Beetles in Stone: the Egyptian Scarab Author(S): William A
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Food Relocation and the Nesting Behavior in Scarabaeus and Kheper (Coleoptera: Scarabaeinae)
ISSN 0065-1737 Acta Zoológica MexicanaActa Zool. (n.s.), Mex. 27(2): (n.s.) 305-324 27(2) (2011) FOOD RELOCATION AND THE NESTING BEHAVIOR IN SCARABAEUS AND KHEPER (COLEOPTERA: SCARABAEINAE) Gonzalo HALFFTER, Violeta HALFFTER & Mario E. FAVILA Instituto de Ecología, A.C., Carretera Antigua a Coatepec No. 351, El Haya, 91070, Xalapa, México. <[email protected]> Halffter, G., V. Halffter & M. E. Favila. 2011. Relocalización de alimento y comportamiento de nidificación en Scarabaeus y Kheper (Coleoptera: Scarabaeinae). Acta Zoológica Mexicana (n. s.), 27(2): 305-324. RESUMEN. En los últimos años se ha avanzado mucho en el conocimiento de las relaciones inter- sexuales y los comportamientos de nidificación de Scarabaeus Linnaeus y Kheper Janseen. En este trabajo, utilizando toda la literatura publicada, aunada a observaciones inéditas, se analizan en detalle las interacciones intrasexuales e intersexuales durante los periodos de alimentación, cópula y nidifica- ción en ambos géneros. En conjunto quedan definidos dos tipos de comportamientos: 1) El que hemos llamado tipo sacer cuyos rasgos más característicos son la ofrenda por el macho de una bola prenupcial que la hembra consume, pero no utiliza para la nidificación; más tarde, con el ovario ya maduro, la hembra puede nidificar sola sin nuevas cópulas ni apoyo del macho. 2) El que se presenta en numerosas especies africanas de Scarabaeus y Kheper, en las que la hembra copula con distintos machos, pero una vez desarrollado el ovario el proceso de nidificación es una actividad conjunta entre un macho y la hem- bra, fabricando la hembra con la bola rodada y enterada una o varias bolas-nido que reciben cuidados postoviposición. -
(Insecta: Coleoptera: Scarabaeidae : Cetoniinae : Tribe, Trichiini)1 Brandon Jones and Andrea Lucky2
EENY-704 Delta Flower Beetle Trigonopeltastes delta (Forster 1771) (Insecta: Coleoptera: Scarabaeidae : Cetoniinae : Tribe, Trichiini)1 Brandon Jones and Andrea Lucky2 Introduction The delta flower beetle, Trigonopeltastes delta (Forster), is a member of the scarab beetle family Scarabaeidae, in the subfamily Cetoniinae. This subfamily is commonly known as flower or fruit chafers because their diet consists mostly of decomposing fruits or pollen (Cave and Ratcliffe 2008). Trigonopeltastes delta belongs to the tribe Trichiini, which contains mostly flower-frequenting species. Although this species is commonly encountered where it occurs, many details of its life cycle and its potential economic importance remain poorly studied. Like many other cetoniines, the delta flower beetle has bright colors and distinctive patterns that distinguish it from other similar species (Figure 1). The delta flower beetle is one of two species in Florida, but while Trigonopeltastes delta is a familiar sight, Trigono- Figure 1. Adult Trigonopeltastes delta (Forster) (dorsal view). peltastes floridana is extremely rare (Woodruff 1960). While Credits: Mike Quinn, TexasEnto.net they are superficially similar, these species are distinguished by distinctive yellow markings on the pronotum. Trigono- Etymology and Synonymy peltastes delta bears a triangular mark whereas Trigono- The name Trigonopeltastes delta is Greek in origin and peltastes floridana has a U- or V-shaped mark. describes the pronotal markings of the species. Trigon translates to triangle, pelt translates to a shield, and delta originates from the letter Δ, or delta. The Greek symbol for delta is a triangle, which resembles the beetle’s pronotal marking, and accounts for its common name. 1. This document is EENY-704, one of a series of the Department of Entomology and Nematology, UF/IFAS Extension. -
Egyptian Quiz
