Redalyc.Glomospirella Cantabrica N. Sp., and Other Benthic Foraminifera from Lower Cretaceous Urgonian-Type Carbonates of Cantab

Geologica Acta: an international earth science journal ISSN: 1695-6133 [email protected] Universitat de Barcelona España

SCHLAGINTWEIT, F.; ROSALES, I.; NAJARRO, M. Glomospirella cantabrica n. sp., and other benthic foraminifera from Lower Cretaceous Urgonian-type carbonates of Cantabria, Spain: Biostratigraphic implications Geologica Acta: an international earth science journal, vol. 14, núm. 2, 2016, pp. 113-138 Universitat de Barcelona Barcelona, España

Available in: http://www.redalyc.org/articulo.oa?id=50549465003

How to cite Complete issue Scientific Information System More information about this article Network of Scientific Journals from Latin America, the Caribbean, Spain and Portugal Journal's homepage in redalyc.org Non-profit academic project, developed under the open access initiative Geologica Acta, Vol.14, Nº 2, June 2016, 113-138, I-III DOI: 10.1344/GeologicaActa2016.14.2.3

Glomospirella cantabrica n. sp., and other benthic foraminifera from Lower Cretaceous Urgonian-type carbonates of Cantabria, Spain: Biostratigraphic implications

F. SCHLAGINTWEIT1 I. ROSALES2 M. NAJARRO2

1Lerchenauerstr., 167, 80935 Munich, Germany. E-mail: [email protected]

2Instituto Geológico y Minero de España (IGME) La Calera 1, 28760 Tres Cantos, Madrid, Spain

ABSTRACT ' & & . ' 2 ( ' 5 ( %&, 4 . ? ? &: ' @ N % & &( ' && ' .&: &: ? > > > , & > . &

KEYWORDS Cantabrian Chain. Aptian. Reocín Formation. Systematics. Orbitolinidae. Dasycladales. Stratigraphy.

INTRODUCTION : ' N' . & & \ ' 2 ( ' 5 ( & !"#$ %& : ' ( ' ) #*+"% , N' ' & . \ - . & / / #*0#1 2 4 54 #*6#% : 2 #*$!1/ . #*0#1 5 #*071 8 > , 5 #*6+%& . &

113 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

12 6 Cuchía 5 N San Vicente de la Barquera 11 Comillas Santillana 3 del mar

8 7 4 Torrelavega 1 9 2 10 El Soplao Ridge NCB Cabuérniga Iberian Peninsula 0 5 10 Km

Carboniferous Jurassic Albian-Cenomanian Reverse fault Permian-Early Triassic Earliest Cretaceous Turonian-Neogene Late Triassic Fault Aptian Quaternary (Keuper facies) Sections with other studied Aptian algae, orbitolinids Studied sections with Glomospirella cantabrica and benthic foraminifera

FIGURE 1. Geological map of the North-Cantabrian Basin with location of the studied samples. 1: Río Nansa section (PN samples), 2: El Soplao section (SOP samples), 3: Novales section (No and NO samples), 4: Cantera de Las Lastrías section (CL samples), 5: Cantera de Cuchía section (CCU samples), 6: Cuchía coast section (CU and PH samples), 7: Santa Eulalia section (SE samples), 8: Ruilobuca section (Ru samples), 9: Rábago section (LA samples), 10: Bustriguado section (Bu samples), 11: San Vicente de la Barquera section (SV samples), 12: Suances section (SU samples).

GEOLOGICAL SETTING #*6*% @ 8 #*+!% 5 #*+$% E #*+$% D !""" !""$% : ' N' ? % (& #% ' & 2 > & > . .. : ? . . ' . A B 5 !"##1 !"#$12 #*6#18 8 #**"%&/ B !"#$%& ? : ' N' "#6 "0" B'D !"#$%&: E #*+$1 . !"##1 !"#$%&( ? . ( &!' F%H%2 ( "NF0 E . N' ( &!1F%& ? ' % - : ' N' 1 % ( "N!$ ? 8 #*!"% \ ' % ? ' & - 2 #*$*% 2 4 54 #*6!% 1%5 ( "N+"

Geologica Acta, 14(2), 113-138, I-III (2016) 114 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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

Geologica Acta, 14(2), 113-138, I-III (2016) 115 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

A Río Nansa-Rábago El Soplao Novales-Ruilobuca-Las Lastrías Cuchía-Suances B Cantera de Las Lastrías section 100.5 Ma SW BIELBA Fm. (Somocuevas Mb.) NE N S Unconformity LAS PEÑOSAS Fm.

