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ISSN 0375-0183 Indexed in Current Contents, Science Citation Index and Research Alert , and covered by most of the major abstracting services ODONATOLOGICA JOURNAL OF THE SOCIETAS INTER- NATIONALIS ODONATO- LOGICA S.I.O. Odonatologica Vol. 42 No. 4 pp. 285-448 December 1, 2013 ODONATOLOGICA publishes original papers in all elds of odonatology. It is a quarterly, published for the International Odonatological Foundation, SOCIETAS INTERNATIONALIS ODONATO- LOGICA (S.I.O.). It is general policy that submitted papers will be refereed. EXECUTIVE EDITOR B. KIAUTA (Bergen/LB, The Netherlands) ASSISTANT EDITORS M. KIAUTA (Bergen/LB, The Netherlands) G. KIAUTA THUCYDIDES (Vancouver, Canada) ASSOCIATE EDITORS M. HÄMÄLÄINEN (Espoo, Finland) P.J. MILL (Leeds, UK) K. INOUE (Osaka, Japan) EDITORIAL BOARD R.J. BECKEMEYER (Wichita/KS, USA) D.R. PAULSON (Tacoma/WA, USA) M. BEDJANIČ (Braslovče, Slovenia) O.N. POPOVA (Novosibirsk, Russia) R.A. CANNINGS (Victoria/BC, Canada) M.J. SAMWAYS (Matieland, SA) A. CÓRDOBA-AGUILAR (Mexico, Mexico) K. SUZUKI (Toyama, Japan) H.J. DUMONT (Gent, Belgium) G. THEISCHINGER (Lidcombe/NSW, Australia) S.W. DUNKLE (Plano/TX, USA) D.J. THOMPSON (Liverpool, UK) R.W. GARRISON (Azusa/CA, USA) C. UTZERI (Roma, Italy) G. JACQUEMIN (Nancy, France) G.S. VICK (Tadley/Hants, UK) R.G. KEMP (Wolverhampton, UK) M. WATANABE (Tsukuba, Japan) O.E. KOSTERIN (Novosibirsk, Russia) H. WILDERMUTH (Rüti, Switzerland) A.B.M. MACHADO (Belo Horizonte, Brazil) K.D.P. WILSON (Brighton, UK) A. MARTENS (Karlsruhe, Germany) ODONATOLOGICAL ABSTRACTS B. KIAUTA (Bergen/LB, The Netherlands) K. INOUE (Osaka, Japan) R.J. ANDREW (Nagpur, India) W. PIPER (Hamburg, Germany) P. BUCZY ŃSKI (Lublin, Poland) C. UTZERI (Roma, Italy) EDITORIAL ADDRESS: Odonatologica Editorial Ofce, P.O. Box 124, NL-5854 ZJ Bergen/LB, The Netherlands, or the address of any Associate Editor, or a member of the Editorial Board. For the addresses see the last page in Notulae odonatologicae. E-mail: [email protected] WEBSITE: www.odonatologica.com SUBSCRIPTION ORDERS should be addressed to the Odonatologica Editorial Office, c/o Mrs M. Kiauta, P.O. Box 124, NL-5854 ZJ Bergen/LB, The Netherlands, – or to the Rep- resentatives in I t a l y (Dr C. Utzeri, Dipto Biol. Anim. & Uomo, Univ. Roma “La Sa- pienza”, Viale dell’Università 32, I-00185 Roma); – J a p a n (K. Inoue, 5-9, Fuminosato 4-chome, Abeno-ku, Osaka, 545-0004); – R u s s i a (Dr O.N. Popova, Inst. Anim. Syst. & Ecol., Siberian Branch, Russ. Acad. Sci., 11 Ul. Frunze, RUS-630091 Novosibirsk); – S l o v e n i a (Dr A. Pirnat, Groharjeva 18, SI-1241 Kamnik); – U n i t e d K i n g d o m (G.S. Vick, Crosselds, Little London, Tadley, Hants, RG26 5ET); – U n i t e d S t a t e s (B. Mauffray, Int. Odon. Res. Inst., D.P.I., P.O. Box 147100, Gainesville, FL 32614-7100). CANCELLATION OF SUBSCRIPTION for the forthcoming year must reach the Editorial Ofce prior to December 1. Cancellations for the current year cannot be considered. ELECTRONIC AVAILABILITY OF THE JOURNAL: all paid subscribers may register at the web- site, which involves entering the subscriber's details, and choosing a user name and password. After conrmation of subscription status has been completed, full access to the online journals will be made available for the subscription period. THE 2015 INTERNATIONAL CONGRESS OF ODONATOLOGY will be convened in La Plata (Argentina). Organizing team: Dr J. Muzon [[email protected]] & Dr F. Lozano [federicolozano82@ gmail.com]. Mailing address: ILPLA-CONICET, cc 712, AR-1900 La Plata, Argentina. THE 2017 INTERNATIONAL CONGRESS OF ODONATOLOGY will be convened in Algeria. Organizing Secretary: Dr B. Samraoui [[email protected]]. Price per volume 2013: € 220.— (postage incl.). Special rates for individual S.I.O. associates. Please inquire! © International Odonatological Foundation S.I.O., Bergen/LB (including the Odonatological Abstracts) Odonatologica 42(4): 285-292 December 1, 2013 EDITORIAL AT THE CONCLUSION OF FOUR DECADES OF EDITORSHIP (1971-2013) In October 1971, I was entrusted by the participants of the First European Sym- posium of Odonatology (Ghent, Belgium) with the organisation and editorship of ODONATOLOGICA. This was to appear quarterly and include, in addition to research papers, the abstracts of contemporary odonatological literature published elsewhere. With the present issue, the 42 nd volume is completed: the 168 issues contain about 1400 research papers and over 19,800 abstracts of works published through- out the world since 1971. The editing and the intensive daily contacts with the odonatological community became my life style, bringing much pleasure and many personal friendships. It was not easy, and with much