a difference from all Malachius-group representa­ tives. For these species, a new genus, Clanomalachius, is B pa6oTe omiCbiBaJOTCJI 2 HOBbiX BH,Aa: Clanoptilus senylia sp.n. described. (Y36eKHCTaH: rHccap, KHprH3HJI : AJiaH) H c. pseudolutescens sp.n. (TaJibiiii, rpy3HJ1) H HOBbiH po.n Cfaii OIIIOiachius, gen.n., All materials were borrowed from, and housed in, the c o63opoM BXO.Allll.(HX a Hero BH.AOB: C!anomalachius dolini (WITT­ following museums and collections: MER, 199 1) comb.n., C. insignicomis (PIC, 1909) comb.n., C. kiesen­ - Institute of Biology and Pedology, Kyrgh wel/eri (SOLSKY, 188 1) comb.n., C. resecatus (WITTMER, 1979) ~BPK yz comb.n., C. debrosi (WITTMER, 1979) stat. ct comb.n., C. kiihne!ti National Academy of Sciences, Bishkek; (I-IlCKER, 1959) comb.n. TiepeOilHCaH pe,AKHH cpe,AHea3HaTCKHH BH,A JSNB - Royal Belgian Institute for Natural Science, Clanopti lus tedschenensis EVERS, 1985, .naHbi pHcyHKH reHHTaJIHH H Bri.issel; Cllel..lH<pli'!eCKHX CTpyKTyp CaMI..la. Ilpe.nnoJKeH HOBbiH CTaryc .[{RJI BH,Aa C/anoptilus (1-/ypopti/us) .foveatus (L. MEDVE DE V, 1964) stat.n. NHMB - Naturhi storisches Museum, Basel; }];aHa onpe.nenHTenbH3JI Ta6JIHI..la po.noa rpynnbi i\1/a/achius, NMP - Narodni Muzeum v Praze; BBO.z:lHTCJI rpynna BH.[{OB C/anoptilus s pinosus, c onpe,AenHTenbHOH SZMN - Siberian Zoological Museum, Institute of Ta6nHI..leH Bli.[{OB. }J;nll OllHCbiBaeMbiX T3KCOHOB npHBO.[{JITCJI pHCYHKH li ROKaJIHTeT. Animal Systematics a nd Ecology, Siberian Branch of the Russian Academy of Sciences, Novosibirsk; Kmo'ICBbiC CJIOBa: Coleoptera , Malachiidae, Malachius-rpynna, ZISP - Zoological Institute, the Russian Academy of C/cmnpti/us, HOBbiH po.n, HOBbie Bli.[{bl, TaKCOHOMHJI. Sciences, St. Petersburg; ZMUM - Zoological Muzeum of the Moscow State University. Introduction For species description some special male structures and genitali a were studied. Under "special male struc­ The present paper is the second part of a work devoted to tures" means: elytral depression with special appendage Ma!achius-group genera. Nine new species were de­ and antenna shape in C!anoptilus; head excavation and a scribed, a new status of the genus Ceratistes and some shape of 1-4 antenna segments in Malachius, head and new combinations and synonyms were presented in the antenna shape in Haplomalachius. Drawings fo r diffe rent first part (TSHERNY SHEV , 1998). Some new data on Siber­ species have been prepared using specimens from the ian species were given in Dahurian work (TSHERNYS HEV, following localities: Clanoptilus tedschenensis EvERS, 1999). 1985 Uzbekistan: Hissar; C. senylia sp. n. (holotype) - 200 Sergei E. TSHERNYSHEV 1 mm 5 3 0.5 mm 0.5 mm 6 Figs. 1-7 - Clanoptilus (s.str.) tedschenensis EVERS, 1985, male: 1- apex of left elytra; 2 - left antenna; 3 - ap ical tergite; 4 - apical sternite; 5 tegmen; 6 - aedeagus, lateral; 7 loca lity. Co ntributions to the knovvl edge of the lv!alachius gro up .. 20 1 Uzbekistan: Hissar; Clanoptilus strangulatus (AB EILL E side a round protuberance bea ring long li ght erect hairs DE PERRIN , 1885) Russ ia: Astakhan Area, Bogdo Mt. ; (arrowed in Fig. I), appendage very small and thin; sur­ C. pseudolutescens sp. n. (holotype) - Talysh, Georgia; face shining, shagreen, with se mi erect li ght pubescence. C/anomalachius dolini (WITTMER, 199 1) (para type) Ka­ Legs of moderate size, posteri or femora reaching the zakhstan: Tugai; C. insignicornis (PIC, 1909) Kyrgyz­ elytral ap ices; tibiae thin, rounded, straight; all tarsi 5- stan : Kazarman; C. kiesenwetteri (SOLSKY, 188 1) - Taji­ segmented, narro w; claw segment the longest, 1. 