Upper BARCENACIONES Fm.

Albian Mid. REOCÍN Fm. Low. LAS PEÑOSAS Fm. Hiatus 113 Ma CL-8 REOCÍN Fm. REOCÍN Fm. CL-7 Rodezas Fm. Aptian San Esteban Fm. San Esteban Fm.

Lower Cretaceous Lower Patrocinio Fm. Patrocinio Fm. Patrocinio Fm. Patrocinio Fm. CL-3 Umbrera Fm. Umbrera Fm. Umbrera Fm. Bedou. Garga. Clansa. 126.3 Ma Rábago Fm. Rábago Fm. Rábago Fm. Barre. Wealden (Pas Group) CL-2 Haut. Vala. Hiatus Hiatus Berri. 145 Ma Jurassic 7 m Triassic

C Unconformity REOCÍN Fm. BARCENACIONES Fm.

CCU-17A CCU-8 5m CCU-7 CCU-6

N Cantera de Cuchía section S

FIGURE 2. A) Lithostratigraphy of the Lower Cretaceous in the North-Cantabrian Basin (modified from Najarro et al. 2011). B) Field picture of the upper part of the Reocín Formation and Las Peñosas Formation in the Cantera de Las Lastrías section, with location of some studied samples. C) Field picture of the Reocín and Barcenaciones formations in the Cantera de Cuchía section, with location of some of the studied samples.

. . (& F%H 2 ( & #0% ' 8 5 ( &F"% 2 ( !"##%&, ' &6" \ !" 4% 5 \ ' % ( &0"% ( & ( ' / ('/% $$% ( & : 4% ' ' 2 5 % ('/ B 4%& 24 ( &!'1F%&

%B &: - &7&+? DESCRIPTION OF THE STRATIGRAPHIC SECTIONS 2 . & : - B . : . B . & & - 2 . &: &!!" ? (&F : , 2 ( & +% (&#%H D % ( & %2 &, - &"&6? 0% 5 ( &!6% 2 . '#+# & ##$ ?& : \ #$ 8 5 2.& , . ? > Q . 2 4 54 #*6!% %2 !"#F% !"##% 2 !"#F%& , 2 ( : ' % . B %& : ( (&F%&:

Geologica Acta, 14(2), 113-138, I-III (2016) 116 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria Cch Cch .

......

...... 40 B 0 m 80 U Bi SE Re W Pa 160 120 section 6: Cuchía coast 6: Cuchía Buntsandstein (Triassic) Buntsandstein Pas Group (Wealden) Group Pas Bielba Fm. (p.p.) Bielba Fm. Fm. Las Peñosas Barcenaciones Fm. Barcenaciones Reocín Fm. Rodezas Fm. Fm. San Esteban Fm. Patrocinio Fm. Umbrera Rábago Fm. B U Ti Bi Cch W SE Pe Re Pa Ra Ro

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......

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. . .

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. . . . B 20 80 60 40 Bi 0 m Re 100 section 5: Cantera Cuchía Cuchía 5: Cantera of sections in Figure 1. Correlation lines are of sections in Figure 1. Correlation lines are Plicatula and rudists Exogyra Exogyra ssil content with biostratigraphic relevance.

. . . .

......

. . .

......

. . . Bioturbated or hardground surface or hardground Bioturbated paleokarstSubaerial exposure: or paleosoils surface/UnconformityErosional Black pebbles and lumps) Bacinellid fabrics (oncoids Bacinellid fabrics (thrombolites/masses) Branching corals domal and platy corals Massive, Chondrodonta Rudists choffati Caprina Oysters, and plant fragments Coal Orbitolinids section Covered

. . .

...... 0 40 section 80 Re Pe 160 Ro 120 Cch 4: Cantera Las Lastrías 4: Cantera . .

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. . .

. .

.. ..

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...... 0 80 40 Pe Pa Ro SE 240 360 Re 280 160 120 200 400 320 3, 8: Composite Ruilobuca-Novales sectionRuilobuca-Novales ...... 0 U 80 40 Ti SE Pa Pe Ra 200 Re W 160 120 2: El Soplao section Stratigraphic cross-section based on the correlation of several measured sections. Numbers sections refer to numbers ...... Cross-bedded, glauconite-rich grainstones-packstone, Albian Albian limestones, and rudist/coral Wackestone Upper Aptian Bacineliid boundstone, Upper Aptian rudstone, to wackestone Bacineliid oncoidal Dolostone Upper Aptian and floatstone, Wackestone Upper Aptian and grainstone, Packstone with siltstones interbedded Limestones Siltstones marls and limestones Interbedded Marls limestones and interbedded Claystone with lenticular bedding Claystone stratification, and ripples flaser bedding with cross Sandstone Aptian Lower Rudist limestones, Aptian Lower and packstone, grainstones Cross-bedded Aptian Lower limestones, and coral gastropod Orbitolinid, claystones and red Sandstones and claystones sandstones Red conglomerates, ...... 0 . . . 40 B 80 . . . Ti Bi Pe Re 120 Ra Pa 200 160 1: Río Nansa section FIGURE 3. established on the basis of regional tracing and mapping lithostratigraphic units main unconformities, fo

