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Explosive Radiation in High Andean Hypericum—Rates of Diversification
ORIGINAL RESEARCH ARTICLE published: 11 September 2013 doi: 10.3389/fgene.2013.00175 Explosive radiation in high Andean Hypericum—rates of diversification among New World lineages Nicolai M. Nürk 1*, Charlotte Scheriau 1 and Santiago Madriñán 2 1 Department of Biodiversity and Plant Systematics, Centre for Organismal Studies Heidelberg, Heidelberg University, Heidelberg, Germany 2 Laboratorio de Botánica y Sistemática, Departamento de Ciencias Biológicas, Universidad de los Andes, Bogotá DC, Colombia Edited by: The páramos, high-elevation Andean grasslands ranging from ca. 2800 m to the snow Federico Luebert, Freie Universität line, harbor one of the fastest evolving biomes worldwide since their appearance in the Berlin, Germany northern Andes 3–5 million years (Ma) ago. Hypericum (St. John’s wort), with over 65% Reviewed by: of its Neotropical species, has a center of diversity in these high Mountain ecosystems. Andrea S. Meseguer, Institute National de la research agricultural, Using nuclear rDNA internal transcribed spacer (ITS) sequences of a broad sample of France New World Hypericum species we investigate phylogenetic patterns, estimate divergence Colin Hughes, University of Zurich, times, and provide the first insights into diversification rates within the genus in the Switzerland Neotropics. Two lineages appear to have independently dispersed into South America *Correspondence: around 3.5 Ma ago, one of which has radiated in the páramos (Brathys). We find strong Nicolai M. Nürk, Department of Biodiversity and Plant Systematics, support for the polyphyly of section Trigynobrathys, several species of which group within Centre for Organismal Studies Brathys, while others are found in temperate lowland South America (Trigynobrathys Heidelberg, Heidelberg University, s.str.). -
CODE of COLORADO REGULATIONS 2 CCR 406-9 Colorado Parks and Wildlife
DEPARTMENT OF NATURAL RESOURCES Colorado Parks and Wildlife CHAPTER W-9 - WILDLIFE PROPERTIES 2 CCR 406-9 [Editor’s Notes follow the text of the rules at the end of this CCR Document.] _________________________________________________________________________ ARTICLE I GENERAL PROVISIONS #900 REGULATIONS APPLICABLE TO ALL WILDLIFE PROPERTIES, EXCEPT STATE TRUST LANDS A. DEFINITIONS 1. “Aircraft” means any machine or device capable of atmospheric flight, including, but not limited to, airplanes, helicopters, gliders, dirigibles, balloons, rockets, hang gliders and parachutes, and any models thereof. 2. “Water contact activities” means swimming, wading (except for the purpose of fishing), waterskiing, sail surfboarding, scuba diving, and other water related activities which put a person in contact with the water (without regard to the clothing or equipment worn). 3. “Youth mentor hunting” means hunting by youths under 18 years of age. Youth hunters under 16 years of age shall at all times be accompanied by a mentor when hunting on youth mentor properties. A mentor must be 18 years of age or older and hold a valid hunter education certificate or be born before January 1, 1949. B. Public Access to State Wildlife Areas 1. Only properties listed in this chapter are open for public access. The Director may open newly acquired properties for public access for a period not to exceed nine (9) months pending adoption of permanent regulations. In addition, the Director may establish and post restrictions based upon consideration of the following criteria: a. The location and size of the area. b. The location, type and condition of roads, vehicle parking areas and the number and type of sanitary facilities available. -
The Buffer Handbook Plant List
THE BUFFER HANDBOOK PLANT LIST Originally Developed by: Cynthia Kuhns, Lake & Watershed Resource Management Associates With funding provided by U.S. Environmental Protection Agency and Maine Department of Environmental Protection,1998. Revised 2001 and 2009. Publication #DEPLW0094-B2009 TABLE OF CONTENTS Page Acknowledgements 1 Introductory Information Selection of Plants for This List 1 Plant List Organization & Information 3 Terms & Abbreviations 4 Plant Hardiness Zone Map 5 General Tree & Shrub Planting Guidelines 5 Tips for Planting Perennials 7 Invasive Plants to Avoid 7 Plant Lists TREES 8 (30 to 100 ft.) SHRUBS 14 Small Trees/Large Shrubs 15 (12 to 30 ft.) Medium Shrubs 19 (6 to 12 ft.) Small Shrubs 24 (Less than 6 ft.) GROUNDLAYERS 29 Perennial Herbs & Flowers 30 Ferns 45 Grasses 45 Vines 45 References 49 ACKNOWLEDGEMENTS Original Publication: This plant list was published with the help of Clean Water Act, Section 319 