BULLETI1 DE L'li STITUT ROYAL DES SCIE 1CES NATURELL ES DE BELG IQUE ENTOi'"IOLOGIE. 70: 199-218.2000 BULLETI! VAN HET KON INKLIJK BELGISCH I STITUUT VOOR NATUU RWETE1 SCI-lAPPEN ENTOMOLOG1E. 70:' 199-21 8.2000

Contributions to the knowledge of the Malachius group of soft-winged flower (Coleoptera, Malachiidae) of the fauna of Russia and the adjacent countries. Pati II by Sergei E. TSHERNYSHEV

Abstract In this work two ne>vv species of the genus Clanoptilus are described. A special interest is atteched to a group of 2 new species: Clanoptilus senylia sp. n. (Uzbekistan: Hissar, Kyrgyz­ species with very similar characters. The shape and sculp­ stan, Alaj) and C. pseudo!utescens sp. n. (Talysh, Georgia) are de­ scribed. A rare species, Clanoptilus tedschenensis Evers, 1985 is re­ ture of the pro no tum and elytra, the coloration and shape described and fi gured. A new status, C/anoptilus (1-/ypopti/w) foveal us of antennae and heads, allows one to regard these species (MEDVEDEV, 1964) stat.n. is proposed. A key to a new Clanoptilus as belonging to a new group, Clanoptilus spinosus . .spinosu s group of species is provided. A new genus, C!anomalachius gen.n. is described ~ov ith the review of species: C/anomalachius dolini Amongst them a special male character developed differ­ (WITTMER, 199 1) comb.n., C. insignicomis (P IC, 1909) comb.n., C. ently: from a very small depression without an appendage kiesel/lvetleri (Sols ky, 188 1) comb.n., C. resecatus (WITTMER, 1979) in C. .foveatus to a deep impression with large and com­ comb.n., C. debrosi (WITTMER, 1979) stat. et comb.n., C. kiil111efti (I-IlCKER, 1959) comb.n. A generic key for the Malachius-group is plicated appendage in C. spinosus. given. Illustrations and distributional maps of species are presented. The same situation with the development of special characters occurs in a group of species with very similar Key words: Coleoptera, Ma lachiidae, ivlalachius-group, C /anopti!us, general characters which neu1ge nus, new species, taxonomy. allows this group to be regarded as Malachius or C!anoptilus according to a scheme of EV ERS ( 1985). The very specific shape of the apical tergite in males, being complexly cu rved and emarginate Pe3JOMe shows a difference from all Malachius-group representa­ tives. For these species, a new genus, Clanomalachius, is B pa6oTe omiCbiBaJOTCJI 2 HOBbiX BH,Aa: Clanoptilus senylia sp.n. described. (Y36eKHCTaH: rHccap, KHprH3HJI : AJiaH) H c. pseudolutescens sp.n. (TaJibiiii, rpy3HJ1) H HOBbiH po.n Cfaii OIIIOiachius, gen.n., All materials were borrowed from, and housed in, the c o63opoM BXO.Allll.(HX a Hero BH.AOB: C!anomalachius dolini (WITT­ following museums and collections: MER, 199 1) comb.n., C. insignicomis (PIC, 1909) comb.n., C. kiesen­ - Institute of Biology and Pedology, Kyrgh wel/eri (SOLSKY, 188 1) comb.n., C. resecatus (WITTMER, 1979) ~BPK yz comb.n., C. debrosi (WITTMER, 1979) stat. ct comb.n., C. kiihne!ti National Academy of Sciences, Bishkek; (I-IlCKER, 1959) comb.n. TiepeOilHCaH pe,AKHH cpe,AHea3HaTCKHH BH,A JSNB - Royal Belgian Institute for Natural Science, Clanopti lus tedschenensis EVERS, 1985, .naHbi pHcyHKH reHHTaJIHH H Bri.issel; Cllel..lH

1 mm

5

3 0.5 mm

0.5 mm 6

Figs. 1-7 - Clanoptilus (s.str.) tedschenensis EVERS, 1985, male: 1- apex of left elytra; 2 - left antenna; 3 - ap ical tergite; 4 - apical sternite; 5 tegmen; 6 - aedeagus, lateral; 7 loca lity. Co ntributions to the knovvl edge of the lv!alachius gro up .. 20 1

