New Land Snails from Montenegro and Albania Introduction
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Arch. Molluskenkunde | 138 | (1) | 103–112 | 14 figures | Frankfurt am Main, 30.06.2009 New land snails from Montenegro and Albania (Mollusca: Gastropoda) PETER SUBAI Abstract Four land snail species from Montenegro and Northern Albania are characterised concho- logically and described as new: Acicula miaphene n. sp., Platyla corpulenta n. sp., Platyla feheri n. sp. and Hiltrudia (?) globulosa n. sp. New records for species of land snails from Montenegro are submitted. Key words: Taxonomy, Anatomy, Acicula, Platyla, Hiltrudia, Albania, Montenegro, new spe- cies, new records. Kurzfassung Es werden 4 Landschneckenarten aus Montenegro und aus Nord-Albanien anhand des Ge- häuses neu beschrieben: Acicula miapheneeschweizerbartxxx sng- n. sp., Platyla corpulenta n. sp., Platyla feheri n. sp. und Hiltrudia (?) globulosa n. sp. Es wird über weitere für Montenegro neue Vorkommen von Landschnecken berichtet. Schlüsselwörter: Taxonomie, Anatomie, Acicula, Platyla, Hiltrudia, Albanien, Montenegro, neue Arten, Neunachweise. Introduction The majority of the molluscan specimens mentioned tors during the past century. The increasing activities of here was collected during vacation trips of 1–2 weeks in malacologists in the past years yielded a various number the years 2004–2006 in Montenegro. The specimens of of species new for science which four of them will be the new taxon Platyla corpulenta n. sp. originate part- described in the present work. ly from collections of the Hungarian malacologists in The spelling of Albanian localities follows FEHÈR et northern Albania1. These areas often been visited by ma- al. (2004). lacologists in the past, have been inaccessible for collec- All measurements in millimetres. Author’s address: PETER SUBAI, Kronenberg 143, D-52074, Aachen; E-mail: [email protected] © E. Schweizerbart’sche Verlagsbuchhandlung (Nägele u. Obermiller), 2009, ISSN 0003–9284 DOI 10.1127/arch.moll/0003-9284/138/103-112 104 SUBAI, P.: New land snails from Montenegro Abbreviations: S private collection P. SUBAI, Aachen SMF Naturmuseum Senckenberg, Frankfurt a. M. E private collection Z. ERőSS, Budapest alk. specimen preserved in alcohol H private collection A. HUNYADI, Budapest AD diameter of aperture HNHM Hungarian Natural History Museum, Buda- D shell diameter pest H height of the shell Maa private collection J.M. W. MAASSEN, Duiven- AH height of the aperture drecht juv. juvenile specimen MMMB Munkácsy Mihály Museum, Békéscsaba W number of whorls NMW Natural History Museum, Wien UTM UTM-Code for the inventory of the European NNM Nationaal Natuurhistorisch Museum, Leiden Invertebrates Systematic account Caenogastropoda COX 1960 tilineata BOETERS et al. 1989 are similar in their overall Aciculidae J.E. GRAY 1850 shell shape, but considerably smaller. Their apertures are more rounded at the base, they have a more strongly Acicula miaphene n. sp. developed bulge of the neck, and the pattern of furrows Figs 1, 4 on the teleoconch is more regular and denser. D i a g n o s i s : Shell large, broad, with narrow shal- L o c u s t y p i c u s : Montenegro, at NW-rim of Mokrine low bulge of the neck, densely and irregularly striped (= NNW from Herceg Novi), at limestone rocks, 575 m alt., with longitudinal furrows, angularis almost reduced, UTM BN 90. without sinulus. Ty p e m a t e r i a l : holotype SMF 329430, paratypes from the type locality: leg. SUBAI 13.10.2004, S 20787/5+5 (juv.); Description: Shell large, low conical; fresh speci- leg. SUBAI 19.9.2005, S 20679/52+35 (juv./damaged); leg. SUB- men are red- to yellowish-brown; apex (the first 2–2.25 AI 17.9.2006, E/5, Maa/5, HNHM 96781/5, NMW/5, NNM/5, whorls) smooth, the teleoconch whorls with dense, ir- SMF 330757/5, S 20966/83+37 (juv./damaged); leg. SUBAI regularly arranged longitudinal furrows; penultimateeschweizerbartxxx sng- 14.4.2009, S 22256/3+4 (juv./damaged). whorl with 55–75 furrows; 6.75–7.25 whorls, the last E t y m o l o g y : miaphene is a fantasy name. whorl occupies 48–52 % of the total height; sutural ridge Distribution p a t t e r n s : This species is only reduced. known from the type locality in the Kotor Bay in Mon- The aperture slightly ascends at the penultimate tenegro (= NNW of Herceg Novi). whorl, and is acutely oval in frontal view (Fig. 4); ap- ertural rim blunt, slightly reinforced at the palatal side, Platyla corpulenta n. sp. more strongly calloused at the basal and columellar side, Figs 2, 5 where it is enlarged to form a small triangle. The parietal callus well developed, covering the