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Dissertaciya Nekipelova Elena « » ______ _____________ , « _____» ________ 2019 . МИК ИЦИ 06.03.01- . , ... .. , , , _________ .. , , ..., ... .. , , , 2019 ..................................................................................................................3 1. ...............................................................................................7 1.1- ( . . ) 7 1.2 10 2. .25 3. …………………………………………………….35 3.1 …………………………………….35 3.2 ……………………………………………………..40 3.3 . …………………44 3.4 ……………………………………………………………45 3.5 50 3.6 52 4. ………………………………………………………………………..54 5. ……………………………………..…..56 2 Thymallus. , Thymallus arcticus Pallas. , Thymallus arcticus , [58]. , , , , , , . , , . (1936) , . , , – [40]. 21 , . .. (2009) 3 – , . . (. , 2009) [29], . 2002 , [72]. 2009 - Thymallus svetovidovi (Knizhin et Weiss, 2009) – . 3 , . [29]. , . , , , , , [40]. , , , , [70]. , ( , ), , [22]. (, , ) ; , , , [65]. , -, [15,70]. [22]. , , , , 4 , [70]. , , - – . . , [50]. , . , , , , . . - , . 5 : , - . : - ; - . ; - . ; - - ; - , ; - ; 6 1. 1.1 - . . – , , . . . , . . . , – . – . 4 . . – . , . - , – . , , , . , . . 7 , [32]. 40-50 . , , , , . , 8. , 0,5 ., , . - 13-14 . , 0,7-1,3. . 1– ( . 64 - ), 2018 ( ) 8 : - Coregonus tugun (Pallas, 1814); Lota lota (Linnaeus, 1758); Lethenteron kesslery (Anikin, 1905); - Esox lucius (Linnaeus, 1758); Rutilus rutilus lacustris (Pallas, 1814); Leuciscus leuciscus baicalensis (Dybowski, 1874); Perca fluviatilis (Linnaeus, 1758); Gymnocephalus cernuus (Linnaeus, 1758); - Gobio gobio cynocephalus (Dybowski, 1869); - Thymallus arcticus (Pallas, 1776); Brachymystax lenok (Pallas, 1773); Hucho taimen (Pallas, 1773); Oncorhynchus mykiss irideus (Walbaum, 1792); Paracottus knerii (Dybowski, 1874); Phoxinus phoxinus (Linnaeus, 1758); - - Abramis brama (Linnaeus, 1758) [4], [7],[60]. , « ». ; , , - [52]. 9 - , . [49] . 2009 68 10 . (Insecta), (. Diptera) – 38 , (. Trichoptera) – 5, (. Ephemeroptera) – 3, (. Coleoptera), (. Heteroptera) – . (. Oligochaeta) – 10 , (. Gastropoda, . Bivalvia) - 4, (. Crustacea) – 3, (. Hirudinea) – 2 . 1.2 чя х х Salmoniformes - Salmonidae - Thymallus - Thymallus articus (Pallas, 1776) - [68]. , . , [29]. , . 400 730.. [5]. , – Thymallus arcticus, 10 . , .. (1936) . (1948) [49]. , ( ) , [58]. , , , , . : , [22]. Thymallus Cuvier, 1829 [24]: 1. Thymallus arcticus (Pallas, 1776) - ; 2. Thymallus baicalensis Dybowski, 1874 - ; 2.1. Thymallus baicalensis nigrescens Dorogostaisky, 1923 - () . 3. Thymallus burejensis Antonov, 2004 - [3]; 4. Thymallus brevicephallus Mitrofanov, 1961 - ; 5. Thymallus brevirostris Kessler, 1879 - ; 6. Thymallus grubii Dybowski, 1869 - ; 6.1. Thymallus grubii grubii Dybowski, 1869 - ; 6.2. Thymallus g. flavimaculatus Khnizhin et al., 2006 - [27]; 7. Thymallus nikolskyii Kaschenko, 1899 - ; 8. Thymallus svetovidovi Khnizhin et al., 2009 - ; 9. Thymallus thymallus (Linnaeus, 1758) - ; 10. Thymallus tugarinae Khnizhin et al. , 2007 – [16]. 11 . , . , , . - -b 9 - . 2,52 – 3,10% -b, 3,38 – 3,92% - , – [53]. , 25 , , , . () (). . [51,52]. , , [21,49,54]. T.grubii grubii – [32],T.grubii flavomaculatus – [70], T. burejensis – [3], T. tugarinae sp. nova – ( ) [71]. .. (Pallas, 1776) — . Salmo arcticus [5]. 1936 .. «- », 12 , : Th. brevirostris Kessler, 1879; Th. nigrescens Dorogostaski, 1923; Th. thymallus (Linne, 1758) Th. arcticus (Pallas, 1776). .. : Th. arcticus pallasi Valenciennes, 1848; Th. arcticus grubei Dybowski, 1869; Th. arcticus grube inatio mertensii Valenciennes, 1848; Th. arcticus baicalensis Dybowski, 1874; Th. arcticus baicalensis infra sub species brevipinnis Svetovidov, 1930 (, 1999). , , .. (1936) , . , , – [58]. .. (1950) , Thymallus , (Th. thymallus, Th. arcticus, Th. nigrescens, Th. brevirostris) , (Th. arcticus signifer, Th. montanus). Th. thymallus – , : , , , , , , , , . (Th. arcticus), , . – , , . : . , , , . 13 , , , , . () , , , , , . (Th. nigrescens ), . (Th. brevirostris), - [44]. (Thymallus arcticus) - Th. arc. pallasii, Th. arc. mertensii Th. arc. signifer [24]. .. Thymallus : Th. thymallus, Th. brevirostris Th. arcticus, : Th. arcticus baicalensis, Th. arcticus pallasi, Th. arcticus grubei [38]. .. [24] , . - 5 : Th. arcticus, Th. baicalensis, Th. baicalensis nigrescens, Thymallus sp. Th. svetovidovi . [29] , . , [30]. .. « (Thymallus) » (2011) , 14 , . , . : Th. arcticus, Th. baicalensis, Th. svetovidovi. , , : , -, , , , , [53]. [11]. . , , . . , , , . , 3-4 , V- [7]. . . , . , , . . , - [7]. . , . , . , . 15 , . , , . , . , . - . . , . - 50 . 1.5 , 3 . , 14 , 51 , 1665 . [60]. [43,48]. [8]. [2]. , , [17]. [36]. , , , [45]. () , , [33] [29], [31]. , , , 16 [37]. () : , , [20]. , , , , [13]. : [37,20]. . 2 5 , 12 [70]. - , , "" [17]. : ( - ), ( - ), ( -). , [64]. 5-11 , 6-9 . , [11]. .
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