Geologica Acta, 14(2), 113-138, I-III (2016) 117 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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

Geologica Acta, 14(2), 113-138, I-III (2016) 118 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

C B D A

G E p.w.

H J I

L K

P O N

R S Q

Y V X W

T U 0.3 mm Z

FIGURE 4. A–P) Glomospirella cantabrica n. sp., Upper Aptian-lowermost Albian of Cantabria. Oblique equatorial sections (A, F–G, M, O–R), axial sections (B–E, H–J, N), equatorial section (K–L). Holotype in K. Note the single planispiral whorl (p.w.) switched within the early Glomospira-type part in J, and the first planispiral whorls differing in their coiling planes in D (arrow). A, E: MGM-10853C-1, -10853C-2 (CCU 6); B: MGM-10863C-1 (No 23); C: MGM-10848C-1 (CL 4); D: MGM-10855C-1 (CCU 8); F–H: MGM-10847C-1, -10847C-2, -10847C-3 (CL 3); I–K: MGM-10862C-1, -10862C-2, -10862C-3 (No 14); L: MGM-10880C-1 (SOP 20); M: MGM-10865C-1 (No 35); N: MGM-10887C-1 (SOP 78); O: MGM-10881C-1 (SOP 40); P: MGM-10866C-1 (PN 13); Q–R: MGM-10854C-1, -10854C-2 (CCU 7). S) Glomospirella sp., (from Setoyama et al., 2011, pl. 2, fig. 18, late Cretaceous of Barents Sea). T) Glomospirella gaultiana (Berthelin, 1880) (from Moullade et al., 2008: pl. 1, fig. 3, late Bedoulian of France). U–W) Glomospira urgoniana Arnaud-Vanneau, 1980, upper Aptian-lowermost Albian of Cantabria.U: MGM-10861C-1 (No 6); V–W: MGM- 10879C-1, -10879C-2 (SOP 19). X–Z) Glomospira? sp. X–Y: MGM-10859C-1, -10859C-2 (La 23); Z: MGM-10864C-1 (No 25).

Geologica Acta, 14(2), 113-138, I-III (2016) 119 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

SYSTEMATIC PALAEONTOLOGY \. & , : ? \ 5 ? (&7S %&, !"#F% ? \ S ?!"#7%& & H( S #+!0 H: !"#F : \ . H' #*+" H' #+0! & , \ @B #*7$ - . : H \ &, #*!6 \ !MF & &(&7'N2% &: Q#*0# &N H&#"6 \ & 7"μ%&: ? Q#*+! &NH \ & ' &#$$ \ & & #*+6 &NH5&! \&0& & ? & : & & 2 - &: . & ( - ? > 8@8#"+76##7%(&7S% A D !"""% . (& F%& . 4 H8@8#"+$F# - &: (&7'% #"+0F#(&7% #"+7+#(&7% S ? #"+$$#(&7/% #"+$F!(&7B% #"+76# @ !""$% (&7(% #"+76!(&7@% #"+76F(&7E% - #"+0!# (& 7,% #"+0!! (& 7A% #"++"# & , (&7% #"+0$#(&78% #"++6#(&7% #"++##(&7O% #"+00#(&75% #"+$7# : #*+6 &$#%&: (&7T% #"+$7!(&72%& . . & &8 4 . &B , & : ? A #**"1 & #$0% \ P - & : Q& E. . ? ? . \ & . S ? !"#7% ? \ & D ( / E 5 ' & !"#$%&

&: "&0$% ? . . /#*0#% #*+!% & : ' ! % 5 & ?& . & %& . & : (&7A%&: %

Geologica Acta, 14(2), 113-138, I-III (2016) 120 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

A B C

D E F

FIGURE 5. Dasycladalean algae from the Aptian of Cantabria. A) Neomeris cf. cretacea STEINMANN. MGM-10874C-1 (Bu 1), lower Aptian Rábago Formation. B) Montiella elitzae (BAKALOVA). MGM-10869C-1 (Ru 3), lower Aptian San Esteban Formation. C) Cylindroporella? pedunculata (JAFFREZO, POISSON AND AKBULUT). MGM-10886C-1 (SOP 77), upper Aptian Reocín Formation. D) Salpingoporella hasi CONRAD, =; AND REY. MGM-10885C-1 (SOP 71), upper Aptian Reocín Formation.E) Genotella pfenderae (KONISHI AND EPIS). MGM-10884C-1 (SOP 69), upper Aptian Reocín Formation. F) Salpingoporella melitae =; MGM-10863C-2 (Nº23), upper Aptian Reocín Formation. Scale bars 0’3mm, except F=1mm.