sorrow and regret, that I had to decide to resign from the of!ce upon the completion of the current volume: the advancing age is being increas- ingly felt and my health condition is not optimal any more. I trust the world odonatological community will share my feelings of thanks to Dr Florian WEIHRAUCH (Jägerstrasse 21/A, D-85283 Wolnzach, Germany; − [email protected]) for his acceptance of the responsibilities for the journal, its administration, production, editorship and circulation as from the 43 rd volume onwards. With his profound experience in worldwide scienti!c communica- tion and odonatological editorship, he is perfectly quali!ed to navigate the journal on the waves of contemporary scienti!c requirements and standards and to guide it to an increasingly high scienti!c level. Since the birth of ODONATOLOGICA , two people have been crucial for its de- velopment: without the support and much more than the manifold help from the late Professor Dr Janny M. van BRINK (Utrecht/Bilthoven, The Netherlands) and my wife, MARIANNE, it would have been unthinkable to continue in the of!ce for so long. For some decades, Marianne has been the backbone of the SIO and ODO- NATOLOGICA , taking care of all the technical matters and much more. Likewise, since the very !rst issue until today, the editorial help rendered by Dr Peter J. MILL (Leeds, UK) has been indispensable. There is hardly an issue in the production of which he has not been involved, either as a peer reviewer, style model- 286 B. Kiauta ler or a language corrector. Kiyoshi INOUE’s (Osaka, Japan) and Professor Dr Minter J. WESTFALL’s (Gainesville/FL, USA) manifold help and continuous encouragement during the four decades were among the cor- nerstones of much of my work. In recent years, my son, Adjunct Professor Dr George KIAUTA- -THUCYDIDES (Vancouver, Cana- da) opened new horizon by managing the website and the electronic edition of the journal. For this I thank him 1 very sincerely. At its very onset, ODONATO- LOGICA enjoyed the active support and assistance of well over a hundred odonatologists from all parts of the world. From among those deceased, at least a few peer reviewers, collab- orators of the Odonatological Ab- stracts section and/or helpers in other editorial matters should be recalled 2 here, viz. [academic titles omitted]: N.N. AKRAMOWSKI (Armenia), S. ASAHINA (Japan), B.I. BAL- INSKY (Sth Africa), J. BELLE (Netherlands), B.F. BELYSHEV (Russia), G.H. BICK (USA), F. CAPRA (Italy), H.-f. CHAO (Chi- na), P.S. CORBET (UK), D.A.L. DAVIES (UK), R.M. GAMBLES (UK), D.C. GEIJSKES (Nether- 3 lands), L.K. GLOYD (USA), A.Yu HARITONOV (Russia), H. KAI- Figs 1-3. Charter Meeting of the Societas interna- tionalis odonatologica (SIO), held during the First SER (Germany), M.A. LIEFT- European Symposium of Odonatology, Ghent, Bel- INCK (Netherlands), C. LONG- gium, 22-23 October 1971, Dr B. Kiauta in Chair: FIELD (Eire), T.T. MACAN (UK), (1) the birth of SIO (!rst row, from left to right: C. S. MIELEWCZYK (Poland), Fischer, Dipl.-Ing. H. Schumann, Dr. J.M. van Brink P.L. MILLER (UK), T.R. MI- and Dr G. Jurzitza); − (2) Dr F. Schaller, proposing the name ‘ Odonatologica ’ for the Society’s journal; TRA (India), E.B. MONTGOM- − (3) discussion on the set-up of the !rst Editorial ERY (USA), P. MÜNCHBERG Board. − [Photographs by D. Smit, Utrecht] (Germany), C. NIELSEN (Italy), Editorial at the conclusion of editorship 287 E.C.G. PINHEY (Rhodesia/UK), A. PORTMANN (Switzerland), G. PRITCHARD (Canada), Z. SPURIS (Latvia), V. TEYROVS- KÝ (Czech Republic), J.A.L. WAT- SON (Australia) and M.J. WEST- FALL (USA). My warm thanks go to the dozens of external peer reviewers and to the past and present members of the Ed- 4 itorial Board, as listed on p. 2 of the cover of each issue. Some of these are currently the sole (or almost so) liv- ing experts in certain narrow elds of odonatology; hence their help remains indispensable. A journal is made by the contrib- uting authors and it is to them I owe profound thanks for the high standard and the impact ODONATOLOGI- 5 CA enjoys, the latter despite its nar- Figs 4-5. Charter Meeting of SIO: (4) Dr M.A. Lieft- row restriction to a very small group inck, one of the four SIO Members of Honour, elect- of organisms. It is also due to them ed at the Charter Meeting; the others were Dr S. that the journal has been in a posi- Asahina (Japan), Dr B.F. Belyshev (USSR) and Dr tion to adhere strictly to the publica- E.C.G. Pinhey (Rhodesia); − (5) Some participants: tion dates, as set-up in 1971. 1: Dr J. Olesen, −2: Dr F. Schaller, −3: Dr A.M. − − rd Moens-Bullens, 4: Dr J. Moens, 5: Dr M. Mouze, The 3 issue of Volume 2 appeared −6: J.C. Andries, −7/8: Mrs & Mr H. Itzerott, −9: in time solely due to M.J.
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  • Neoneura, Species, Neoneura Jurzitzai

    Neoneura, Species, Neoneura Jurzitzai

    Odonatologica 28(4): 343-375 December I, 1999 The genus Neoneura, with keys and description of a new species, Neoneurajurzitzai spec. nov. (Zygoptera: Protoneuridae) R.W. Garrison Research Associate, Natural History Museum of Los Angeles County, 900 Exposition Blvd., Los Angeles, CA 90007, United States Received April 10, 1999 / Reviewed and Accepted May 11, 1999 A all both synopsis of 23 spp. includes keys to sexes, based primarily on caudal appendage morphology in males and morphology of the hind lobe of the prothorax in females, diagnoses, distributional notes and diagnostic illustrations. N. jurzitzai sp.n. (holotype 6: Brazil, Santa Catarina state, Nova Teutonia, 5 XI-I942, in UMMZ) is described, and the 6 and 9 ofN. rufithorax Selys are described based on specimens from Peru. INTRODUCTION include The genus Neoneura was first proposed by SELYS (1860) to two new species, N. rubriventris and N. bilinearis. Later, SELYS (1886) expanded the ge- nus to include eight species, of which four were new. CALVERT (1903, 1907) described three species (N. aaroni, N. amelia, and N. paya) from southern Texas, Mexico, and Central America. WILLIAMSON (1917), in his seminal revision monographed all known species, described six new species, and included diagnos- tic illustrationsof all known species. It is the only paper which attempted to key all work is valuable for Williamson’s species. Besides the illustrations, the attempt to determine the identity of some of the poorly known Selysian species. Although Williamsonkeyed the males for all species, he did not have material ofN. rufilhorax Selys or N. waltheri Selys. At about the same time, KENNEDY (1917) illustrated the penes of all known species.
  • A Checklist of North American Odonata, 2021 1 Each Species Entry in the Checklist Is a Paragraph In- Table 2

    A Checklist of North American Odonata, 2021 1 Each Species Entry in the Checklist Is a Paragraph In- Table 2

    A Checklist of North American Odonata Including English Name, Etymology, Type Locality, and Distribution Dennis R. Paulson and Sidney W. Dunkle 2021 Edition (updated 12 February 2021) A Checklist of North American Odonata Including English Name, Etymology, Type Locality, and Distribution 2021 Edition (updated 12 February 2021) Dennis R. Paulson1 and Sidney W. Dunkle2 Originally published as Occasional Paper No. 56, Slater Museum of Natural History, University of Puget Sound, June 1999; completely revised March 2009; updated February 2011, February 2012, October 2016, November 2018, and February 2021. Copyright © 2021 Dennis R. Paulson and Sidney W. Dunkle 2009, 2011, 2012, 2016, 2018, and 2021 editions published by Jim Johnson Cover photo: Male Calopteryx aequabilis, River Jewelwing, from Crab Creek, Grant County, Washington, 27 May 2020. Photo by Netta Smith. 1 1724 NE 98th Street, Seattle, WA 98115 2 8030 Lakeside Parkway, Apt. 8208, Tucson, AZ 85730 ABSTRACT The checklist includes all 471 species of North American Odonata (Canada and the continental United States) considered valid at this time. For each species the original citation, English name, type locality, etymology of both scientific and English names, and approximate distribution are given. Literature citations for original descriptions of all species are given in the appended list of references. INTRODUCTION We publish this as the most comprehensive checklist Table 1. The families of North American Odonata, of all of the North American Odonata. Muttkowski with number of species. (1910) and Needham and Heywood (1929) are long out of date. The Anisoptera and Zygoptera were cov- Family Genera Species ered by Needham, Westfall, and May (2014) and West- fall and May (2006), respectively.