funds, under a grant awarded to the Androscoggin Valley Soil and Water Conservation District and with help from the Maine Department of Environmental Protection and the U.S. Environmental Protection Agency. Graphics and ‘clip-art’ used in this document came from the University of Wisconsin-Extension and from Microsoft Office 97(Small Business Edition) and ClickArt 97 (Broderbund Software, Inc). This publication was originally developed by Cynthia Kuhns of Lake & Watershed Resource Management Associates. Substantial assistance was received from Phoebe Hardesty of the Androscoggin Valley Soil and Water Conservation District. Valuable review and advice was given by Karen Hahnel and Kathy Hoppe of the Maine Department of Environmental Protection. Elizabeth T. Muir provided free and cheerful editing and botanical advice. -
Assessment Report on Hypericum Perforatum L., Herba
European Medicines Agency Evaluation of Medicines for Human Use London, 12 November 2009 Doc. Ref.: EMA/HMPC/101303/2008 COMMITTEE ON HERBAL MEDICINAL PRODUCTS (HMPC) ASSESSMENT REPORT ON HYPERICUM PERFORATUM L., HERBA 7 Westferry Circus, Canary Wharf, London, E14 4HB, UK Tel. (44-20) 74 18 84 00 Fax (44-20) 75 23 70 51 E-mail: [email protected] http://www.emea.europa.eu © European Medicines Agency, 2009. Reproduction is authorised provided the source is acknowledged TABLE OF CONTENTS I. REGULATORY STATUS OVERVIEW...................................................................................4 II. ASSESSMENT REPORT............................................................................................................5 II.1 INTRODUCTION..........................................................................................................................6 II.1.1 Description of the herbal substance(s), herbal preparation(s) or combinations thereof 6 II.1.1.1 Herbal substance:........................................................................................................ 6 II.1.1.2 Herbal preparation(s): ................................................................................................ 7 II.1.1.3 Combinations of herbal substance(s) and/or herbal preparation(s)........................... 9 Not applicable. ................................................................................................................................9 II.1.1.4 Vitamin(s) ................................................................................................................... -
Byword(USA) Chestnut, 2006 • Height 16Hh
Byword(USA) chestnut, 2006 • height 16hh Nearctic - Nearco NORTHERN DANCER Natalma - Native DANCER RACE RECORD Nureyev 1977 Forli - Aristophanes In France, UK, USA and Hong Kong, 7 wins, Special 1600-2000m (from 16 starts; 15 races in Graded and Thong - Nantallah Listed races, for earnings of £699.385). Peintre Celebre Champion Older Horse France 2010 (USA)1994 RAISE A Native - Native DANCER Alydar Timeform 126 – “effective on firm ground as well as soft; Sweet Tooth - On-And-On very best efforts at around 1¼m” Peintre Bleue At 3 WON Prix Pelleas-(L) (2000m) 1987 Habitat - Sir Gaylord 4th Gr2 Prix Guillaume d’Ornano (200m) Petroleuse 4th Gr3 Prix du Prince d’Orange (2000m) Plencia - Le Haar At 4 WON Gr1 Prince Of Wales S. (2000m) RAISE A Native - Native DANCER WON Gr2 Prix du Muguet (1600m) Mr Prospector WON Prix Jacques Laffitte-(L) (1800m) Woodman Gold Digger - Nashua 2nd Gr1 Prix d’Ispahan (1850m) 1983 Buckpasser - Tom Fool 3rd Gr1 Juddmonte Int. S. (2050m) Playmate 4th Gr2 Prix Foy (2400m) Binche (USA) Intriguing - Swaps At 5 WON Gr2 Prix Dollar (1950m) 1999 Blushing Groom - Red God WON Gr3 Prix du Chemin de Fer du Nord (1600m) Rainbow Quest 2nd Gr2 Prix du Muguet (1600m) Binary I Will Follow - Herbager 1993 Nijinsky - NORTHERN DANCER SIRE LINE Balabina PEINTRE CELEBRE – European Horse Of The Year 1997. Peace - Klairon European Champion 3yo, won 5 of 7 starts incl Gr1 GP de Paris, Gr1 Prix du Jockey Club, Gr1 Prix de l’Arc de SALES ANALYSIS 4th dam Triomphe (TFR 137). Sire of sires & broodmare sire of Gr1 BORN colt avg sold SI filly avg sold SI PEACE (66f Klairon): won Blue Seal S.-(L) (only start at 2; TFR winners worldwide. -
Toxicity Assessment of Hypericum Olympicum Subsp. Olympicum L. On
J Appl Biomed journal homepage: http://jab.zsf.jcu.cz DOI: 10.32725/jab.2020.002 Journal of Applied Biomedicine Original research article Toxicity assessment of Hypericum olympicum subsp. olympicum L. on human lymphocytes and breast cancer cell lines Necmiye Balikci 1, Mehmet Sarimahmut 1, Ferda Ari 1, Nazlihan Aztopal 1, 2, Mustafa Zafer Özel 3, Engin Ulukaya 1, 4, Serap Celikler 1 * 1 Uludag University, Faculty of Science and Arts, