Exploration D Pare Nati Nal Albert Exploratie Va Het Nati Aal Albert Park
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INSTITUT DES PARes NATIONAUX INSTITUU T DER NATI ONA LE PARKEH DU CONGO BELGE VA N BE LGISC H CONGO Exp lo ration d Pare Nati nal Albert M ISSION G. F. DE W ITTE (1933-1935) F ASCICU LE 63 Exploratie va het Nati aal Albert Park ZENDI G G. F. DE WITTE (1933-1935) AFLEVERI NG 63 PA PICORNIA J. 1I LFU{;I -Bl 0 \- •.'E, .r. \. (Lo (Iun ) BRUXELLES BRUSSEL 1950 1950 INSTITUT DES PARCS NATIONAUX jlNSTlTUUT DER NATIONALE PARKEN DU CONGO BELGE VAN BELGISCH CONGO Exploration du Parc National Albert MISSION G. F. DE WITTE (1933-1935) FASCICULE 63 Exploratie van het Nationaal Albert Park ZENDING G. F. DE WITTE (1933-1935) AFLEVERI NG 63 PALPICORNIA GY .J. BALFOUH-BROvVNE, M. A. (London) BRUXELLES BRUSSEL 1950 1950 Imprimerie M. RAYEZ. Bruxelles 112. rue de Louvain. 112 Dom. légal: r. de la Chancellerie, 4 PARC NAllONAL ALBERT NATIONAAL ALBERT PARK 1. MISSION G. F. DE WITTE 1933-1935 I. ZENDING G. F. DE WITTE 1933-1935 Fascicule 63 Aflevering 63 PALPICORNIA Il)" J. BALFOUR -BROWNE M. A. (London). The Palpicornia of the Albert Natiollal Park collected by the .Mis sion G. F. DE WITTE had been, in part, determinecl hy the late M. A. D'OHCHYMONT before his regretted death carly in i\Vl6. Shortly lhereafter Dr. VAN STHAELEN asked me to undertake the completion of the determination of the DE WITTE collections and also those of the Mission H. DAMAS, and to prepare a report thereon. l wish here to express my sincere thanks to Dr. VAN STHAELEN for the opportllnity to study these very rich collections, both numerically and seientifically, representing a total of over 103 species, which furnish the richest single material l have yet seen from Afr·ica. In those cases where fj'ORCHYl\IONT had assigned, in collection, a name to new species recognised by him in his preliminary study l have retainecl the name proposed hy him. Of the total of 103 named species dealt with hereinafter 22 species are descrihed as new to science, foUI' belonging to the Hydréenid3' , one to the llydrochina:?, fifteen to the Sph,rridiillir and two to the H1fdro philinée. The high proportion of new species occuring in the Hydr3'nidrc emphasises that this family is still but very imperfectly known from Africa, the emphasis heing the greater for the fact that only 20 specimens belonging to this group were obtainecl out of the very large total of the whole collection. Again, the fifteen new species of Sp}uEl'idù:n,r, out of a total of 44 species taken, compared with only two new species of Hydrophilin,E, out of 48 species taken, suggests that the terrestrial forms of the IIydrophilidiE are still relatively unknown whilst the truly aquatic forms from « normal» habitats are relatively weIl known; this, perhaps, is but a measure of the faet that previous collections have tended to ignore the smaller forms of the Sphéerid'iinœ from non-aquatic habitats. 