P Q - ] & S ? #**!1S ? !"""1@ ? ? !""71 8 !""+%& ' 2 ( %& O ? ' 5 B @ !""7%& ' 5 ( & : . > 4U"&F - &' ' "&6F 8 S ? !"""% 2 ( " # & % Q (&7:%& % (& $% $ 8. . 75 (& $/% %(&$B%&: & $ 75 . (&$(% @ - E . ' & !""0 : #%&, > !"##H&! \&#+% S. - ( (& 7% @ . & ? . @4 ? #+*+1 !"##%& % &

Geologica Acta, 14(2), 113-138, I-III (2016) 121 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

TABLE 1. Distribution of taxa from the Aptian–lower Albian formations of Cantabria

Formation Rábago San Rodezas Reocín Las Species Esteban Peñosas Choffatella decipiens X X Palorbitolina lenticularis X X Orbitolinopsis? simplex X Praeorbitolina cormyi X Nautiloculina cretacea X X X Glomospira urgoniana X X X X Charentia cuvillieri X X X X Akaya minuta X X X? Mesorbitolina parva X Mesorbitolina texana X X X Mesorbitolina birmanica X? Dictyoconus? pachymarginalis X Pseudochoffatella cuvillieri X Dobrogelina? carthusiana X Coskinolinella daguini X Orbitolinopsis reticulata X Simplorbitolina aquitanica X Simplorbitolina manasi X X Cuneolina cf. parva X X Glomospirella cantabrica X X Involutina hungarica X Simplorbitolina conulus X

2 ( % (& $'% & ' % B ( . 2 %(&$%& 24 B . 5 ( & 0N+%& ( & : . - : #&' S ? !""+% - . - .% 8 . & - 2 . . ? H #++$ . #**!1 H #*+$ ? !""*%& @B #*F+ ' ' % #*60(&0'N% 2 ( &/ 2 ( % 5 #*60% E . O 5 & & : % . % % Biostratigraphic remarks on some other benthic . ' ) 5 #*6+%& foraminifera % & & &06% P - Q ( & : 2 ( . 1 : #*+6%& \ ' E. \ &: -

Geologica Acta, 14(2), 113-138, I-III (2016) 122 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

A B C D

H G E F

I J L

FIGURE 6. Benthic foraminifera from the upper Aptian Reocín Formation. A–B), Nautiloculina cretacea PEYBERNÉS, slightly oblique equatorial section (A) and axial section (B). C–D), Cuneolina cf. parva HENSON, axial section showing complex embryo (C), longitudinal section parallel to plane of biseriality (D). E–F) Charentia cuvillieri NEUMANN, equatorial section (E), axial section (F). G) Akcaya minuta (HOFKER). H, K, L) Dobrogelina? carthusiana ARNAUD-VANNEAU, transverse section of the dorsal side (H), axial section (K), subaxial sections (L). Note Glomospira urgoniana ARNAUD- VANNEAU in the lower left side of K. I) Pseudochoffatella cuvillieri DELOFFRE, tangential section. J) Akcaya capitata (ARNAUD-VANNEAU) (lower part of the picture) and Akcaya minuta (HOFKER) (upper part). M) Coskinolinella daguini DELMAS AND DELOFFRE. A, K: MGM-10856C-1, -10856C-2 (CCU 17A); B: MGM-10850C-1 (CL 8); C: MGM-10860C-1 (PH 14B); D: MGM-10862C-4 (No 14); E, H, M: MGM-10887C-2, -10887C-3, MGM- 10887C-4 (SOP 78); F: MGM-10854C-3 (CCU 7); G: MGM-10849C-1 (CL 6); I: MGM-10883C-1 (SOP 68); J: MGM-10866C-2 (PN 13); L: MGM- 10882C-2 (SOP 51). Scale bars 0.3mm, except I= 1mm and M= 0.5mm.