Uzbekistan: Hissar; Clanoptilus strangulatus (AB EILL E side a round protuberance bea ring long li ght erect hairs DE PERRIN , 1885) Russ ia: Astakhan Area, Bogdo Mt. ; (arrowed in Fig. I), appendage very small and thin; sur­ C. pseudolutescens sp. n. (holotype) - Talysh, Georgia; face shining, shagreen, with se mi erect li ght pubescence. C/anomalachius dolini (WITTMER, 199 1) (para type) Ka­ Legs of moderate size, posteri or femora reaching the zakhstan: Tugai; C. insignicornis (PIC, 1909) Kyrgyz­ elytral ap ices; tibiae thin, rounded, straight; all tarsi 5- stan : Kazarman; C. kiesenwetteri (SOLSKY, 188 1) - Taji­ segmented, narro w; claw segment the longest, 1. 2-1 .4 kistan: Ishkashim; C. resecatus (WITTM ER, 1979) (bolo­ times longer than I , 2 and 3 taken together; all segments type) Uzbekistan: Buchara; C. debrosi (W ITTMER, 1979) simple, lacking co mbs or appendages; claws narrow and (holotype) Uzbekistan: Samarkand; C. kiihnelti (H ICKER, long, with small dent and pellucid lamell ae at the base. 1959) Kyrgyzstan: Kichik-Aiaj . Ventral surface of body dense ly punctated, with sparse white long adpressed hairs; apical sternite transverse, narrowed at apex and strongly emarginate in the middle Descriptions (F ig. 4) ; apical tergite strongly transverse, sli ghtl y nar­ rowed on sides, straight (Fig. 3), tegumen (Fig. 5) almost Clanoptilus (s.str.) tedscllenensis EVERS, 1985 pellucid, thin and transverse, with thing short appen­ (F igs. 1-7) dages; aedeagus wide and short (F ig. 6), almost pellucid with a visi ble inn er sac arranged with small teeth above Material. S Uzbekistan, Aki·abat Pass, near Shurob, and a narrow cornutus inside. h~ 1450 m, 25.Y.l997, leg. D. Milko I c3' (SZMN); female, Length (male) 3.0 mm , width (at elytral base) 1.0 mm . Uzbekistan, NW Hissar Mts, lgri su ca nyon Kok-Kishl ak Fe mal e. Similar to th e male, except as follows: antenna viii ., h~J500 m, 38°36'N 66°55 ' E, 7.YI.l997, leg. short, reaching the anterior third of elytra, serrate, almost D.Milko - I ~ (SZMN ). completely yellow; pronotum dark coloration divided onto two very sma ll spots; sc utellum ye ll ow; elytra more Description. Male. Head ye ll ow with black occiput I 51 strongly widened posteriorl y with simple apices; tibia of and 2"d antenna! joints ye ll ow, the remaining ones with all legs almost completely ye ll ow. dark sinu ate outer sides (Fig. 2). Pronotum yellow with Length (female) 2.7 mm , width (at elytral base) 0.8 5 longitudinal dark spot in th e middle, scutellum dark, mm. elytra completely yellow. Ventral sid es ye llow with dark margins; legs ye llow with dark upper sides of fe mora, Distribution. Mountain part of Uzbekistan (F ig. 7). half darkened tibia in anterior and intermediate legs and almost completely dark posterior tibia. Surface with Remarks. This species is quite specific but was purely dense light adpressed pubescence. Vesicl es, trochanters described and has not been drawn by the auth or (E VE RS, and thorax mesepimeres yellow. Dark body part lack ing 1985), the reason for the redecription of thi s species. metallic luster. Head little wider than the pronotum, front flat with three sli ght impressions: one in the middle and one above Clanoptilus (Hypoptilus) lutescens (KRAU SS, 1902) each eye; genae short and straight, clypeus narrow, trans­ (Figs. 8-15) ve rse, labrum short, transverse; palpi elongate, 2"d joint / transversa l, less than half the length of the 3 rei; apical joint Malach ius viridis ab. lutescens KR AUSS , 1902: 22 , 31 . thin, cylindrical, 1.2 times longer than th e 1'\ with thin Ma lachius viridis ab. lutescens KR AUSS, 1902 - Greiner, stretched apex; surface shining, punctures sparse, micro­ 1937: 11 8. sculpture smooth, pubescence as on pronotum. Cordy lepherus lutescens (KRAUSS, 1902) - Evers, 1985:7. Antennae (F ig. 2) long, reaching the posterior third of Clanoptilus (Hypoptilus) lutescens (KRAUSS , 1902) the elytra, the I st joint larger than th e second , ob longo­ TSHERNYS HEV, 1998: 13 9. clavate, th e 2"ci joint short, transverse, less than half of the length of th e previous joint, intermedi ate segments 3-10 Material. Uzbeki stan, Zeravshan Mts. , h ~ 1200 m, elongate, with the outer edges sinu ate , apical joint cy­ 2 l.Y. l990, Em. Strej cova leg. - I male (SZMN), river lindrica l; evenl y covered with short li ght semi erect pub­ Zeravshan va ll ey, (pr. Samarkand), Y.l990, Em. Strej­ escence. cova leg. - I fema le (SZMN). Specimens from these Pronotum transverse, ante ri or margin pronounced, pos­ localiti es are depos ited also in the Nati onal Muse um , terior alm ost straight, th e depress ions just in sid e th e Prague and Naturhi storisches Mu eum , Base l. posterior angles di stin ct; all angles rounded; surface den­ se ly punctured, .smooth and shining, with sparse long Description. Hol otype, male. Head (F ig. 8) bl ack-green li ght and adpressed pubescence. with metalli c luster, stomal part and basa l segments of Scutellum very short and transverse, almos t hidden by palpi ye ll ow. Antenn ae dark with 2"ci -4 111 ye llow in the pronotum, covered with li ght adpressed pubescence. part, 5111 -7 111 with ye ll ow apices (F ig. I 0). Pronotum, Elytra obl ong, at the base not wider than the pronotum ; sc ute llum and almost compl etely elytra bl ac k-gree n shoulders di tin ct, not protruding; api ces with compl ex with metallic lu ster; elytral tips ye ll ow. Legs almost dark impress ion (Fig. I) being ova l and formin g on th e lower with tarsi and tibiae somew hat li ghter in anterior and 202 Sergei E. TSHERNYSHEV

.....,. I .. E ··. E ·; ..--

9 1 mm

1 mm

80° 15 / !

Figs. 8-1 5 - C/anoptilus (H. ) lutescens ( K RA USS, 1902), male: 8 head dorsall y; 9 - apex of left elytra; 10 - ri ght antenna; ll - api cal terg ite; 12 - apical sternite; 13 tegmen; 14 - aedeagus, dorsal; 15 locality. Contributions to the knowledge of the Ma la chius group I I 203