Glasgow Museums Resource Centre Ancient Egypt Quiz Notes Round 1 - Life in Ancient Egypt Question 1 The Nile is the longest river in the world at over 4000 miles long Question 2 Khemet - The Black Land, named after the black tar-like silt that was dredged up when the Nile flooded every year, bringing nutrients to the soil and allowing crops to grow Interesting point - the word ‘Deshret’ is where we get our word for ‘desert’ from Question 3 There are different ways of measuring it, but most scholars will say around 760 hieroglyphs Hieroglyphs can be read up, down, left and right - you look at which way the figures are looking to guide you. The figures will always be looking towards the beginning of the ‘sentence’. Object notes This limestone stela fragment was given to Glasgow Museums by Miss J. May Buchanan in 1912.It preserves parts of six columns of a hieroglyphic inscription separated by vertical lines. The surviving text is part of a hymn to the sun in the form of the god Ra-Horakhty, a combination of the sun-god, Ra, with the falcon god, Horus-of-the-Two-Horizons. The inscription reads: '[Giving praise to Ra]-Hor-akhty, when he goes to his rest in life in this his great mountain […] he says, ‘Hail to you, great one of the Ennead, who comes into being […] in peace to the land of the Western mountain […]' '[… when] he rises in the eastern horizon of the sky, by the servant / deputy […] praise […]' Hymns invoking Amun-Ra-Horakhty do not appear before the 20th Dynasty, suggesting a date of the 20th Dynasty for this fragment. -
Evaluation of Beetles Scarabaeus Sacer Derived- Chitosan, Anti-Cancer and Anti-Bacterial Potentials: in Vitro Study
Vol. 6(1), pp. 1-7, October 2018 DOI: 10.14662/ARJB2018.038 Academic Research Copy © right 2018 Author(s) retain the copyright of this article Journal of Biotechnology ISSN: 2384-616X https://www.academicresearchjournals.org/ARJB/Index.htm Full Length Research Evaluation of beetles Scarabaeus Sacer Derived- Chitosan, Anti-Cancer and Anti-bacterial Potentials: In Vitro Study Amr Abdelkader A.1*, Mohamed Maged1**, Aly Fahmy Mohamed2*** 1,2Faculty of Biotechnology, MSA university 3Head of International Center of Advanced Researches (ICTAR-Egypt) Cairo, Egypt. [email protected]* [email protected]** [email protected] /[email protected]*** Accepted 26 September 2018 Lung and colorectal cancer represent a major health problem all over the world, in addition, bacterial infections impose a serious medical and health concern. Thus, in this study, we evaluate the cytotoxic effect, anti-cancer and anti-bacterial properties of beetles derived chitosan. The study conducted using lung (A549) and colorectal (HCT-116) cancer cell lines to identify the anti-cancer effect and antimicrobial activity against gram-positive bacteria (Streptococcus pyogenes, streptococcus Aureas) and gram- negative bacteria (Salmonella typhimurium). Cytotoxicity was evaluated by describing and measuring recoding morphological changes. The viability and related IC50 were cell type and concentration dependent. Also, related cell apoptosis was monitored using PI stain where early and late apoptosis of treated A549 cells was significantly elevated than in case of HCT-116 cell line (P<0.05). In the meantime, the necrosis % of treated cells didn't perform any changes between the two cell lines but significantly elevated than that of cell control (P<0.05). -
The Beetles Story
NATURE The Beetles story They outshine butterflies and moths in the world of insects and are a delight for their sheer variety—from the brilliantly coloured to the abysmally dull. But they have their uses, too, such as in museums, where flesh-eating beetles are used to clean off skeletons. Text & photographs by GEETHA IYER THE GIRAFFE WEEVIL (Cycnotrachelus flavotuberosus). Weevils are a type of beetle and they are a menace to crops. 67 FRONTLINE . MARCH 31, 2017 HOW was this watery planet we so much love born? Was it created by God or born off the Big Bang? While arguments swing between science and religion, several ancient cultures had different and interesting per- spectives on how the earth came to be. Their ideas about this planet stemmed from their observations of nature. People living in close prox- imity to nature develop a certain sen- sitivity towards living creatures. They have to protect themselves from many of these creatures and at the same time conserve the very envi- ronment that nurtures them. So there is constant observation and in- teraction with nature’s denizens, es- pecially insects, the most proliferate among all animal groups that stalk every step of their lives. The logic for creation thus revolves around differ- ent types of insects, especially the most abundant amongst them: bee- WATER BEETLE. The Cherokees believed that this beetle created the earth. tles. Beetles though much detested (Right) Mehearchus dispar of the family Tenebrionidae. The Eleodes beetle of by modern urban citizens are per- Mexico belongs to this family. ceived quite differently by indige- nous cultures. -