Geologica Acta, 14(2), 113-138, I-III (2016) 123 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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

Geologica Acta, 14(2), 113-138, I-III (2016) 124 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

D E

F G H

I J K

FIGURE 7. Orbitolinids from the lower Aptian Rábago (B) and San Esteban formations (A, C, F–G, I), and the upper Aptian Reocín Formation (D–E, H, J–K). A–B) Palorbitolina lenticularis (BLUMENBACH). MGM-10858C-1 (CU-10), and MGM-10867C-1 (PN-4). C) Praeorbitolina cormyi SCHROEDER. MGM-10871C-1 (RU-12). D–E, H) Dictyoconus? pachymarginalis SCHROEDER. D: MGM-10852C-1 (CCU-1), E: MGM-10851C-1 (CL 14), H: MGM- 10868C-1 (PN-14), F–G, I) Orbitolinopsis? simplex (HENSON). F: MGM-10875C-1 (SU-8), G: MGM-10872C-1 (Ru-29), I: MGM-10870C-1 (Ru-6). J–K) Orbitolinopsis reticulata MOULLADE AND PEYBERNÈS, oblique transverse sections. J: MGM-10866C-3 (PN-13), K: MGM-10878C-1 (SOP-14). Scale bars 0.5mm.

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D( & E?#*0$% & : 4 "& ' ' #7$M"&#$M"M"&!#M"&!M"�% &' . . \ #*+$% \ . & #*+$ &$F% #&7 #&0 -& ' @ 6 #*+# %& !&"% #&$ E?#*0$%&: \ B 4% 4 F&! . @ !"#F% & , # ##% .. ) % . % .& : & ' ' #*+$%& @B #*07 ' ) 5 . ! #*07 (&6% #**$% 5 \ A & E. \ 2 ! & : . ' % B : ( ] ) ' N' 5 4 \ ? ? & : 8 #**!%&: ) #*+$% @B #*0! - & #+7*%(&+E A% - &' . : \ & % @ !"#"%&, \ . 4 %&E. E?#*0$% & . O 24 ( 2 ( & & : #*+6 &0*6% ) & #*0"%(&+SN% \ . S ? !"#7% 2 & . &, 24 ( 2 #*+$% . ( &: O \S ?!""7 !"#7%& ? \ \ & HO #+*" & & & & \ @B #*0F ! #+"$% @ & (&# (&6'N% !"#F%&

! % & #*F6%(&+, 8N% . 2 ( 5 7 2 ( . ? - 1 O5# B % D !"#7%& 2 4 54 B ( #" #*6#% & \ %& : O5 # ! +& , ' .& & - . ? # . @ B 4 D & , > . 2 ( &: !""7% \ . % 4 . & &, 2 ( & &: ! & . . ( 2 #*6FH % & &'%&

Geologica Acta, 14(2), 113-138, I-III (2016) 126 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

A B D

E F

G H I

J M

K L N

FIGURE 8. Orbitolinids from the upper Aptian Reocín Formation (A–E, H–N) and the lower Albian Las Peñosas Formation (F–G). A) Simplorbitolina aquitanica (SCHROEDER), oblique subaxial section (specimen on the left) and oblique section (specimen on the right). MGM-10878C-2 (SOP- 14).B–E) Simplorbitolina manasi CIRY AND RAT. Subaxial section (B), axial sections showing protoconch and initial trochospire (with up to 12, ?13 chambers; compare with Schroeder, 1985, pl. 20, fig. 15) (C–D), transverse section (E). B: MGM-10888C-1 (SOP-81B); C: MGM-10882C-1 (SOP-51-1); D: MGM-10887C-6 (SOP-51), E: MGM-10887C-5 (SOP-78). F–G) Simplorbitolina conulus SCHROEDER. Oblique transverse section (F), oblique subaxial section (G). MGM-10877C-1, -10877C-2 (SV-31). H, J) Mesorbitolina texana (ROEMER), axial sections. H: MGM-10876C-1 (SE-2-7); J: MGM-10873C-1 (Ru-69). K–L) Mesorbitolina parva (DOUGLASS). Axial section (K), oblique section (L). MGM-10851C-2, -10851C-3 (CL-14). I, M–N) Mesorbitolina birmanica (SAHNI). Oblique section passing through the marginal zone (lower part) and the central zone (upper part) (I), axial sections of juvenile specimens (M–N). I: MGM-10857C-1 (CCU-17B); M–N: MGM-10856C-3, -10856C-4 (CCU-17A).