4 l'ABC NATIONAL ALBERT A nalysis of the vertical and horizontal distributions as detailed in the accompanying Tables, indicates that for the Palpicornia as a whole (Table 1) the numerical preponderance of species occurs at the low inter mediate zone (1.001 to 2.000 ml, the next preferred zone being the lowest (0 to 1.000 ml; contrarily only two species attain to the highest zone (3.001 to 4.000 ml. This altitudinal preference holds true (Table 2), but without significant value, for the HydréPnida? and for the subfamilies I1J/drochiiléP and Hydrophilinœ. The Spcrcheùla? shew a decided preference for the lowest zone, no specimens occurring above 1.000 m. On the other hand the Spha?ridiina?, whilst shewing the same maximum preference for the low intermediate zone at 1.000 to 2.000 m, have their next, highest pre ference for the high intermediate zone (2.001 to 3.000 m) and are the only group to attain to the highest zone ove1' 3.001 m. This difference in preference must undoubtedly be significant, having regard to the numerically rich material on which the opinion is based. Horizontally, as is to be expected, a greater number of species are seen to be common to the East African Highlands and to the Albert National Park than are common to the Park and to the relatively low lying area of the Congo basin with altitudes rarely exceeding 600 m. TABLE 1. 0 1 1lJl ci s S S 0 d 0" .S .S S P E C 1 E '" '" '" ... .;; S '" '" '"~ .. '" '" .... d -< -< " '"g ~ ...:. " :t ~ ..00 t5 '"~ '" <Il "'"" '" p:j .0 ~ '"",i ..;'" >il" i;:'" Hydreen idee. Or:h tl/l'Iritis l'l'y i III {wrti K,\ISCH x + - andreinü? s. sp. RÉGIMBART X + Hydr:ena (s. str.) pet/onga nov. "p.. X - albeTti nov. sp. X X Li-mnebius damasi sp. nov. X X + wit/ei sp. nov.. X - sp.. X Spercheidee. 1 SpeTcheus cerisy'i GUÉRI'i X + T seneua/ensis \.ASTEL:\AU X + + + III/méroUs nü;rMBAnr ' X + Hydrophilidee. 1 Hy,iroclws aUJicaTls RÉGIMBART X T - al/JatillT/us sp, HOV. ... ... , .. X Cœlosto1ll11 plllJ!IICToides Tubrocinctlllil RÉGIMBART, ,..... '" X >~ + + + m,ocquerljsi [)'OnCHYMONT '" X + -- Tu/Uars/' ROIIE\IA~. .. X X ;< + + \ \ NATIONAAL ALBERT PARK 5 SPECIES Cœlostoma wittei sp. nov. X subsphœroides HÉGIMBART X + - deplexum sp. nov. ... x - sp. ... -1 -' sp. ............ X DactylostfrnUl/I 11litellllale D'OHCHY· MONT X X + - raTum J(\ISCH + + - a!JI/omillrrle f'ABHICIUS X + + + Splucridilu/I IJO tleqoi MAHCUZZI X X + caftrl/1/! CASTELNAU ... X x X + + + si.1II pliri/I('s l\IARCUZZI X -+ liisimwlulII sp. no\'. + olJs('urnm HÉGIMIlART ... X X + + aliditum D'ORCHY~IOl\T X + + nbductum sp. nov. X X 1 exile BOHEMAN - ... .. X X + " thornsoni D'OnCHYMQNT X + -+ + Sphœridium sp. X Cerc!Jon dieganus HÉGIMBART . X + putricoln WOLLASTON X + + + picius RÉGIMBAHT .. X X -+ + + sp. ............ X tigreanus GEMMINGER & HAHOLD X + vnlCllnif1!S sp. no\'. X sturmi ROTH X + + soilo/is sp. nm'. X X wittei sp. no\'. x X x atriN/I!Wl/s I\IAHSHAM X + + + sp. X sJl. x sulJfilis KNISCH X + ('ostl'rnoiill's K\ISCH ... X + parastromoides J. BALFOUH·BHOWNE X + Parnstromus crislatus .1. BALFOUR· BROWNE ............ X pelocyon jeanneli O'ORCHYMOXT X + Oosternum horni n'ORCHYMONT X Quadristernum africanum sp. nov. X Tectosternum brunneum Sp. nov. X Pachysternum capense Ml'LSANT ._. X + + + - gestroi D'ORCHYMONT ... X + - wittei sp. nov. X - slllc'atl1m sp. no\'. X CT!JJltoplClITlITI/ sutural1l1ll RÉGIMBART. X + + - costatum. BOHEMAN :.< + - rotun (/UIII sp. nov. X - histl'roirles sp. nov. X + Pyretus gm.tlls sp. nov. X Paro('!JlIIusminor RÉGIlIIBART X + - ('haleeus RÉGI~mAIIT X X + + - ollullUiliallus SCOTT X + T.,accolJius gradUs I\IOTSCHULSKY ab. col. ......... ... ... ... .. X X + Helorl/.ares (H.) 71otaticollis RÉGIMBART X X =+= - strirtlls O'OHCHYMO>;T .. ... X X + =+= ... PARC NATIONAL ALBERT Il 0 1 1 El 1: ~ i oi .i s S 0 1 .S: <:> <:> <:> 0 1 .S 1 <:> Q Q .. .. 