intermediate legs, knees yellow. Vesicles and thoracal 1998) these spec imens were recorded for Tajikistan. Type mesepimeres yellow. locality - Buchara. (Fig. 15). Head (Fig. 8) of the sa me width as pronotum, de­ pressed in the middle with di stal part swollen only near Diagnosis. This species is close to Cordylepherus viridis the antenna (arrowed in Fig. 8). Genae straight, clypeus and differs by di stinctly emarginate elytra with ye llow wide, transverse, bea ring a long light hairs. Palpi elon­ apices, short antenna, smaller I 51 antenna! joint. gate, its I 51 segment almost of the same length as the 3'd, and hvice bigger than the 2nd_ Apical joint narrowed and cut at the tip. Labrum long, narrow and transverse, with Clanoptilus (Hypoptilus) pseudolutescens light long hairs ahead. Surface densely punctured, micro­ TSHE RNYS HEV , sp. n. sculpture distinct, forehead with double pubescence: light (Figs. 16-23) adpressed fine hairs and sparse erected black setae. The row of black setae are set behind th e eyes. Antennae Description. Holotype, male. Head (Fig .. 16) black-green (Fig. I 0) short, reac hing; the base of elytra; I 51 joint with metallic luster, stomal part, pal pi yellow. I 51, 9' 11 , swollen, rectangular, 211 the smallest, 4 times shorter I 0111 and II '11 antenna! joints dark, the remaining yellow than the previous, the 3rd is 2.5 times bigger than the with dark basis (Fig. 17). Pronotum, scutellum and almost 2nd and 1.2 times longer than the 4111 , remaining segments completely elytra black-green with metallic luster; elytral almost equal in size, cylindrical; evenly covered with tips orange. Legs dark, only anterior tarsi somewhat short light adpressed pubescence, only the basal segments lighter, knees yellow. Vesicles and thoracal mesepimeres bear a few erect black setae. yellow. Pronotum almost equilateral, with produced anterior Head (Fig. 16) of the same width as pronotum, di s­ and posterior margins; all the angles rounded. Impres­ tinctly depressed in the middle with swollen di stal part sions near the posterior angles distinct. Surface with (arrowed in Fig. 16). Genae straight, clypeus wide, trans­ minute and dense punctures, microsculpture di stinct, pub­ verse, bearing a long light hairs. Pal pi elongate, its I 51 escence as on forehead. segment almost of the same length as th e 3rd, and twice Scutellum short and transverse, shining, bearing short bigger than the 2nd. Apical joint narrowed, stretched to adpressed hairs. the apex and cut at the tip. Labrum narrow, transverse, Elytra parallel, not wider th an pronotum at the base, with light long hairs. Surface dense ly punctured, micro­ evenly expanded posteriorly. Shoulders distinct, slightly sculpture visible, forehead with double pubescence: light protruding. Elytral tips sli ghtly impressed (Fig 9). Surface adpressed hairs and sparse erect black setae. The row of granulate, covered with fine light adpressed hairs and black setae are set behind the eyes. Antennae (Fig. 17) black erect setae. short, reaching the base of elytra; I 51 joint swo llen, rec­ Legs of a moderate size, posterior femora almost tangular, 2'"1 the smallest, 3 times shorter than the pre­ reaching the elytral tips. Tibiae thin, rounded, straight, vious, the 3rd twice bigger than the 2'"1 and 1.2 times posterior slightly curved inside. All tarsi 5-segmented, longer than the 41h, remaining segments almost equal in narrow; apical joint is the longest, its length being equal size, slightly serrate; evenly covered with short light to the total length of the I 51+2nd in anterior legs, 1.2 times adpressy d pubescence. shorter in intermediate and 1.1 times longer than the I '' P!fonotum almost equilateral, with produced anterior joint in posterior legs. Claws narrow, with pellucid la­ and straight posterior margins; all the angles rounded. mellae at the base. Depressions at the posterior angles distinct. Surface with Ventral side of thorax with light hairs, abdomen with minute and dense punctures, microsculpture smooth, pub­ short adpressed and sparse pubescence. Apical sternite escence as on forehead. (Fig. 12) strongly elongate, emarginate in the middle. Scutellum short and transverse, shining, bearing short Apical tergite (Fig. II) longitudinal, with rounded sides. adpressed hairs. Phallus (Fig. 14) simple, with th e number of small teeth Elytra parallel, not wider than pronotum at the base, on the inner sac; co rnutus straight, rounded and thin slightly ex pand ed posteriorly. Shoulders di stinct, slightly (arrowed in Fig. 14). Tegum en (Fig. 13) longitudinal, protruding. Elytral tips strongly impressed (Fig 18). Sur­ 2.2 times longer than wide, inner side emarginate; thin face finely granulate, covered with fin e li ght adpressed appendages widely flatten ed at th e apices, appendage­ hairs and black erect setae. base length ratio is 1.1. Legs of a moderate size, posteri or femora almost Length (mal e) 4.3 mm , width (at elytral base) 1.4 mm. reaching th e elytral tips. Tibiae thin , rounded, straight, Female. Similar to male, except as follows . Antennae posterior sli ghtly curved inside. All tarsi 5-segmented, narrower, almost completely dark . Elytra more strongly narro w; aFical join is the longest, its length being equal to widened posteri orl y. Elytral apices simple, with wide of I 51+ 2'" total length of anterior legs, somewhat shorter ye ll ow spot. in intermediate and almost of the sa me length as I 51 joint Length (female) 5.1 mm , width (at elytral base) in posterior legs. Claws narro w, short, with small lamel­ 1.7 mm. lae at the base. Ventral side of thorax with long li ght hairs, abdomen Distribution. In the first part of th e article (TSHERNYSHEV, with short adpressed and sparse pubescence. Apical ster- 204 Sergei E. TSHERNYSHEV I I

E E E E

1 mm 16

17

19 1 mm

Fi gs. 16-23 - Clanoptilus (H.) pseudolutescens TSHERNYS HEV , sp. n., male: 16 head dorsally; 17 - ri ght antenna; 18 - apex of left elytra; 19 - apical tergite; 20 - apical sternite; 2 1 tegmen; 22 - aedeagus, dorsal; 23 loca li ty. Contributions to th e knowledge of the Malachius group I I 205

nite (Fig. 20) longitudinal, emarginate in the middle. bei Malachius foveal us MEDVEDEV keine A nhange besit­ Apical tergite ( Fig. 19) rectangular, longitudinal, with zen". The investigation of a serial material shovvs, that slightly emarginate anterior side. Phallus (Figs. 22) sim­ small size is a strong, but not variable c haracter in this ple, with the number of small teeth on the inner sac; species. All males have no appendages in slight depres­ cornutus straight, rounded and wide (arrowed in sions of elytral apices. A differences also can be found in Fig. 22). Tegumen (F ig. 21) longitudinal, 2.2 times longer aedeagus inner structure. These facts seem to be enough than wide, inner side twice emarginate; thin appendages to reestablish this species as a good one. The species was flattened at the apices, appendage-base length ratio is 1.1. previously known only from Kazakhstan, this is a first Length (male) 5.1 mm , width (at elytral base) 1.8 mm. record for Russia and Mongolia. Female. Similar to male, except as fo llows. Antennae narrower and shorter, not reaching t he base of elytra. Elytra more strongly widened posteri orl y. Elytral apices Clanoptilus (s.str.) simple, with very small round orange spot. (ABE ILL E DE PERRIN, 1885) Length (female) 5.3 mm, width (at elytral base) 1.9 mm. (Figs. 24-32)

Distribution. As shown under Material onl y (Fig. 23). Cyrtosus strangulatus AB EILLE DE PERRIN, Rev. d'Ent. IV, Probably it is limited in range by Caucasus region. 1885, p. 143 Holotype, d', Caucasus, Talysh, Lerik district, Gosmaljan Malachius slrangulatus (ABEILLE DE PERR I , 1885) GRE I­ · env., II. Vl.1988, S.Kasantsev leg. (SZMN), para types: NER, 1937:11 6. idem- I d' (SZMN); Caucasus, Talysh, Lenkoran district, Clanoptilus strangulatus (ABE IL LE DE PERRIN, 1885) Avrora e nv., 13 .VI.I988, S.Kasantsev leg. - ~ (SZMN); EVERS, 1985 : 36. Caucasus, Georgia, Shiraki near Tsiteli-Tskaro, = Malachi us vulneratus ABEI LLE DE PERR IN, 1891 EVERS, 7.V.I983, W.Dolin 5 d'd' (NHMB), I " (ISNB); Georgia, 1985: 39-40. Vale, ~ 1700-2100 m, 23.VI.I978, W.Dolin 3 d'd' 2 ~~ = Malachius pseudospinosus L.M EDVEDEV, 1964 EVER S, (NHMB); I d' I ~ (ISNB); I d' I ~ (SZMN). 1985: 39-40.

