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Zach Griebenow

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Zachary Griebenow PhD Candidate, Ward Laboratory, Dept. of Entomology & Nematology One Shields Avenue, Davis, CA 95616 502-902-7290, [email protected] EDUCATION Doctor of Philosophy, Entomology, the University of California-Davis, September 2017-present • Qualifying Exam passed December 2, 2019 B.S. Agriculture, Major: Entomology, The Ohio State University, August 2013-May 2017, magna cum laude • Minor: Music Homeschooled PROFESSIONAL EXPERIENCE Graduate Research Ward Lab, the University of California-Davis, September 2017-present • Advisor: Philip Ward o Thesis: systematic revision of the Leptanillinae (Hymenoptera: Formicidae) jointly based upon 1) phylogenomic and 2) male morphological data . Phylogenomic inference: • Acquired genome-scale data from leptanilline morphospecies using the ultra-conserved element (UCE) probe set hym-v2 (Branstetter et al. 2017) and inferred phylogeny of all UCE-enriched taxa using IQ-Tree ver. 1.6.10 or ExaBayes ver. 1.5 • Inferred partition schemes for phylogenomic data using SWSC algorithm of PartitionUCE (Tagliacollo & Lanfear 2018) and the best-fitting substitution models thereof with PartitionFinder2 (PF2) ver. 2.1.1 or ModelFinder • Using RevBayes ver. 1.0.9-10, inferred phylogeny of male leptanilline morphotaxa jointly from 11 nuclear loci (partitioned using PF2 v. 2.1.1) and de novo matrices of binary morphological characters o Nuclear loci recovered from UCE-based dataset and aligned on a by-locus basis using online MAFFT interface o Modelled morphological substitution using Mk/v+Γ models with accommodation of asymmetrical state frequencies . Morphological techniques: 1 Zachary Griebenow, C.V. • Imaged external morphological characters of select leptanilline males with Hitachi desktop scanning electron microscope, non-coating • Acquired morphometric data and imaged specimens for upload to AntWeb using AutoMontage program • Segmented micro-computed tomographic (μ-CT) scans of male leptanilline genitalia using ITK-SNAP ver. 3.6.0, Dragonfly and β version of Biomedisa Independent Undergraduate Research with Distinction Rothenbuhler Bee Lab, The Ohio State University, September 2015-August 2017 • Conducted research project on morphometric parameters of Caribbean Heterotermes spp. (Blattaria: Termitoidae: Rhinotermitidae: Heterotermitinae) • Imaged specimens using AutoMontage program • Measured morphological indices across multiple specimens • Calculated comparative morphometric indices utilizing Microsoft Excel • Analyzed morphometrics using SPSS Research Assistant Triplehorn Insect Collection, The Ohio State University, September-December 2014, March-April 2015 • Converted preexisting dataset originating in Ohio Odonata Survey to format necessary for entry in OSU’s Hymenoptera Online Database • Located geographical coordinates for collection localities using United States Geological Survey and/or Google Earth • Modeled probable distributions for selected Aeshnidae upon the basis of collection data using Maxent software Curatorial Assistant Triplehorn Insect Collection, The Ohio State University, September 2013-April 2014, June-July 2016, January-July 2017 • Labeled and databased specimens • Mounted microhymenopterans on cardstock points • Identified specimens to familial level or lower • Helped reorganization of Coleoptera collection 2 Zachary Griebenow, C.V. HONORS • Received the Ernst Mayr Travel Grant from the Museum of Comparative Zoology, May 2020 • Member of the University of California-Davis’ Linnaean Games team (2018- present), winner of 1st place in the Entomological Society of America’s (ESA) 2018 national championship • Member of The Ohio State University’s Linnaean Games team (2017), winner of 2nd place in ESA national championship • Placed 