Geologica Acta, 14(2), 113-138, I-III (2016) 127 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

@B #*"" @ %& , ( Q #*0$ P ! (&6/ B E% Q - 2 ( & #*0$ % #*6*% @ 2 @ ( &, 5 4 '4 8 , & , 5 $ # 4 8 #**!% @ 8 4 . . #**!%& , - @ 4 8 !"#71 & #6"% %& 2 , . $ Q # % 7 ! 4 ' - ( > !"#" \&#"H % ' 4 % & & & & F%& ' ' !"#F%&2 2 ( Q . & . . & - (48 !"#$%& 2 ( . 2 5#F , 2 ( . #7% # . -2 '#6 2 #7 5#F1B O5 #7%& : ? & &:. \ ? - & \ - & @B #*$! @B #*7+ $ # Q #*7+%(&6( @ ,% 5 #*07%(&+'%

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Geologica Acta, 14(2), 113-138, I-III (2016) 128 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

2 ( \ ' - % ! &\ 2 ( @ . 5 ( & 8 !"#$%& ( ' ( 2 ( ' #*+$%&: 2 ( ( 48 !"#$%& ( + #*+$H & !" \& #$% : % 70 2 ( . & , . 24 ( #!Q#F% . 4 (&+/%& . 4 8 #*!"1 8 !"##% $ #*0$(&+(N@% @ 2 !"#7%(&*%&

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Geologica Acta, 14(2), 113-138, I-III (2016) 129 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

Sea level Benthic Foraminifera events (Haq, 2014) Las Peñosas Formation

* (0-200m) Albian ? KAp 7 ?

Simplorbitolina conulus

carthusiana

Clansayesian Reocín

? Formation

Mesorbitolina birmanica (9-300m)

pachymarginalis

Dobrogelina

Orbitolinopsis reticulata

Simplorbitolina aquitanica

Mesorbitolina texana

Glomospirella cantabrica

Dictyoconus

Mesorbitolina parva

fatella cuvillieri

Simplorbitolina manasi

Aptian Gargasian

Pseudochof

Coskinolinella daguini

Rodezas (2) Formation (0-105m)

KAp 2 Orbitolinopsis simplex San Esteban ? Palorbitolina lenticularis Formation Praeorbitolina cormyi (0-55m) Choffatella decipiens* Patrocinio Fm. deep-water facies (30-80m) (no larger benthic foraminifera) (1)

Bedoulian Umbrera Fm. Palorbitolina lenticularis (0-50m) Rábago Fm. Palorbitolina lenticularis (0-18m) Choffatella decipiens*

FIGURE 9. Biostratigraphy of the Cantabrian Urgonian (Aptian) and distribution of selected benthic foraminifera (orbitolinids and others). The boundary Bedoulian-Gargasian is tentatively placed based on the biostratigraphy of orbitolinids and other benthic foraminifera. The Gargasian- Clansayesian and Aptian-Albian boundaries are intentionally left open. Ammonite data: 1) Deshayesites forbesi and D. deshayesi zones (Collignon et al., 1979; Najarro et al., 2011; García-Mondejar et al., 2015). 2) Epicheloniceras martini Zone, Epicheloniceras gracile subzone (Collignon et al., 1979; Moreno-Bedmar et al., 2011).

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Geologica Acta, 14(2), 113-138, I-III (2016) 130 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

Main depositional sequences North-Cantabrian Iberian Basin Basque- Global SB Basin (this study) Galve Benicàs- Cantabrian (Haq, 2014) area* sim area** Basin ***

Bielba Fm. VIII Lower Cenomanian (Cóbreces Mb. ) 100.5 ma TST RST SB 7 KAl8 Bielba Fm. VII (Somocuevas Mb.) VII 6 TST SB KAl7 KAl6 Late RST VI KAl5 Barcenaciones Fm.

~107.5 ma VI KAl4

Albian 5

Middle V KAl3 ~110 ma 4 Las Peñosas Fm. KAl2 TST IV Lower RST SB 3 KAl1 V TST V III 113 ma SB 2 RST KAp7 Clansayesian IV IV KAp6 IV KAp5 ~117 ma Reocín Fm. TST SB KAp4 RST II Gargasian Aptian III III III Rodezas Fm. MFZ KAp3 TST ~123 ma SB KAp2 RST 1 San Esteban Fm. Bedoulian II II II Patrocinio Fm. MFZ I Umbrera Fm. TST SB KAp1 126.3 ma Rábago Fm. I I I

FIGURE 10. Chrono-, lithostratigraphic and depositional sequences chart showing comparison of the depositional sequences of the North-Cantabrian Basin (this study) with the global cycle boundaries (after Haq, 2014) and the main depositional sequences of the Basque-Cantabrian Basin (BCB) after García-Mondéjar et al., 2004 (***), and the Iberian Chain (Benicàssim and Galve areas) after Martín-Martín et al., 2013 (**) and after Bover-Arnal et al., 2009 (*). I to VIII: depositional sequences. SB: Sequence Boundaries. The dashed lines of the SB indicate uncertainty in the chronostratigraphic dating. TST: Transgressive System Tract, RST: Regressive System Tract, MFZ: Maximum Flooding Zone. Absolute ages are taken from Haq (2014).