Diagnosis. It can be distinguished from the true C. lutes­ Material. , Donau-Au, date-?,- ld' (NHMB), cens by long a ntenna, reaching the anterior fourth of Ukraine, Azov Sea, Berdiansk, 9.V!. I981, B.Pavlov-Ver­ elytra, its Ii ghter coloration; strongly impressed elytral evkin leg. I d' (SZMN); Ukraine, Crimea, Golaia Pristan', ap ices colored in orange; a shape of swoll en part in head 3.V. I995, - S,Vasilenko leg. I d' (SZMN); Russia, K.ras­ (arrowed in Figs. 8, 16) a nd wide inner structure of nodar Area, Ub inskaia Station, Papai Mt., ~ 1700 m, aedeagus (arrowed in Figs. 14, 22). 30.4. 1989, A.Solodovnikov leg. I d' (SZMN); Russia, Astrakhan Area, steppe near Nizhiy Baskunchak viii. and Bogdo Mt., 30.V.I996, V.Dubatolov leg. I d' 2 ~ Clanoptilus (Hypoptilus) foveatus (SZMN); E-Kazakhstan, S Bank of Zaisan Lake, 5 k m (L.MEDVED EV, 1964) stat. n. ESE ofrriozernoe, II.VI.I 997, R.Dudko & V.Zinchenko leg. 2 " "', I (SZMN); E-Kazakhstan, E Prizaisanje, 20 Malachiusfoveatus L.MEDVED EV, 1964: 156. km JNE of Karatal, sands of Bozalgyrkume, 12- Clanoptilus s trangulatus ssp. fovea/us MEDVED EV, 13.V 1.1 997, R.Dudko & V.Zinchenko leg. ld' (SZMN); EV ERS, 1985: 39-40 Kyrgyzstan, N slope of Kyrgyzsky Mt. Range, Valley of Karabalta Riv., ~ 1800 m, 30.V.I995, D.Milko leg. 2d'd' Material. Kazakhstan, I 0 km N of Zharkol Lake, (SZMN). I OVI.I957 holotype, d' (ZISP), para types: 2 ~~ (ZISP); £-Kazakhstan, Zaisan Di strict, Rozhkovo viii., sweeping, Description. Holotype, male. Head black-green with me­ 15 .V!.I986, V.S hilenkov leg. I d' I (SZMN); Mongo li a, · tallic luster, clypeus and mandibles yellow, antennae South-Gobi Aimak, Toirym Rav ine, 68 km NE of Tsa­ black. Pronotum and almost completely elytra black­ gan-Bogdo Mt. , desert with Tamari x and Calli gonum, green with metallic luster, only the elytral tips red. Scu­ 23 .V!.I 973, G.Medvedev leg. I d' (ZISP); Russia, Volga­ tellum and legs black with metallic luster. Ventral side of grad Area, 1-5 k m E of Pallasovka viii. , dry steppe with thorax, coxae and trochanter black, borders of sternites Euphorbi a and Tanacetum, 25 .V. I996, V.Dubatolov leg. and thorax mesepimeres yellow. Vesicles red. - ld' (SZMN). Head not wider than the pronotum, with thin light adpressed pubescence a nd bunches of erect black hairs Remarks. The special male structures of new species are on the temples; forehead slightly impressed between very well shown in MEDVED EV's description ( 1964): antennae and sparsely punctuated with di sti nct micro­ "elytra with di stinct, but not deep impression ... appen­ sculpture a mong punctures; labrum transverse, 1.3 ti mes dages lack ing". EvERS ( 1985) decided that this spec ies is longer than the c lypeus, rounded anteriorl y and beari ng only a va riation of C. strangulatus, a usual r esident of white sparse long hairs; genae short, straight; eyes of the Central As ian steppes, but he had included the phrase: moderate size, ro und, w ith slightly produced anterior " ... Int eressant ist aber die Tatsache, daf3 die Excitatoren side. Pal pi elongate, the I st joint is 1.3 times longer than 206 Sergei E. TSHERNYSHEV

1 mm,

Figs. 24-32 C!anoptilus (s.str. ) strangulatus (As.-P ERR. , 1885), male: 24 - ri ght antenna; 25 - apex of left elytra; 26 appendage in elytral apex; 27 - api cal tergite; 28 - ap ical stern ite; 29 tegmen; 30 - aedeagus, dorsal; 3 1 aedeagus, lateral ; 32 - locali ty. Contributions to the knowledge of the tvfa/achius group 207

