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Bell and Bell 1979 Qev15n4 377 446 CC Released.Pdf This work is licensed under the Creative Commons Attribution-Noncommercial-Share Alike 3.0 United States License. To view a copy of this license, visit http://creativecommons.org/licenses/by-nc-sa/3.0/us/ or send a letter to Creative Commons, 171 Second Street, Suite 300, San Francisco, California, 94105, USA. This work is licensed under the Creative Commons Attribution-Noncommercial-Share Alike 3.0 United States License. To view a copy of this license, visit http://creativecommons.org/licenses/by-nc-sa/3.0/us/ or send a letter to Creative Commons, 171 Second Street, Suite 300, San Francisco, California, 94105, USA. RHYSODINI OF THE WORLD PART II. REVISIONS OF THE SMALLER GENERA (COLEOPTERA: CARABIDAE OR RHYSODIDAE) ROSS T. BELL Department of Zoology University of Vermont Burlington, VERMONT 05401 JOYCE R. BELL 24 East Terrace Quaestiones Entomologicae South Burlington, VERMONT 05401 15:377-446 1979 This paper is the second of a series which will constitute a revision of the Rhysodini of the world and consists of revisions of all genera except for the four largest ones. Treated are: Leoglymmius Bell and Bell; Tangaroa Bell and Bell; Dhysores Grouvelle; Neodhysores Bell and Bell; Rhysodes Dalman; Kupea Bell and Bell; Kaveinga Bell and Bell; Grouvellina Bell and Bell; Xhosores Bell and Bell; Yamatosa Bell; Shyrodes Grouvelle; Srimara Bell and Bell; Arrowina Bell and Bell; and Plesioglymmius Bell and Bell. Some of the genera have been redescribed, incorporating new information. Yamatosa NEW NAME replaces Yamatoa Bell 1977 which is preoccupied by Yamatoa Kiriakoffl967 In Kaveinga Bell and Bell, four subgenera are recognized: Angekiva, NEW SUBGENUS, rype-Kaveinga frontalis (Grouvelle), one sp., Australia; Ingevaka, NEW SUBGENUS, rype-Kaveinga orbitosa (Broun), one sp., New Zealand; Vakeinga, NEW SUBGENUS, two spp., New Zealand, New Caledonia; Kaveinga (sensu stricto), 14 spp., New Guinea, Bismark Archipelago, Solomon Islands, Santa Cruz Island, Moluccas, and Mindanao. In Plesioglymmius Bell and Bell, three subgenera are recognized: Plesioglymmius sensu stricto two spp., Sumatra, Borneo, Mindanao; Ameroglymmius NEW SUBGENUS, 0>/>e-Plesioglymmius meridionalis (Grouvelle), three spp., South America, Cuba; Juxtaglymmius NEW SUBGENUS, rype-Plesioglymmius jugatus NEW SPECIES, one sp., Java. Rhysodes parvus Grouvelle, accidentally omitted from Part I, is assigned to Kaveinga (sensu stricto). The following new species are described (and type localities indicated): Dhysores pan (ZAIRE, Kivu, Itombwe, 2300 m, Terr. Uvira, Nyalengwe); Dhysores liber (LIBERIA, crest of Mt. Nimba, 1400 m); Kaveinga fibulata (NEW BRITAIN, Rabaul): Kaveinga pignoris (SOLOMON IS., Bougainville, Kokure, 690 m) Kaveinga nudicornis (SOLOMON IS., Russell Is., Yandina); Kaveinga kukum (SOLOMON IS., Guadalcanal, Kukum): Kaveinga ulteria (SOLOMON IS., Santa Cruz Group, Reef Is.); Kaveinga lupata (NEW GUINEA, Mt. Kaindi, 16 km SW Wau); Kaveinga okapa (NEW GUINEA, Kamira, Eastern Highlands); Kaveinga marifuanga (NEW GUINEA, Marifuanga, Asaro-Chimbu Divide); Kaveinga histrio (PHILIPPINE IS., Mindanao, e. slope Mt. McKinley, 3300', Davao