Ivermectin Residues Disrupt Dung Beetle Diversity, Soil Properties and Ecosystem Functioning: an Interdisciplinary field Study
Science of the Total Environment 618 (2018) 219–228 Contents lists available at ScienceDirect Science of the Total Environment journal homepage: www.elsevier.com/locate/scitotenv Ivermectin residues disrupt dung beetle diversity, soil properties and ecosystem functioning: An interdisciplinary field study José R. Verdú a,⁎, Jorge M. Lobo b, Francisco Sánchez-Piñero c, Belén Gallego a, Catherine Numa d, Jean-Pierre Lumaret e, Vieyle Cortez a, Antonio J. Ortiz f,MattiaTonellia, Juan P. García-Teba a, Ana Rey b, Alexandra Rodríguez g, Jorge Durán g a I.U.I. CIBIO, Universidad de Alicante, Alicante E-03690, Spain b Department of Biogeography and Global Change, Museo Nacional de Ciencias Naturales-CSIC, José Abascal 2, Madrid E-28006, Spain c Departamento de Zoología, Universidad de Granada, Granada E-18071, Spain d IUCN-Centre for Mediterranean Cooperation, Marie Curie 22, Campanillas, Málaga E-29590, Spain e UMR 5175 CEFE, CNRS - Université de Montpellier - Université Paul-Valéry Montpellier 3 – EPHE, Université Paul-Valéry Laboratoire Zoogéographie, Route de Mende, 34199 cedex 5 Montpellier, France f Departamento de Química Inorgánica y Química Orgánica, Universidad de Jaén, Campus Las Lagunillas, Jaén E-23071, Spain g Center for Functional Ecology (CEF), Department of Life Sciences, University of Coimbra, Calçada Martim de Freitas, 3000-456 Coimbra, Portugal HIGHLIGHTS GRAPHICAL ABSTRACT • At the short term, ivermectin residues cause a strong decrease in dung reloca- tion and dung spreading by dung beetles. • Conventional use of ivermectin disrupts diversity by affecting species richness, abundance and biomass of dung beetles. • Reduction in the functional efficiency of dung degradation resulted in the long- term accumulation of manure. -
The Importance of Ecological Memory for Trophic Rewilding As an Ecosystem Restoration Approach
Biol. Rev. (2019), 94,pp.1–15. 1 doi: 10.1111/brv.12432 The importance of ecological memory for trophic rewilding as an ecosystem restoration approach 1,2,4 1,3 1,2 Andreas H. Schweiger ∗ , Isabelle Boulangeat , Timo Conradi , Matt Davis1,4 and Jens-Christian Svenning1,4 1Section for Ecoinformatics and Biodiversity, Department of Bioscience, Aarhus University, Ny Munkegade 114, 8000, Aarhus C, Denmark 2Plant Ecology, Bayreuth Center for Ecology and Environmental Research (BayCEER), University of Bayreuth, 95440, Bayreuth, Germany 3University Grenoble Alpes, Irstea, UR LESSEM, 2 rue de la Papeterie-BP 76, F-38402, St-Martin-d’H`eres, France 4Center for Biodiversity Dynamics in a Changing World (BIOCHANGE), Aarhus University, Ny Munkegade 114, 8000, Aarhus C, Denmark ABSTRACT Increasing human pressure on strongly defaunated ecosystems is characteristic of the Anthropocene and calls for proactive restoration approaches that promote self-sustaining, functioning ecosystems. However, the suitability of novel restoration concepts such as trophic rewilding is still under discussion given fragmentary empirical data and limited theory development. Here, we develop a theoretical framework that integrates the concept of ‘ecological memory’ into trophic rewilding. The ecological memory of an ecosystem is defined as an ecosystem’s accumulated abiotic and biotic material and information legacies from past dynamics. By summarising existing knowledge about the ecological effects of megafauna extinction and rewilding across a large range of spatial and temporal scales, we identify two key drivers of ecosystem responses to trophic rewilding: (i) impact potential of (re)introduced megafauna, and (ii) ecological memory characterising the focal ecosystem. The impact potential of (re)introduced megafauna species can be estimated from species properties such as lifetime per capita engineering capacity, population density, home range size and niche overlap with resident species. -
Dung Beetles (Coleoptera: Scarabaeidae: Aphodiinae) of the Mpala Research Centre and Environs, Laikipia District, Kenya