I I

the 2"d, being the same length as the 3rd; apical joint fer by the rough black pubescence of surface, bright red narrowed at apex; surface s hining with light semierect elytral apices and a s hape of elytral appendage (Fig. 26) pubescence. Antennae long, reaching the half way along attached on a round vertical plate and being small above the elytra; the I 51 segment clavate, not swollen, 3 times with short hairs distally. longer than the 2"d and about 1.5 times longer than the remammg· · , -'"ret -5' 11 segments o f t h e equa IS .izes , t I1 e otI 1ers a little longer, all segments, excepting I 51 and 2"d, cylind­ Clanoptilus (s. str.) senylia TSHERt YSHEV, sp. n. rical (Fig. 24); covered with thin light adpressed pubes­ (Figs. 33-41) cence with sparse erect hairs a mongst them. Pronotum almost equilateral, somewhat wider than Description. Holotype, male. Head black-green with me­ long, anterior margin strongly pronounced, posterior tallic luster, clypeus, mandibles and anterior part of lab­ straight, all angles rounded, posterior with distinct im­ rum yellow, antennae black. Pronotum and elytra almost pressions; surface densely punctate with di stinct micro­ completely black-green with metallic luster, only the sculpture and double pubescence: light adpressed and elytral tips yell ow. Scutellum and legs black with slight black erect. metalli c luster. Ventral side of thorax, coxae and trochan­ Scutellum transverse-rectangular, small , distinctly de­ ters black, borders of sternites, vesicles and thorax me­ pressed in the middle; anterior side straight, surface shin­ sepimers yellow. ing, densely punctured and pubescent. Head narrow, with slight interocular depression, cov­ Elytra oblong, parall el, very slightly widened poster­ ered wi th thin light adpressed pubescence, the row of iorly; shoulders distinct; apices evenly rounded and erect bl ack hairs is o n the temples; forehead sparsely strongly impressed (Fig. 25) with double lamellate ap­ punctuated, microsculpture distinct; labrum short, trans­ pendage inside ( Fig. 26), its vertical lamella rounded, versal, twice longer than the clypeus, rounded anteriorly horizontal very small bearing a bunch of short white and bearing white long erect hairs; genae short, straight; curved hairs; the upper side of each impression angular eyes protrudent, round. Pal pi elongate, the I 51 segment is pronounced, the lower side rounded, with stretched angu­ 1.5 times longer than the 2"d, the 2"d segment is twice lar apex; surface moderately shining, fi nely granulate, shorter than the 3rd; apical segment narrow, slightly with fine, light and short adpressed pubescence and dense stretched at apex; surface shining with light semierect dark and very strong erect setae. long hairs. Antennae (Fig. 33) long, reaching the middle Legs long and thin, posterior femora not reaching the of elytra; the I 51 segment clavate, not swollen, 3.5 times elytral apices, covered with short light adpressed hairs; longer than the 2"d, the 3rd is twice longer than the claw segments bear long black setae a nteriorly. Tibiae previous, of the same length and shape as 4 111 ,5 111 or 6111 , thin, slightly expanded anteriorly, rounded. All tarsi 5- somewhat widened at apex, being triangular-shape, the segmented , narrow; 4111 segment I. S sma II est, b e.m g twi.ce remaining, segments elongate, almost cylindrical, very shorter than the apical which is largest, being 1. 2- 1.3 slightly widened at apex excepting the a pical, which is times longer than the I 51• Claws thin, with oval pellucid pointed. All segments covered with th in light and sh01i lamellae at the base. adpressed pubescence and sparse light erect hairs. Apical sternite (Fig. 28) narrow, transversal, deeply Pronotum almost equilateral, 1.2 times wider than emarginate in the middle; apical tergite (Fig. 27) elon­ long, anterior margin arcuate, posterior straight, all angles gate, rectangular, 1.2 times longer than wide, anterior side r~H1cled , with distinct depressions just at the posterior very s li ghtly emarginate, covered with dark long hairs. angles; surface densely punctate, microsculpture distinct, Phallus (Figs. 30, 3 1) simple, thin and long, with small covered with double pubescence: light short and ad­ teeth on the inner sack, the only medial cornutus narrow pressed and sparse black erect. (arrowed in Fig. 30); apex straight. Tegumen longitudi­ Scutellum transverse-rectangular, small , almost com­ nal, 2 times longer than wide, with moderate e margina­ pletely hidden by the pronotum, slightly depressed in the tion in the inner si de, with long thin appendages, dis­ middle; its surface shining, densely punctate and pubes­ tinctly fl attened and widened at the tips (Fig. 29). cent with light hairs. Length (male) 5.5-5.9 mm, width (at elytral base) 1.6- Elytra subparallel, strongly widened posteriorly; 1.7 mm. shoulders distinct; apices evenly rounded and strongly Female. Similar to male, except as fo ll ows. Antennae depressed (Fig. 34); lamellate appendage consisting of narrower. Elytra more strongly w idened posteriorl y. Ely­ two parts, horizontal, being rectangular and beari ng a tral apices simple, with large round red spot. bunch of black long straight hairs, and a vertical which Length (female) 5.8 mm, width (at elytral base) is pointed on a distal side (Fig. 35); the upper side of each 1.8 mm. evenly rounded, slightly produced, the lower side rounded, with stretched angular apex; surface shin ing, Distribution. Ranged from Europe (, , Hu­ shagreen, covered with fine, light and adpressed pubes­ gary, Ukraine, Russia) to Central Asia (Kazakhstan, Uz­ cence and sparse semierect d ark strong setae. beki stan, Kyrgyzstan) (Fig. 32). Legs long and thin, posterior femora not reaching the e1y tra l a pices, covered w ith short lig ht adpressecl hairs; Diagnosis. All representatives of the spinosus-group dif- claw segments bear l ong black setae on the upper si de. 208 Sergei E. TSHERNYSHEV

E E E E L!) 0

38 1 mm

Figs. 33-41 - Clanoptilus (s.str.) senv lia T SHERNYS HEV, sp. n., male: 33 - ri ght antenna; 34 - apex of left elytra; 35 appendage in elytral apex; 36 - api ca l tergite; 37 - api cal sternite; 38 tegmen; 39 aedeagus, lateral; 40 - aedeagus, dorsal; 4 1 - locality. Contributions to the knowledge of the Malachius group 1 I 209

Tibiae thin, slightly expanded anteriorly, rounded. All - Appendage in lower side of elytral impression lacking tarsi 5-segmented, narrow; 4111 segment is smallest, being horn (male only) . . strangulatus (AB.-P ERR., 1885) twice short as the apical joint, which is largest and being 1.2-1.3 times longer than the I 5\ 2"ci or 3'"ci segments. Claws narrow, sharp, with di stinct oval pellucid lamellae C/anoma/ac/tius TSHERNYSHEV, gen. n. at the base. Type species: Malachius resecatus WITTi\IIER, 1979 Apical sternite (Fig. 37) transversal, wide, with deep narrow emargination in the middle; apical tergite (Fig. 36) Definition. Quite big malachiid beetles, ranging from 5 to rectangular, 1.3 times longer than wide, anterior side 8 mm in length . Surface with double coloration: almost evenly rounded, covered with sparse black hairs. Phallus black-green with metalli c luster and partly ( legs, elytral (Figs. 39, 40) simple, slightly curved ventrally, with small apices, stomal part etc.) o range-yellow; covered with teeth on the inner sac, the medial cornutus big and wide bl ack erected and light adpressed hairs. Antenna I ! -seg­ (arrowed in Fig. 40); apex of aedeagus wide, pointed. mented (Fig. 61 ), flabell ate in male and slightly toothed Tegumen longitudinal, 1.9 times longer than wide, emar­ in female, long, reaching a fi rst quarter of the elytra; the ginate in the middle, with long thin appendages, slightly I ' 1 joint swollen, cylindrical, ventrall y depressed. Re­ flattened and widened at the tips ( Fig. 38). mai ning segments with the outer edges extremely Length (holotype) 5.0 mm, width (at elytral base) 1.4 stretched, except the apical, being parallel. Jnterocular mm. pl ate on head between antenna slightly elevate. Pronotum Female. Similar to male, except as follows. Antennae with orange-yellow margins on sides. Thorax mesepimers narrower. lnterocul ar d epression feeble. Elytral apices and anterior trochanters yellow. First tarsi 5-segmented si mple, very s li ghtly depressed, with small yellow spot. covered with very long and fine white hai rs, all segments Length (female) 5.1 mm, width (at elytral base) 1.4 mm. simple. Elytral apices with depressions which are small (as in C. dolini) or strong (as in C. debrosi). Appendage in Distribution. Holotypus: (j (SZMN), Uzbekistan, NW the depression lacking in C.dolini, very s mall in C. in­ Hi ssar Mts., Jgrisu canyon, Kok-Kishlak viii., signicornis and C. kiesenwetteri and quite big in remain­ h- 1500 m, 38°36'N66°55 'E, 7.YI.1997, D.Milko leg.; ing species. Apical tergite strongly emargi nate (Fig. 64) all otype -