2nd in undergraduate category at the Ohio Entomological Association’s Research Forum, presenting systematic research in the termite genus Heterotermes (Blattaria: Rhinotermitidae); November 4th, 2016 • Received Research Scholar Award ($1,000) to aid undergraduate research project; August 2016 • Dean’s List, College of Food, Agricultural, & Environmental Sciences (The Ohio State University; Spring 2014, Spring 2015, Spring 2016, Autumn 2016, Spring 2017 PROFESSIONAL PRESENTATIONS • Griebenow, Z. H. (2020). Total-evidence Bayesian Inference and the phylogeny of aberrant ants (Formicidae: Leptanillinae). On-demand presentation at the virtual annual meeting of the Entomological Society of America. • Griebenow, Z. H., Fischer, G., & Economo, E. (2019). Total-evidence analysis of male Leptanillinae (Hymenoptera: Formicidae) clarifies phylogeny of male-based taxon Phaulomyrma. Presentation at the annual meeting of the Entomological Society of America, November 18, St. Louis, MO. • Griebenow, Z. H., Fischer, G., & Economo, E. (2019). Preliminary characterization of male morphology in the Leptanillinae (Hymenoptera: Formicidae) for phylogenetic inference. Presentation at the annual North American meeting of the International Society of Hymenopterists, November 17, St. Louis, MO. • Griebenow, Z. H., Fischer, G., & Economo, E. (2018). Preliminary characterization of male morphology in the Leptanillinae (Hymenoptera: Formicidae) for phylogenetic inference. Presentation at the annual meeting of the Entomological Society of America, November 6, Vancouver, BC. • Griebenow, Z. H., Jones, S. C., & Eaton, T. D. (2017). Revision of Diagnostic Morphological Traits in Three Caribbean Species of Heterotermes (Dictyoptera: Rhinotermitidae). Virtual poster at the annual meeting of the Entomological Society of America, Denver, CO. 3 Zachary Griebenow, C.V. • Griebenow, Z. H., Jones, S. C., & Eaton, T. D. (2017). Revision of Diagnostic Morphological Traits in Three Caribbean Species of Heterotermes (Dictyoptera: Rhinotermitidae). Presentation at the annual meeting of the North Central Branch of the Entomological Society of America, June 7, Indianapolis, IN. • Griebenow, Z. H., Jones, S. C., & Eaton, T. D. (2016). Revision of Diagnostic Morphological Traits in Four Caribbean Species of Heterotermes (Isoptera: Rhinotermitidae). Presentation at the annual meeting of the Ohio Valley Entomological Society, November 4, Purdue University, West Lafayette, IN. • Griebenow, Z. H., Jones, S. C., & Eaton, T. D. (2016). Revision of Diagnostic Morphological Traits in Four Caribbean Species of Heterotermes (Isoptera: Rhinotermitidae). Poster presented at the 21st annual Denman Undergraduate Research Forum, March 30, The Ohio State University, Columbus, OH. • Griebenow, Z. H., Jones, S. C., & Eaton, T. D. (2016). Revision of Diagnostic Morphological Traits in Four Caribbean Species of Heterotermes (Isoptera: Rhinotermitidae). Poster presented at the CFAES Undergraduate Research Forum, February 18, The Ohio State University, Columbus, OH. EXTENSION Skype a Scientist, April 10, 2021 • Made general presentation on entomology via Zoom for Girl Scout Brownie Troop DCCNS Cooperative Preschool, Davis, CA; February 11, 2020 • Showed pre-K through kindergarten students specimens from personal insect collection under microscope Bohart Open House, Bohart Museum of Entomology; UC-Davis; January 18, 2020 • 1:00-4:00 p.m. o Presented leptanilline specimens and explained research to public audience Picnic Day, University of California-Davis, volunteer at Entomology & Nematology Department’s Picnic Day open house; April 13, 2019 • 9:00 a.m.-5:00 p.m. o Handed out samples of honey o Refereed blattodean races o Answered general entomological queries o Aided sales of T-shirts to gain revenue for the Entomology Graduate Student Association (EGSA) Biodiversity Museum Day, Bohart Museum of Entomology; UC-Davis; February 16, 2019; February 15, 2020; February 13, 2021 (virtual) 4 Zachary Griebenow, C.V. • 7:30 a.m.