Prov.); Kaveinga strigiceps (Buru); Grouvellina gigas (MADAGASCAR, Chutes de la Mort); Grouvellina cuneata (MADAGASCAR, Tamatave and Foret Alahakato); Grouvellina hova (MADAGASCAR, Tsaramainiandro); Grouvellina ranavalona, Grouvellina edentata, Grouvellina cooperi, Grouvellina montana (MADAGASCAR, only); Grouvellina radama (MADAGASCAR, Mahatsinjo); Grouvellina descarpentriesi (MADAGASCAR, Annanarivo); Grouvellina divergens (MADAGASCAR, Mt. d'Ambre); Grouvellina dentipes (MADAGASCAR, Fizono); Grouvellina cinerea (MADAGASCAR, Perinet); Arrowina pygmaea (SRI LANKA, Dikoya); Plesioglymmius silus (PHILIPPINE IS., Mindanao, Zamboanga, Kabasalan); Plesioglymmius reichardti (VENEZUELA, 378 Bell and Bell Suapure, Caura R.); Plesioglymmius compactus (CUBA); and Plesioglymmius jugatus (JA VA). Cet article est la deuxieme d'une serie qui constitueront une revue taxonomique des Rhysodini du monde. La deuxieme partie est composee des revisions taxonomiques de tous les genres saufles quatre les plus grands. Les genres discutes sont: Leoglymmius Bell et Bell; Tangaroa Bell et Bell; Dhysores Grouvelle; Neodhysores Bell et Bell; Rhysodes Dalman; Kupea Bell et Bell; Kaveinga Bell et Bell; Grouvellina Bell et Bell; Xhosores Bell et Bell; Yamatosa Bell; Shyrodes Grouvelle; Srimara Bell et Bell; Arrowina Bell et Bell; et Plesioglymmius Bell et Bell. Nous decrivons certains genres de nouveau, comprendant des renseignements nouveaux. Yamatosa NOUVEAUNOM remplace Yamatoa Bell 1977, car il est preoccupee de Yamatoa Kiriakoff 1967. Nous divisons le genre Kaveinga Bell et Bell parmi quatre sous-genres; Angekiva NOUVEAUX SOUS-GENRE, type-Kaveinga frontalis (Grouvelle), une sp., Australie; Ingevaka NOUVEAU SOUS-GENRE, 0>/>e-Kaveinga orbitosa (Broun), une sp., Nouvelle Z'elande; Vakeinga NOUVEAU SOUS-GENRE, type-Kaveinga setosa (Grouvelle), deux spp., Nouvelle Zelande et Nouveau Caledonie; Kaveinga (sensu stricto) 14 spp., Nouvelle Guinee, Archipel de Bismarque, lies de Solomon, lies de Saint Croix, lies Moluques, et Mindanao. Nous divisons le genre Plesioglymmius parmi trois sous-genres: Plesioglymmius sensu stricto, deux spp., Sumatra, Borneo, et Mindanao; Ameroglymmius NOUVEAU SOUS-GENRE, fy/?e-Plesioglymmius meridionalis (Grouvelle), trois spp., Amerique du Sud et Cuba; Juxtaglymmius, NOUVEAU SOUS-GENRE, rype-Plesioglymmius jugatus NOUVELLE ESPECE, une sp., Java. Par hazard, nous omlmes Rhysodes parvus Grouvelle de la premiere partie. C'appartient au genre Kaveinga, sous-genre Kaveinga (sensu stricto) On d'ecrit les especes nouvelles que void, en indiquant pour chacune la localite du specimen type: Dhysores pan (ZAIRE, Kivu, Itombwe, 2300 m, Terr. Uvira, Nyalengwe); Dhysores liber (LIBERIA, cr'ete du mont Nimba, 1400 m); Kaveinga fibulata (NOUVELLE BRETANGE, RABAUL): Kaveinga pignoris (ILES DE SOLOMON, Bougainville, Kokure, 690 m) Kaevinga nudicornis (ILES DE SOLOMON, He Russell, Yandina); Kaveinga kukum (ILES DE SOLOMON, Guadalcanal, Kukum): Kaveinga ulteria (ILES DE SOLOMON, groupe Santa Cruz, He Reef); Kaveinga lupata (NOUVELLE GUINEE, Mt. Kaindi, 16 km S.O. Wau); Kaveinga okapa (NOUVELLE GUINEE, Kamira, Highlands de I'Est); Kaveinga marifuanga (NOUVELLE GUINEE, Marifuanga, Asro-Chimbu Divide); Kaveinga histrio (ILES