E E

E E

47

42

1 mm

48

Figs. 42 -49 - Clanomalachius dolini (WITTMER, 199 1) , male: 42 - ri ght antenna; 43 - apex of left elytra; 44 - ap ica l tergite, dorsal; 45 - apica l tergite, ventral ; 46 - ap ica l sternite; 47 tegmen; 48 aedeagus, lateral; 49 - locality. Contributions to the knowledge of the Malachius group 21 I

E E

1 mm

Figs. 50-59 - Clanomalachius insignicornis (PIC, 1909), male: 50 h ead, dorsal; 5 1 - right ante1ma; 52 - apex of left elytra, author drawing; 53 - apex of right elytra, from Wittmer, 1979; 54 - apical tergite, dorsal; 55 - apical tergite, ventral; 56 - apical sternite; 57 tegmen; 58 aedeagus, lateral; 59 - locality. 212 Se rgei E. TSHERNYS HEV ''

4. C. resecatus (WITTM ER, 1979) co mb.n. of Susa myr Ri ver, 27.V. l9 14, Mihalevskaya - 2 5. C. debrosi (W ITTM ER, 1979) stat. et comb.n. c3' "(Z ISP); Naryn, N ri verside, 12 k m W A k-Tal_ Tchat 6. C. kiihnelti (Hl CKER , 1959) comb .n. Viii. , h- 1600 m, 4 1°26 'N 74° 56' E, l.VII.I 999, D.M ilko leg. - 2c3'c3' ; 6 ~ (SZMN). Tajikistan: Teriklitau, near Key to species (males onl y) Kum san Vill age, 27.IV. l99 1, S.Ovchinni kov leg.- I c3'; I. A ppendage in elytral depression absent (F ig. 43) 2 ( SZM N)...... C. dolini - Appendage in elytral depress ion present, bein g very Diagnosis. Th e species ca n be easil y di ffe renti ated by the small or quite bi g (F igs. 52 , 87) ...... 2 bl ack colorati on of th e interantennal part of head (Fi g. 50), 2. lnterocul ar pl ate between antenna a bove clypeus whil e thi s c haracter in other species is light (Fi g. 60). bl ack (Fi g. 50) ...... C. insignicomis Elytra with very deli cate impression (F ig. 53), appendage - In te rocul ar pl ate betvvee n ante nna above clypeus ye l­ small , fl at, bea rs a long bl ack chaeta fixed in underside low (F ig. 6 . .0 ) ...... 3 (F ig. 52). The 3'"d antenna! segment with strongly 3. Appendage in elytral depress ion ve ry s mall , not stretched outer side (Fi g. 51 ). Api ca l tergite s ubparall el, reaching the lower margin (Fi g. 62, 71) ...... 4 (F ig. 54), possesses long and short dense hairs in bunches - Appendage in elytral depression bi g, reaching th e on ante ri or side (F ig. 55). Api cal stern ite t ransversa l, lower lnargin (Fi g. 79, 87) ...... 5 trapezium-shape ( Fi g. 56). Tegumen and aedeagus as in 4. Append age very s mall , pellucid , with very thin and Fi gs. 57, 58. long bl ack bristl es fixed under it . C. kiesenwelleri - Appendage short, strong, with 3-4 thin bl ac k bri st les Distribution . Ranged in Afghani stan, Kyrghyzstan and fi xed on sides and below ...... C. resecatus Turkmeni stan. This is a first record in Tajikistan. 5. Upper side of elytra l depression with a stretched pl ate (F ig. 87) ...... C. kiihnelti - Upper side of elytral depression evenl y rounded, with- Clanoma/achius kiesemvetteri (SOLSKY , 188 1) comb. n. out stretched pl ate (F ig. 79) ...... C. debrosi (Fi gs. 60-69)

Ma lachius kiesenwetteri SOLSKY , Horae Soc. Ent. Rossi­ Clanomalachius dolini (WI TTM ER, 199 1) comb. n. cae, 188 1, XII (3-4): 242-245 (F igs. 42-49) Clanoptilus kiesenwelleri (SOLSKY) : EvE RS, 1985 :35 .

Clanoptilus (Hypoptilus) dolini Wi TTM ER, Mi tt. EGB , 4 1, Ma terial. Uzbeki stan: Hi ssar Mt. Range, Turaps~ Ra­ 199 1: 29 Abb. 4 1-46 vin e, 4 km SS W Sanga rdak Vill age, H- 1550 m, 28 32 'N 67°3 1'E , 3.Vl.l 99 7, D.Mil ko leg.- 3 c3'c3' , 5 ~ (SZMN, Material. Kazakhstan: Tugaj on Charyn Ri ver, H-620 m, IBPK); Kyrghyzstan: Sa ry-Chelek Natural Reserve, 5 km 25 .V.I990, W.Dolin - holotypus, c3' (NHMB), paratypes ­ S Arkyt, H- 1200 m, 41 °47'N 71°59'E, 29.V l.l 996, I c3' , I ~ (S ZM N); Kyrghyzstan: NE Fergana, Kara-Un kur O.Milko leg. - 3 c3' "; 5 ~~ (SZMN); Ferga nsky Mt. Rav ine, 3 km SS W Charvak Viii. , 18. V. I993, D.Mil ko Range, Kara-A lma va ll ey, H- 1600 m, 5-7.VI.l 997, leg. I ~ (SZMN) \Y<'6olin leg. - I c3' ; 3 ~ (SZMN ); Ferga nsky Mt. Range, Al as h Vill age, A las h fo restry, I O.Vl.l 997, S.Ovchinni­ Diagnosis. Di ffers f rom all congeners by t he a bsence of kov leg. - I "(SZMN); Tajikistan: Gorno-Badak hshans­ append age in elytral api ces (Fi g. 43) and co lorati on of kaya Avtonomn aya Oblast, 8 km SE from Ishkashim to surface with green-bronze metalli c lu ster. The 3'd anten­ Dasht, 23.V l.l 984, Co li.-? - 2 c3'c3' (SZMN ). na! segment evenl y rounded (F ig. 42). Api ca l t ergite parall el (Fi gs. 44) with a few hairs in bunches at th e tips Diagnosis. Similar t o C. insignicornis (Pic), excepting of anteri or side. Api cal sternite (F ig. 46) short, transver­ characters as foll ows. Elytra more strongly impressed sa l. Tegum en and phallus as in fi gs. 47, 48 . (F ig. 63). Append age di stinct, lame! al te, wi th short strong chaetae. Head between antenn ae ye ll ow (F ig. 60). Outer Distribution. Kn own from its type locality onl y (Fi g. 49). side in 3'd antenna! joint less sinu ate ( Fi g. 6 1). Apical tergite wid e, its anteri or side evenl y emargin ate (F ig. 64) and pubescent ( Fi g. 65). Api ca l sternite transversal, tri ­ C!anomalachius insignicomis (PI C, 1909) comb. n. angul ar-shape ( Fi g. 66). Tegumen and aedeagus a re (F igs. 50-59) show n in Figs. 67, 68.