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Geologica Acta, 14(2), 113-138, I-III (2016) 131 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 132 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 133 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 134 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 135 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 136 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

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Geologica Acta, 14(2), 113-138, I-III (2016) 137 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

2& #*0F & ! 2& 8&'& A& #*0+& - @ O (O2'8&%& A @ ( 5 ' ##6 F70F$*& . & S O H 2& #*0F& @ O * $ & & & &1 ! % ? @4 %& A ! %&' . @ 5 8 F F!"F!0& !"!% #**!!!& 2& #*07 & S 2& ) (&&5& '& /& O & 2 ) & , '& @ & !"#"& 2. > ( #" 7##7#F& 4 N 2& #*07&O ' ' 5 . : %& A & @' @ 5 8 #*07 70!76!& 5 7 7**0& 2& #*0$ & & & B& S ? 8&'& : 4? A& !"##& ' B 4@ , % . O ,,,%& B @ E. $+!% *60*6*& &,HS ? 8&'& ( & 2& #*0$& % & & %& 5 B , D? 5 O &B ' ( &@4 ?( @ E. $+#% $6*$+*& 5 #0 !$#F"*& 2& #*6!& R O ( &% '&E& D& #*0!& S& A @ ! &5 7 $+#$*#& 5 8 ! #"+##*& A& 2& 8 '& !""!& 2& #*+$& @ $ p 2 #*$F& 2 ,H 2& 8& &%& @ O %&5 & ( 8 8 & B &: 8 @ ## @ 6 70$"& 06*$& 2& 5 '&(& #*07& # D 8& !"""&B. 8 ( ?'> D H ' % ( ? % .O ,%&B %& @ #FF @ E. $6!% $$F$$+& #*$!!0& 2& 8& &% #*+$& D 8& !""$& ( 8 8 & ' %& A @ 8 6 ##0"& , @ F#!% !$F!6$&

Manuscript received August 2015; revision accepted December 2015; published Online May 2016.

Geologica Acta, 14(2), 113-138, I-III (2016) 138 F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

ELECTRONIC APPENDIX I

TABLE I. Geographic coordinates and stratigraphy of the studied sections

Name of the Geographic Previous Stratigraphic Sample Remarks stratigraphic section studies/ succession ID coordinates descriptions

1: Río Nansa section Top: 43º 18' 35.89'' N; Ramírez del Rábago Fm.: ca. PN The Rábago Fm. 4º 28' 11.76'' W Pozo, 1972. 16m overlays Patrocinio Fm.: unconformably Triassic (Bunt) Najarro et al., ca. 30m Base: 43º 18' 09.94'' N, 2011a siliciclastics. Reocín Fm.: ca. Absence of the 4º 27' 46.01'' W 70m Umbrera, San Rosales, 2013. Las Peñosas Esteban and Fm.: ca. 60m Rodezas fms. Barcenaciones Fm.: ca. 55m

2: El Soplao section Top: 43º 18' 05.54'' N; Najarro, 2015. Rábago Fm.: ca. SOP Absence of the 4º 25' 10.57'' W 8m Rodezas Fm. Rosales et al., Umbrera Fm.: 2013 ca. 6m Base: 43º 17' 44.31'' N; The first 35m of Patrocinio Fm.: 4º 24' 31.28'' W the Reocín Fm. ca. 27m are pervasively San Esteban dolomitized. Fm.: ca. 15m Reocín Fm.: ca. 100m Las Peñosas Fm.: ca. 60m (top covered)

3: Novales section Top: 43º 22' 33.74'' N; Najarro, 2015. Reocín Fm.: ca. No and The first 160m of 4º 09' 46.94'' W 230m NO the Reocín Fm. Las Peñosas are dolomitized. Fm.: ca. 70m Base: 43º 22' 58.25'' N; (lower part, top 4º 10' 13.14'' W covered)

4: Cantera de Las Top: 43º 20' 16.02'' N; Najarro, 2015. Reocín Fm.: ca. CL Lastrías section 4º 09' 05.23'' W 130m Las Peñosas Fm.: ca. 5m Base: 43º 20' 11.41'' N; (lower part) 4º 09' 15.35'' W

5: Cantera de Cuchía Top: 43º 25' 42.23'' N; Collignon et Reocín Fm.: ca. CCU Absence of the section 4º 01' 03.74'' W al., 1979. 80m (base Las Peñosas Fm. covered).