University of Nebraska - Lincoln DigitalCommons@University of Nebraska - Lincoln Center for Systematic Entomology, Gainesville, Insecta Mundi Florida March 2008 Dung beetles (Coleoptera: Scarabaeidae: Aphodiinae) of the Mpala Research Centre and environs, Laikipia District, Kenya Robert D. Gordon Northern Plains Entomology Patrice Bordat Lagasse, F-82300 Saint-Cirq, France Giovanni Dellacasa Genova, Italy Marco Dellacasa Università di Pisa, via Roma Follow this and additional works at: https://digitalcommons.unl.edu/insectamundi Part of the Entomology Commons Gordon, Robert D.; Bordat, Patrice; Dellacasa, Giovanni; and Dellacasa, Marco, "Dung beetles (Coleoptera: Scarabaeidae: Aphodiinae) of the Mpala Research Centre and environs, Laikipia District, Kenya" (2008). Insecta Mundi. 119. https://digitalcommons.unl.edu/insectamundi/119 This Article is brought to you for free and open access by the Center for Systematic Entomology, Gainesville, Florida at DigitalCommons@University of Nebraska - Lincoln. It has been accepted for inclusion in Insecta Mundi by an authorized administrator of DigitalCommons@University of Nebraska - Lincoln. INSECTA MUNDI A Journal of World Insect Systematics 0028 Dung beetles (Coleoptera: Scarabaeidae: Aphodiinae) of the Mpala Research Centre and environs, Laikipia District, Kenya Robert D. Gordon Northern Plains Entomology P. O. Box, 65 Willow City, ND 58384 USA Patrice Bordat Lagasse, F-82300 Saint-Cirq, France Giovanni Dellacasa C.P. 921, I-1612 Genova, Italy Marco Dellacasa Centro Interdipartimentale, Museo di Storia Naturalle e del Territorio Università di Pisa, via Roma 79, I-56011 Calci (Pisa) Italy Date of Issue: March 3, 2008 CENTER FOR SYSTEMATIC ENTOMOLOGY, INC., Gainesville, FL Robert D. Gordon, Patrice Bordat, Giovanni Dellacasa, Marco Dellacasa Dung beetles (Coleoptera: Scarabaeidae: Aphodiinae) of the Mpala Research Centre and environs, Laikipia District, Kenya Insecta Mundi 0028: 1-15 Published in 2008 by Center for Systematic Entomology, Inc. -
Dung-Beetles-FS-NYSIPM.Pdf (752.1Kb)
From the NYSIPM Publications collection: hdl.handle.net/1813/41246 2020 New York State Integrated Pest Management Cornell Cooperative Extension Program FIELD CROPS Dung Beetles Aid in Reducing Flies and Gastrointestinal Parasites in Pastures Ken Wise1, Michael Baker2, and Jaime Cummings1 1New York State Integrated Pest Management Program, Cornell University 2Department of Animal Science, Cornell University Dung beetles are important insects for pasture ecology and There are three categories of dung beetles. soil health. They move manure into the soil, thus increasing Rollers (telecoprids) organic matter, improving soil structure, increasing water infiltration and providing essential nutrients for grass growth. Geotrupes species form balls of manure which they push from As the majority of cow/calf and stocker operations are pasture the pat to bury as brood balls. This group of insects comprise based, and with the increased interest in finishing beef on nesting species. The male and female beetles work together to grass, and grass-based dairies, there has been renewed interest bury the brood ball to feed their young. surrounding the importance of dung beetles in pasture Dwellers (endocoprids) ecology. There are hundreds of organisms that call a manure pat home. Some organisms are not beneficial, but many of Aphodius species consume the manure as they tunnel within them are. Dung beetles are in the Scarab beetle insect family, the dung pat and lay eggs directly in the manure or surrounding and are known for their digging abilities. Dung beetles exist soil. Most dung beetles found in New York are dwellers. everywhere there is fecal matter. The beetles are attracted by Tunnelers (paracoprids) the smell, and can find a new cowpat within seconds. -
Contents Introduction to the Touch Tour