8 P E C 1 E S El <:> Q 1 .... .... ~ 1 -1j Q 1 ;; Q -< .., <:> ~ '" ! 'bI, 1 ... Q ~ Q .... <:> Q Q -; ~ '" 0 .... Coi ..,; >!:i f"l ~" Il ! 1 1 1 lIetoc/wres merSl1S D'ORCHYMO'lT ... X x - - 1 + + -_. steni.us D'ORCHYMONT ... ... .. ... - x x - li + - liviŒ/WS D'ORCHY1VlONT ... ... ... - x x - - -. + 1 - col/ortl D'ORCHYMONT ... ... ... ... - 1 1 - _. _. x x 1 minax D'ORCHYMmT ... ... ... .. -- + - x 1 + - dimorphus D'OHCHYMmT - - + ... ... ... x 1 - + + - lnenlinotus KUWERT ... ... ... ... x - - - - 5p. ... ... ... ... ... ... ... -, _. - + +_. x - -- sp. ... ... ... ... ... ... ... ... _ - x - -_. - - (C.) alricanl1s D'ORr:lIYMONT ... ... x - - - + - - sp. ... ... ... ... ... ... ... .. x - - - - - sp. ... ... ... ... ... ... ... .. -- - x - - - - (s. str.) longipalpis MURRAY ... ... X X - + + + - (S. str.) nilut1Js ERICHSON ... ... X X - _. + + + - (8. str.) ellipticus RÉGIMBART ... - - - + + - (S. str.) pallens MAC LEAY X -- -- + + - (s. str.) Ji. /œvlllscllius RÉGIMBART _.X - - - - + - PellochŒres conspicU1LS RÉGIMBART ... - - - + + Enochrlls (M.) lwtalensis GEMMINGER & HAROLD ... ... ... ... ... X X - + + + - ci'rcumnuctus RÉGIMBART ... ... - X X -- + + - Ilnticus RF.GIMnART ... ... ... ... X X _. - 1 + + - Il cspcridll'J/l \VOLLASTO-' ... ... X X - - - +_. + -- alberti Sp. nov. ... ." ... ... ... - - X - - S ternoloplws soUer; CASTELNAU ... X X - + + +1 - angolensis ERICHSON ... ... ... X X X - + ,.. + H ydro]Jhilus senegalensis PERCHERON X - - - + + + - aeulca/us SOLIER ... ... .. ... X X - - -' '" + + Hydrochara elliptica FABRICIUS ... ... X X X - +_. + Neohydrophilus strael.eni sp. nov. ... X - -- - neplanatus D'ORCHYMONT .. ... .. X X - - - _.. + - distinc/us HOPE ... ... ... ... .. - X - + + + - wehnckei D'OLIVEIRA ... ... - X - - + + AmpIl iOps phalliclls D'ORCHYMO:'iT X X - -- + + - - - - - 1 wittei D'OnCHYMOI'IT ... ... .. - - Il + - seneylllensis CASTELNAU ... ... X X X - + + + - y/obus EmcHsoN ... ... ... .. - X - _. + + + Aliocotocerlls seriatus RÉGIMBART ... X X X + + + - semirotllnrtns D'ORCHYMONT ... ... - -- - + + - sllbœncl1s ERICHSON ... ... ... X X -- + + + llerosl1S (E.) cuspidatus ERleHSON X X X - + -- + - fnrcatus BOHEMA:'i . " ... ... ... X X -- 1 1 + + 1 - (5. str.) nigriceps FABRICIUS ... ... X X -- - + - (s. st1'.) vitticoUis BOHEMAN ... ... X -- - ..L+ + - 1 + - (s. str.) nigropunctatus sp. nov. - _. - - + - Ile!l'Î'lnbartia elliplicll RÉGIMBART ... ... X - - X 1 + + - nilotica SHARP ... ... ... ... ... X X - - 1 + ,1 + + . 1 1 TOTALS. 1 50 74 1 34 2 1 56 1 71 33 \ ----- ----- .,. NATIONAAL ALBERT PARK 1 TABLE 2. ci !>Il 1 :::; d â S â 0 d <:> <:> <:> 0 .;:; <:> <:> <:> " ,::" SPECIES â <:> :::; ... <:> co co ~ 1 "1" os ...: <:> ~ ..!.'" .... .... t; co <:> <:> <:> j en .... 1 co co <:> os c; Ô .... N ..,; P<'" ~ 1 1 1 '" 1 1 j Hydrœnidœ ... :~ [) t 0 0 3 0 Spercheidœ ... :3 Ù 0 0 2 3 1 Hydrophilidœ Sph:eridiina' ... ... ... 1 10 32 22 2 16 , 24 14 Hydrophilidœ : 1 1 , Hydra('llinél~ + HydrophUin:e ... ... 34 37 11 0 3H 1 H 18 1 1 1 The Holotype specimen of aIl new species will be deposited in the « Institut Royal des Sciences Naturelles de Belgique» (Brussels), paratypes in the same institution and in the British Museum. The localities between [ lare without the Park's boundaries. 8 PARC NATIONAL ALBERT HYDR.E:N 1D.E: Subfam. HYDRtENINtE. Ochthebius (Homalochthebius) regimbarti KNISCIJ. Ochlhebius (Homalochthebius) regimbarti KNISCH 1924, Coleopt.