-1:00 p.m. o Ushered groups of visitors into the Bohart Insect Collection o Displayed specimens to visitors with Hitachi desktop SEM o Explained research project in lay language Roseville Bug & Bird Bonanza, volunteer for Bohart Museum of Entomology exhibit; February 2, 2019 • 10:00 a.m.-2:00 p.m. o Handled and displayed live arthropod specimens o Explained contents of given drawers to visitors Explorit Science Center, volunteer for Bohart Museum of Entomology exhibit; October 15, 2018 • 4:00 p.m.-7:00 p.m. o Handled and displayed live arthropod specimens o Explained contents of given drawers to visitors Picnic Day, University of California-Davis, volunteer at Entomology & Nematology Department’s Picnic Day open house; April 21, 2018 • 9:00 a.m.-5:00 p.m. o Handed out samples of honey o Refereed blattodean races o Answered general entomological queries o Performed in entomological graduate student band (“Alienoptera”) “Engaging with Insects” 4-H Camp, volunteer, June 26-June 30, 2017; June 26-June 30, 2016 • Mentored group of four middle-school-aged children along with another adult, teaching curatorial practices and aiding creation of student insect collections “Life, et al.”, Entomology Blogger, December 2012-December 2018 • Personal blog written almost exclusively about entomological topics o http://gentlecentipede.blogspot.com/ “The Daily Entomologist” Blogger, University of Kentucky, July 2012-May 2016 • Writer by invitation of Dr. Blake Newton for official university entomology blog o http://ukyentomology.blogspot.com/ Museum of Biological Diversity: Triplehorn Insect Collection, Museum Open Houses, February 8, 2014; April 23, 2016; and April 23, 2017 5 Zachary Griebenow, C.V. • Answered visitors’ queries as regarded the contents of given display drawers, 10:00 a.m.-4:00 p.m.
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  • The Ant Larvae of the Subfamily Leptanillinae (Hymenoptera, Formicidae)

    The Ant Larvae of the Subfamily Leptanillinae (Hymenoptera, Formicidae)

    THE ANT LARVAE OF THE SUBFAMILY LEPTANILLINAE (HYMENOPTERA, FORMICIDAE) BY GEORGE C. WHEELER AND JEANETTE WHEELER University o,f North Dakota, Grand Forks The Leptanillinae are a small subfamily comprising x4 species in 3 genera: Leptanilla Emery, species; Leptomesites Kutter, species; Phaulomyrma G. C. & E. W. Wheeler, 2 species. This subfamily has been recorded only from the warmer parts o the Old World: Corsica, Sardinia, North Africa, India, Malaya, Java, Queensland, Western Australia and Japan. Seemingly it is rare, but the paucity of the records may be due to minute size. and hypogeic habits. As W. M. "Wheeler optimistically remarked (932, P. 54) "We should expect careful collecting with the Berlese funnel to bring additional forms to light in South Africa, Madagascar, Asia Minor and India, or even, perhaps, in the warmer.parts of the New World." Brown (954, p- 28) noted: "The habits of the species are such as to render their discovery highly fortuitous under present collecting methods." In the 3o years since Wheeler wrote, only two species have been discovered (one in India and one in Japan), which would rather support another statement by Wheeler in the same article (p. 57-58): "The Leptanillinae... must be very ancient, like many other components of the microgenton L. swani is particularly interesting in this connection, because the extreme south- western corner of Australia, in which it was taken, is known to possess the oldest and least disturbed fauna ) any portion of the. continent." Antiquity is compatible with a disjunct distribution and with rarity. The genus Leptanilla was established by Emery in 87o and then or half a century was kicked about over the taxonomic table.