PHILIPPINES IS., Mindanao, versant est du mont McKinley, 3300', province du Davao); Kaveinga strigiceps (INDONESIE, Buru); Grouvellina gigas (MADAGASCAR, Chutes de la Mort); Grouvellina cuneata (MADAGASCAR, Tamatave et Foret Alahakato); Grouvellina hova (MADAGASCAR, Tsaramainiandro); Grouvellina ranavalona, Grouvellina edentata, Grouvellina cooperi, Grouvellina montana (MADAGASCAR, sans localites specifiees); Grouvellina radama (MADAGASCAR, Mahatsinjo); Grouvellina descarpentriesi (MADAGASCAR, Annanarivo); Grouvellina divergens (MADAGASCAR, mont d'Ambre); Grouvellina dentipes (MADAGASCAR, Fizono); Grouvellina cinerea (MADAGASCAR, PERINET); Arrowina pygmaea (SRI LANKA, Dikoya); Plesioglymmius silus flLES PHILIPPINES IS., Mindanao, Zamboanga, Kabasalan); Plesioglymmius reichardti (VENEZUELA, Suapure, Riv. Caura); Plesioglymmius compactus (CUBA); et Plesioglymmius jugatus (JAVA). Classification of Rhysodini 379 TABLE OF CONTENTS Introduction 379 Sources of Material 379 Subtribe Leoglymmiina 380 Subtribe Dhysorina 381 Fig. 1 - 12 383 Subtribe Rhysodina 386 Fig. 13-24 391 Fig. 25 - 39 392 Fig. 40-51 393 Subtribe Clinidiina 408 Fig. 52 - 63 414 Fig. 64-74 415 Subtribe Omoglymmiina 423 Fig. 75 - 87 426 Fig. 88-102 433 Fig. 103-114 440 Acknowledgements 443 Supplementary References 443 Indices 444 INTRODUCTION This paper, the second of a series of five, includes revisions of all the genera of Rhysodidae except Clinidium Kirby, Rhyzodiastes Fairmaire, Omoglymmius Ganglbauer, and Pyxiglymmius Bell and Bell. Some of the genera have been redescribed, incorporating new information. Otherwise, subtribal and generic descriptions have not been repeated; instead, the reader is referred to the appropriate pages in Part I (Bell and Bell 1978). A short list of additional references is added to supplement the extensive list in Part I. SOURCES OF MATERIAL The following abbreviations designate collections cited in this paper. The names in parentheses are the curators of the respective institutions. AIM Auckland Institute and Museum, New Zealand (K.A.J. Wise) ; AMS Instituut voor Taxonomische Zoologie, Amsterdam, Netherlands (J. Duffels); BMNH British Museum, Natural History, London (R. Pope); BPBM Bernice P. Bishop Museum, Honolulu (G Samuelson); Quaest.Ent., 1979, 15 (4) 380 Bell and Bell BSL Naturhistorisches Museum, Basel, Switzerland (W. Wittmer); CAS California Academy of Sciences, San Francisco, CA (D. Kavanaugh); CMP Carnegie Museum of Natural History, Pittsburgh, PA (G. Wallace); CNHM Field Museum of Natural History, Chicago, IL (H. Dybas); DSIR Department of Scientific and Industrial Research, Auckland, N.Z. (J. Watt); DY Daniel K. Young, Michigan State University, E. Lansing; LCC Lincoln College, Canterbury, N.Z. (R.M. Emberson); LEI Rijksmuseum von Natuurlijke Historie, Leiden, Netherlands (J. Krikken); MCZ Museum of Comparative Zoology, Cambridge, MA (J. Lawrence); MNHB Museum fur Naturkunde der Humboldt-Universtat, Berlin, DDR (F. Hieke); MNHN Museum National d'Histoire Naturelle, Paris, France (A. Descarpentries); MRAC Musee Royal de l'Afrique Centrale, Tervuren, Belgium (P. Basilewsky); MZSP Museu de Zoologia da Universidade Sao Paulo, Brazil (U.R. Martins);
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