2-1 .4 kistan: Ishkashim; C. resecatus (WITTM ER, 1979) (bolo­ times longer than I , 2 and 3 taken together; all segments type) Uzbekistan: Buchara; C. debrosi (W ITTMER, 1979) simple, lacking co mbs or appendages; claws narrow and (holotype) Uzbekistan: Samarkand; C. kiihnelti (H ICKER, long, with small dent and pellucid lamell ae at the base. 1959) Kyrgyzstan: Kichik-Aiaj . Ventral surface of body dense ly punctated, with sparse white long adpressed hairs; apical sternite transverse, narrowed at apex and strongly emarginate in the middle Descriptions (F ig. 4) ; apical tergite strongly transverse, sli ghtl y nar­ rowed on sides, straight (Fig. 3), tegumen (Fig. 5) almost Clanoptilus (s.str.) tedscllenensis EVERS, 1985 pellucid, thin and transverse, with thing short appen­ (F igs. 1-7) dages; aedeagus wide and short (F ig. 6), almost pellucid with a visi ble inn er sac arranged with small teeth above Material. S Uzbekistan, Aki·abat Pass, near Shurob, and a narrow cornutus inside. h~ 1450 m, 25.Y.l997, leg. D. Milko I c3' (SZMN); female, Length (male) 3.0 mm , width (at elytral base) 1.0 mm . Uzbekistan, NW Hissar Mts, lgri su ca nyon Kok-Kishl ak Fe mal e. Similar to th e male, except as follows: antenna viii ., h~J500 m, 38°36'N 66°55 ' E, 7.YI.l997, leg. short, reaching the anterior third of elytra, serrate, almost D.Milko - I ~ (SZMN ). completely yellow; pronotum dark coloration divided onto two very sma ll spots; sc utellum ye ll ow; elytra more Description. Male. Head ye ll ow with black occiput I 51 strongly widened posteriorl y with simple apices; tibia of and 2"d antenna! joints ye ll ow, the remaining ones with all legs almost completely ye ll ow. dark sinu ate outer sides (Fig. 2). Pronotum yellow with Length (female) 2.7 mm , width (at elytral base) 0.8 5 longitudinal dark spot in th e middle, scutellum dark, mm. elytra completely yellow. Ventral sid es ye llow with dark margins; legs ye llow with dark upper sides of fe mora, Distribution. Mountain part of Uzbekistan (F ig. 7). half darkened tibia in anterior and intermediate legs and almost completely dark posterior tibia. Surface with Remarks. This species is quite specific but was purely dense light adpressed pubescence. Vesicl es, trochanters described and has not been drawn by the auth or (E VE RS, and thorax mesepimeres yellow. Dark body part lack ing 1985), the reason for the redecription of thi s species. metallic luster. Head little wider than the pronotum, front flat with three sli ght impressions: one in the middle and one above Clanoptilus (Hypoptilus) lutescens (KRAU SS, 1902) each eye; genae short and straight, clypeus narrow, trans­ (Figs. 8-15) ve rse, labrum short, transverse; palpi elongate, 2"d joint / transversa l, less than half the length of the 3 rei; apical joint Malach ius viridis ab. lutescens KR AUSS , 1902: 22 , 31 . thin, cylindrical, 1.2 times longer than th e 1'\ with thin Ma lachius viridis ab. lutescens KR AUSS, 1902 - Greiner, stretched apex; surface shining, punctures sparse, micro­ 1937: 11 8. sculpture smooth, pubescence as on pronotum. Cordy lepherus lutescens (KRAUSS, 1902) - Evers, 1985:7. Antennae (F ig. 2) long, reaching the posterior third of Clanoptilus (Hypoptilus) lutescens (KRAUSS , 1902) the elytra, the I st joint larger than th e second , ob longo­ TSHERNYS HEV, 1998: 13 9. clavate, th e 2"ci joint short, transverse, less than half of the length of th e previous joint, intermedi ate segments 3-10 Material. Uzbeki stan, Zeravshan Mts. , h ~ 1200 m, elongate, with the outer edges sinu ate , apical joint cy­ 2 l.Y. l990, Em. Strej cova leg. - I male (SZMN), river lindrica l; evenl y covered with short li ght semi erect pub­ Zeravshan va ll ey, (pr.
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