Contents Introduction to the Touch Tour................................2 Description and plan of Room 4 ............................4 1. Seated statue of Amenhotep III .........................6 2. Lion statue of Amenhotep III ..............................8 3. Sarcophagus ...................................................... 10 4. Statue of King Senwosret III ............................11 5. Left arm from a colossal statue of Amenhotep III .......................................................... 13 6. Seated statue of the goddess Sekhmet ........14 7. Block statue of Amenhotep ............................. 16 8. Boat sculpture of Queen Mutemwia ............. 18 9. Colossal scarab beetle .................................... 20 1 Introduction to the Touch Tour This tour of the Egyptian Sculpture Gallery is a specially designed Touch Tour for visitors with sight difficulties. This guide gives you information about nine highlight objects in Room 4 that you are able to explore by touch. The Touch Tour is also available to download as an audio guide from the Museum’s website: britishmuseum.org/egyptiantouchtour If you require assistance, please ask the staff on the Information Desk in the Great Court to accompany you to the start of the tour. The sculptures are arranged broadly chronologically, and if you follow the tour sequentially, you will work your way gradually from one end of the gallery to the other moving through time. Each sculpture on your tour has a Touch Tour symbol beside it and a number. 2 Some of the sculptures are very large so it may be possible only to feel part of them and/or you may have to move around the sculpture to feel more of it. If you have any questions or problems, do not hesitate to ask a member of staff. -
Ancient Egypt Handling Box
Ancient Egypt Handling Box Ancient Egypt handling collection Teacher Notes Welcome to the National Museum of Scotland. This box contains handling objects and resources designed to support learning about ancient Egypt. It contains 33 real and replica objects relating to the ancient Egyptian collections of National Museums Scotland. The objects are grouped into four themes: 1. Life 2. Mummification 3. Afterlife 4. Egyptology This Teachers Resource Pack for the Ancient Egypt handling collection is also available to download from our website: www.nms.ac.uk/nmosvisit#handlingbox National Museums Scotland Teachers’ Resource Pack Ancient Egypt handling collection Introduction The ancient Egyptian collections of National Our ancient Egyptian collections were begun Museums Scotland comprises around in 1819, and many of the objects derive from 6,000 items, including many unique and archaeological excavations by Alexander Henry internationally significant objects, such as Rhind (Fellow of the Society of Antiquaries royal statuary, reliefs, mummies, coffins, papyri, Scotland), the Egypt Exploration Society, British furniture, jewellery and textiles. School of Archaeology in Egypt, and Oxford Ancient Egypt was one of the earliest and and Liverpool Universities. The museum also longest-lived civilisations, spanning almost has the only intact ancient Egyptian royal burial 4000 years of history. The River Nile and the group outside of Egypt. surrounding desert shaped ancient Egyptian The ancient Egyptian collections of National culture and how they saw the world. Egypt’s Museums Scotland are represented in this success derived from its natural resources, such informative and interactive learning resource. as fertile agriculture from the annual flood, Use the handling objects to create an exciting stone for building monuments, and precious and educational experience for your group and gold. -
17 the Bull Hunt Scarab of Amenhotep III
17 The Bull Hunt Scarab of Amenhotep III He lunged from side to side Watching for his chance to seize the horns. The bull frothed in its rage at this dance And suddenly Enkidu seized its tail And twisted it around, until the bull Stood still, bewildered, out of breath, And then Enkidu plunged his sword behind its horns Into the nape of the bull’s neck, and it fell dead. Gilgamesh, translated by Herbert Mason Hsbt 2 xr Hm n anx Hrw kA nxt xa m mAat nbti smn hpw sgrH tAwi Hrw-nbw aA xpS Hw sstiw (n)sw(t)-bit(i)nb tAwi NB-MAa&-Ra zA ra IMN-@&P HqA-wAst di anx Hmt-(n)sw(t) wrt Tiy anx.ti biAit xprt n Hm.f iw(t).tw r Dd n Hm.f iw wn smAw Hr xAst nt w n St(a) nat Hm.f m xd m wiA-(n)sw(t) xa-m-mAat Hr tr n xAwi Szp tp-wAt nfrt spr m Htp r w n St(a) Hr tr n dwA xat Hm.f Hr ssmt mSa.f tm m xt.f sHnt srw mSa anxw nw mSa r Dr.f mi qd.f Xrdw n kAp r irt rsw Hr nA n smAw ist wD.n Hm.f rdit itH.tw nn smAw m sbti Hna Sdy wD.in Hm.f r nn smAw r Aw.sn rxt iri smAw 170 rxt in.n Hm.f m bHs m hrw pn smAw 56 wAH.in Hm.f hrw 4 m wS rdit srf n ssmwt.f Xat Hm.f Hr ssmt rxt nn smAw in.n.f bHs smAw 40 tmt smAw 96 The Bull Hunt Scarab of Amenhotep III.