Najarro, 2015. Barcenaciones Base: 43º 25' 55.94'' N; Fm.: 26m 4º 00' 55.68'' W Fernández- Mendiola et al., 2015.

Geologica Acta, 14(2), 113-138, I-III (2016) I F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

TABLE I. Continued.

Name of the Geographic Previous Stratigraphic Sample Remarks stratigraphic section studies/ succession ID Coordinates descriptions

6: Cuchía coast section Top: 43º 26' 10.52'' N; Mengaud, Umbrera Fm.: CU and Very reduced 4º 01' 47.38'' W 1920. ca. 22m PH Rodezas and Patrocinio Fm.: Reocín fms., and absence of the Collignon et ca. 80m Base: 43º 26' 42.00'' N; Rábago and Las al., 1979. San Esteban Peñosas fms. 4º 01' 19.20'' W Fm.:. ca. 26m Wilmsen, Rodezas Fm.: 2005. ca. 1.5m (top disrupted by fault) Najarro, 2015. Reocín Fm.: ca. 11m García- Barcenaciones Mondéjar et Fm.: ca. 18m al., 2015.

7: Santa Eulalia Top: 43º 20' 52.94'' N; Najarro, 2015. Rodezas Fm.: SE2 The middle part section 4º 11' 47.03'' W ca. 76m of the Reocín Fm. Reocín Fm.: ca. exhibits pervasive Blázque- dolomitization. Fernández, 211m Base: 43º 20' 14.70'' N; 2013. 4º 12' 03.69'' W

8: Ruilobuca section Top: 43º 22' 37.49'' N; Najarro, 2015. San Esteban Ru The top of this 4º 15' 12.75'' W Fm.: ca. 55m section correlates Rodezas Fm.: with the base of ca. 105m the Novales Base: 43º 22' 08.63'' N; section. Reocín Fm.: ca. 4º 15' 47.16'' W 20m (base only).

9: Rábago section Top: 43º 18' 21.22'' N; Najarro et al., Rábago Fm.: ca. LA Absence of the 4º 26' 02.95'' W 2011b. 16m Umbrera, San Patrocinio Fm.: Esteban and Rodezas Najarro, 2015. ca. 33m Base: 43º 18' 03.32'' N; formations. Reocín Fm.: ca. 4º 26' 12.30'' W 78m Las Peñosas Fm.: ca. 70m Barcenaciones Fm.: ca. 40m

10: Bustriguado Top: 43º 18' 19.83'' N; Najarro, 2015. Rábago Fm.: ca. Bu Absence of the section 4º 21' 18.80'' W 36m Rodezas Fm. Umbrera Fm.: ca. 16m Base: 43º 17' 55.34'' N; Patrocinio Fm.: The first 150 m of 4º 21' 01.42'' W ca. 30m the Reocín Fm. San Esteban exhibits pervasive Fm.: ca. 38m dolomitization. Reocín Fm.: ca. 267m Las Peñosas Fm.: ca. 125m Barcenaciones Fm.: ca. 70m

Geologica Acta, 14(2), 113-138, I-III (2016) II F. Schlagintweit et al. Lower Cretaceous foraminifera from Cantabria

TABLE I. Continued.

Name of the Geographic Previous Stratigraphic Sample Remarks stratigraphic section studies/ succession ID Coordinates descriptions

11: San Vicente de la Top: 43º 23' 42.03'' N; Najarro, 2015. Reocín Fm.: ca. SV The Reocín Fm. Barquera section 4º 23' 47.35'' W 152m overlays Las Peñosas unconformably Fm.: ca. 80m Carboniferous Base: 43º 23' 42.05'' N; (top covered) limestones, with 4º 25' 50.42'' W absence of the Rábago, Umbrera, San Esteban and Rodezas formations.

12: Suances section Top: 43º 26' 35.33'' N; Rat, 1959. San Esteban SU Stratigraphy 4º 02' 43.76'' W Fm.: ca. 17m similar to the (base covered) Cuchía coast Najarro, 2015. Rodezas Fm.: section. Base: 43º 26' 39.72'' N; ca. 4–6m Absence of the 4º 02' 45.17'' W Reocín Fm.: 8m Las Peñosas Fm. Barcenaciones Fm.: ca. 12m.

Geologica Acta, 14(2), 113-138, I-III (2016) III