periomphalum with D i a g n o s i s : shell medium-sized shell, very broad, a faint stripe, a columellar callus is missing; angularis white; angularis and sinulus reduced, bulge of the neck reduced. The apertural rim is slightly curved in lateral missing; subterranean mode of life. view, a sinulus is missing. Description: shell medium-sized, broad, conical, The bulge of the neck is moderately developed. It colour white (fresh specimens translucent); surface of starts with a small channel behind the apertural rim. the teleoconch smooth, the few furrows mark interrup- Measurements (n = 171): H = 5.25–6.12; tion of shell growth; whorls 5.5–6.25, regularly increas- D = 1.93–2.18; AH = 1.56–1.81; AD = 1.18–1.37; meas- ing, sutural ridge weak or completely reduced; last whorl urements of the holotype: H = 6; D = 2.06; AH = 1.75; occupies 56–60 % of the total shell height. AD = 1.31; W = 7.25. The aperture slightly ascends at the penultimate whorl, Differential d i a g n o s i s : Due to its size and and is acutely oval in frontal view; apertural rim blunt, its dense pattern of furrows on the teleoconch, A. mia- slightly reinforced at the palatal side, more strongly cal- phene is easily separated from all related species. Ac- loused at the basis and strong at the columellar side. The icula disjuncta BOETERS et al. 1989 has the same shell parietal callus is thin, and covers the periomphalum with height, but it is considerably more slender and has a a narrow stripe; the angularis is reduced or missing, the much lesser number of furrows (13–20 on the penulti- sinulus is always missing. The apertural rim is slightly mate whorl). Additionally, the bulge of its neck is weak- curved in lateral view. The bulge of the neck is always er. Both, A. hausdorfi BOETERS et al. 1989 and A. mul- lacking. SUBAI, P.: New land snails from Montenegro 105 from locus typicus: leg. SUBAI, 21.9.2005, S 20495/1 (dam- aged); leg. SUBAI, 16.9.2006, S 21050/4+6 (juv./damaged). Montenegro, Rumija Mountains, 4 km from Arbneš/ Arbneshi following the road to the south (= approx. 2 km S of pass), in crevices in limestone rocks, approx. 350 m alt., UTM CM 65, leg. SUBAI 21.9.2005, S 20522/1+1 (damaged); leg. SUBAI 16.9.2006, HNHM 96782/1, S 21013/1+4 (juv./dam- aged). Albania, Periferi Kukës, Gurri i Arrënit E of Arrën (= 35 km S of the Shkodër–Kukës road), limestone rocks, degrad- ed beech forest, 1600 m alt., UTM DM 44, leg. DELI, ERőss, FEHÉR & MURÁNYI 8.10.2005, E/1 (juv.), HNHM 96803/1 (juv.), MMMB/1 (juv.). N o t e : The locus typicus of P. corpulenta and hither- to only record of its closest relative P. procax is approx. 26 km apart. E t y m o l o g y : The name corpulenta describes the very broad shell which is characteristic for this species. Distribution p a t t e r n s : This species is only known from three localities in Montenegro, in the north- ern part of the Rumija-Mountains and from the Kroništar Mountain, (both localities are south of Lake Skadar) as well as in Northern Albania, approx. 35 km SSW of Kukës. The Montenegrian and Albanian localities are lo- cated in a distance of approx. 130–135 km. The species Figs 1–3: New species of Aciculidae. — Fig. 1: Acicula mia- lives subterraneous. phene n. sp., Montenegro, at NW-rim of Mokrine (= NNW of Herceg Novi), at limestone rock and in a rock cavity, 575 m alt., UTM BN 90. (holotype SMF 329430, H= 6, D= 2.06 mm). Fig. 2: Platyla corpulenta n. sp., Montenegro, Kroništar Moun- Platyla feheri n. sp. tain, southern slope, approx. 10 km E of Ðuravci (= in direc- Figs 3, 6 tion to Ostros, S of Lake Skadar), at limestone rocks,eschweizerbartxxx sng- approx. 350 m alt., UTM CM 56. (holotype SMF 329428, H= 3.93, D= D i a g n o s i s : shell medium-sized, white, smooth, 1.8 mm). Fig. 3: Platyla feheri n. sp., Montenegro, Kroništar aperture subquadrate without sinulus, bulge of the neck Mountain, southern slope, approx. 10 km E of Ðuravci (= in narrow, slightly upraised; subterranean mode of life. direction to Ostros, S of Lake Skadar), at limestone rocks, ap- Description: shell medium-sized, slender conical, prox. 350 m alt., UTM CM 56. (holotype SMF 329429, H= colour white (fresh specimens translucent); surface of the 3.62, D= 1.37 mm); all phot. E. NEUBERT, × 15. teleoconch smooth, the few furrows mark interruption of shell growth; whorls 6.5–7.25, regularly increasing, su- tural ridge distinct; last whorl occupies 44–49 % of the Measurements (n = 8): H = 3.68–4.3; D = 1.68– total shell height. 1.93; AH = 1.31–1.5; AD = 1–1.25; measurements of The aperture slightly ascends at the penultimate whorl, the holotype: H = 3.93; D = 1.8; AH = 1.43; AD = 1.12.