Ma lachius insignicom is PI C, Ec hange, XXV, 1909, Distribution. The species was described from Turkestan p. 129. (th e upper strea m of Zeravshan Ri ver and near Alai Clan optilus insignicom is (Pic): EVE RS, 1985 :35. vill age). Later, recorded in Tajikistan (W ITTME R, 1979). Ranged widely in moun ta in ous parts of Uzbekistan and Material. Ky rghyzsta n: Kazarm an environs, H-2000 m, Ky rghyzstan: a nd , probabl y, can b e fo und in th e s ame 13.V lll.l 972, Yu. Tarbin sky leg. - I J (SZMN); vall ey landscapes of Eastern Kazakhstan (F ig. 69). Contributions to the knowledge of the Malachius group , • 2 13

E E ~tsv 63 60

70° so•

69 i

Figs. 60-69 - C/anoma/achius kiesenwetteri (SOLSK Y, 188 1) , male: 60 head, dorsal; 61 - ri ght antenna; 62 - apex of left elytra, author drawing; 63 - apex of ri ght e lytra, from Wittmer, 1979; 64 - apical tergite, dorsal; 65 - api cal tergite, ventral; 66- apical sternite; 67 tegmen; 6 8 aedeagus, lateral; 69 - locality. 214 Sergei E. TSH ERNYSHEV

E E

•• 0 ••••• .·. · . .... • :. 0 . . . . ; ..

0 .. ·::: .;: . . :·. .. . 0: ... . · .·

1 mm 76

right antenna; 71 - apex of left elytra; Figs. 70-77 - Clanoma/achius resecalus (WITTMER, 1979), male: 70 - 75 tegmen; 76 aedeagus, 72 - apical tergite, dorsal; 73 - apical tergite, ventral; 74 - apical sternite; latera l; 77 - locality. Contributions to the knowledge of the Malachius group , , 215

Clauomalachius resecatus (WITTM ER, 1979) comb. n. species in the shape of antenna! joints. Investigations of (Figs. 70-77) special structures in all congeners allows Clanomalachius debrosi to be regarded as a good species. In the original Malachius resecatus WiTTM ER, Ent. Arb. M. Frey 28, description (WITTM ER, 1979) the e lytral apex was 1979:3 Abb. 5-6 wrongly shown in Fig. 8. The paratype from Tajikistan, Clanoptilus resecatus (WITTM ER) : EVERS, 1985 :36 used for the preparation of this figu re is surely C. kiihnelti (HICKER) and sho uld be e liminated from the type series. Material. Uzbekistan: Buchara, Khodzha-fil '-ata, Investi gation of the holotype of C. debrosi has shovvn the ll.V. l910, N.Zarudny - holotypus, o (ZISP); Buchara, differences in shape of the e lytral impression of this Kim·is, 22.V.l91 0, N.Zarudny - 4 para types, ~ (ZISP); species. Zeravshan Mt. Range, Aman-Kutan Pass, H-1250 m, 39°18'N 66°54'E, 8.VJ.l997, D.Milko leg. - I o (SZMN); Kughitang Mt. Range, Surkhansky Reserve, Clauomalachius kiilme/ti (HICKE R, 1959) comb. n. Kompyrtepa, H-2400 m, 37°42 ' N 6 6°36' E, 30.V.I 997, (Figs. 86-93) D.Milko leg. - I 0 (SZMN). Ma/achius kiihnelti HI CKER, Ent. Nachrichtenbl. I I, Diagnosis. The species differs by its s hort wide appen­ 1959:7 1 dage and strongly impressed elytral tips (Fig. 71 ). The 3rd Clanoptilus kiihnelti (HICKER): EVE RS, 1985:35. antenna! joint with slightly stretched outer side (Fig. 70). Apical tergite wide, complicatedly emarginate on anterior Material. Kyrgyzstan: Kichik-Alaj Mt. Range, Kirgh iz­ side (Fig. 72) and bearing a bunches of hairs mainly on Ata Ravine, H-2 150 m, 40°07'N 72°35 'E, 23.Vl.l 990, the inner lateral sides (Fig. 73). Apical sternite (Fig. 74) D.Milko leg. - I o (SZMN). narrow, somewhat elongate. Tegumen and phallus as in Figs. 75-76. Diagnosis. Special male characters strongly di stinct in C. kiihnelti. Elytral impression deep, with stretched lamellae Distribution. Was described from Buchara. New collec­ in upper side and long curved lower side ( Fig. 87). Ap­ tions show it is wide spread in Uzbekistan (Fig. 77). pendage strong and long, bears a bunch of curved chaetae on t I1 e t ip.. T l1 e 3 rd antenna! segment large, tn.angu lar (Fig. 86). Apical tergite strongly emarginate (Fig. 88) Clauoma/achius debrosi (WITTMER, 1979) stat. et comb. n. and pubescent ( Fi g. 89). Apical sternite narrow, with (Figs. 78-85) wide e margination in the middle ( Fig. 90). Tegumen and phallus as in Figs. 91, 92 . Malachius resecatus ssp. debrosi Wi TTMER, Ent. Arb. M. Frey 28, 1979:5 Abb. 7 Distribution. Th is rare species was known only from its type locality, according tothe description: Buchara, Uz­ Material. Uzbeki stan: Samarkand Prov., Galliaaralsky beki stan. A new finding in Kirghizia w idens notions of Distr., Zhi li bu lak, on Vexibia sp. , 20.V. I964, Haidarova the range. Probably, the speci es can be found in the other - holotypus, o (ZJSP); Nuratau Mountains, Nuratau State pal):s- ofCentral Asia as in the care of C. kiesenwetteri, for Reserve, 5 km S Fari sh, 2. VI.I 985, Andreeva - ~ia mpl e (Fig. 93). lo (SZMN); SY RDA RI A PROV, AU LI E-ATA, 12.V 1.1 913, CHERNA­ VIN; KYRGYZSTAN: ALA I, 5.VI.I 885 - I c)(Z ISP); CHU RI VE R, Key to Malachius-group genera 3. VI.I 955, COL-? - I J(SZMN). TCHANDALASH RAV INE. 6 N JCT. I. Male spec ial structures not distinct ...... 2 TC HATKA L RIV., H- 1630 M, 41°44'N 70°52'E, 21. VI.I 999, D.MILKO Male special structures distinct, located on antennae, LEG.- 33'J , W'f: (SZMN). head or elytral apex ...... 3 2. Pronotum transverse f-laplomalachius Ev ERS, 1985 Diagnosis. The species di ffers by very long narrow ap­ Pronotum longitudinal .. Microlipus LECONTE, 1852 pendage and deep vertical impression (Fig. 79). The 3'd 3. M ale s pecial structures located in elytral apex as antenna! segment almost tri angular (Fig. 78). Ap ical impressions or appendages ...... 4 tergite wide (Fig. 81) , bunches of hairs disposed on each Male special structures located on head as excavations tip of tergite inner structure (Fig. 80). Apical sternite or protuberances ...... 5 narrow, almost tri angul ar (Fig. 82). Tegumen and phallus 4. First antenna! joint swoll en, cyl indrical, apical tergite as in Figs. 83 , 84. twice emarginate on distal part, with bunches of ha irs ...... C/anomalachius, gen.n. Distribution. Ranged in Uzbeki stan, Ky rgyzstan and Ta­ First antenna! joint not swollen, clavate or paral lel, jikistan (F ig. 85). apical tergite with evenly rounded di stal part, bunches of hai rs lacking . . Clanoptilus MOTSCI-IULSKY, 1853 Remarks. Was described as subspecies (WI TTM ER, 1979). 5. Male head excavati on lacking ...... 6 The author h as shown the main character, a very long Male head with deep transverse excavation . . . . 7 appendage, but di d not find differences with the other 6. Male head with smooth protuberance between anten- 2 16 Sergei E. TSHERNYSI-IEV

E E

1 mm 81 84

Figs. 78-85 - Clanomalachius debrosi (WITTMER, 1979), male: 78 - ri ght antenna; 79 - apex of left elytra; 80 - api cal tergite, ventral; 8 1 - api cal t ergite, dorsal; 82 - apical sternite; 83 tegmen; 84 aedeagus, lateral; 85 - loca lity. Contributions to the knowledge of the Ma/a chius group '' 217

. ·.·• • ..• • 0 •••. . 0 •••

• • 0 • . .• .• · ...0 • ••

E E

87

1 mm 92

Figs. 86-93 - Cfanomalachius kiihnelti (H lCKER , 1959), male: 86 - ri ght antenna; 87 - apex of left elytra; 88 - apical tergite, dorsal; 89 - apical terg ite, ventral; 90 - apical sternite; 9 1 tegmen; 92 aedeagus, lateral; 93 - locali ty. 218 Sergei E. TSHERNYSHEV I>

na, 5111 segment simple . Co rdylepherus EVERS, 1985 EV ERS, A.M.J., 1985. Auftei lung der pa laa rkti schen Arten des Male head with strong hom betvveen antenna which is Gattungcomplexes Malachius F. Entomologische 8/iiller, 81( 1- round and cut at the tip, 5111 segment large, funnel - 2): 1-40. shape ...... Ceratistes FISHER DE WALHEIM , 1844 GRE INE R, J., 1937. Ma lac hiidae. In : Junk-Schenking. Co leop­ 7. Mal e head excavation located behind the antenna in a terorum Catalogus. Pars 159 : 1-1 99. distal part of head, protuberance bearing long hairs KRAUSS , 1-1 ., 1902. Ge nus Ma /a chius. In: Bestimmungs-Tabel­ disposed just inside the excavation, antenna wi th len der europaischen Co leopteren 49 : 1-33. appendages ...... Ma lachius FABRI CIUS, 1775 MEDVEDEV, L.N. , 1964. New species of Ma lachiidae (Coleop­ Male head excavation located before the antenna, tera) from Central Kazakhstan. Trudy ZIN, 34: 155-1 58 [in distal part lacking protuberance, antenna simple, ap- Ru ss ian] pendages lacking ...... SOLSKY , S.M., 188 1. New and little known Co leoptera of the ...... A nhomodactylus MAYOR et WITTMER , 1981 suburbs of the Russ ian Empire and adjacent coun tries. Horae Societatis Entomo/ogicae Rossicae, X11(3-4) : 237-26 1 (In Rus­ sian with Latin descriptions) Aknowledgements TSHERNYSHEV, S.E ., 1998. Towa rd s the knowledge of the Ma ­ lachi us group soft-winged flower beetl es (Coleoptera, Ma la­ The main new material cited is give n by co ll eagues from th e Institute chiidae) of the fauna of Ru ss ia and the adj acent countries. Part of Biology and Pedo logy, Kyrgyz Academy of Sc iences (B ishkek), I. Russian Entomological Journal, 7(3-4): 129-1 46. Dmitri Milko and Serge i Ovchinnikov. I am pl eased to ex press many th anks for their kind help . TSHERNYS HEV , S.E., 1999. Beetles of the fa mili es Canthari dae, I am gratitude to Drs G.S.Medvedev, B.A.Korotyaev (ZIN RAN , St.­ Mal ac hiidae, Oedemeridae, Meloidae, Chrysomelidae of the Petersburg), N.B.Nikitsky, G.Y u.Lubarsky (ZMUM , Moscow), V.vih­ Dahurian Nature Reserve. of Dalwria and adjacent la, NM P (Praha), V.G.Shilenkov (Irkutsk University) for th e loa n of the territories. Proceedings of th e Dahurskii Stale Biosphere, ll: va lu abl e materi al under their cure. 94-111 (In Russian wi th Engli sh summary) I am sin cerely grateful to Professo r E.G. Be llinger, Central Europea n University, Budapest, for th e kindest help in correcti on of th e manu­ WITTMER, W., 1979. Zur Kenntnis der Gattung i\lfa/a chius F. sc ript. sensu Jato (Co l. Malachiidae). Entomo/ogischen Arbeiten aus Thi s work is partly supported by a gra nt of Siberian Branch Pre­ dem Museum G.Frey, Tutzing, 28: 1-8. sidium of Russian Academy of Sciences, ac ts 83 of I 0.3.2000.

Dr Sergei E. TSHERNYS HEV References Siberian Zoological Museum, Institute of Animal Systematics and Ecology, ABEILLE DE PERR IN, E., 1890-9 1. Malac hiidae. Mal achides Russ ian Academy of Sciences, d' Europe et pays vo isins. Annates de Ia Societe entomo/ogique Siberian Branch, de . 6 Serie. Tome X. I ( 181) - 680(284), Tome. LX. Frun ze street, II , 285(115)- 446(442). Novosibirsk 630091 Russ ia

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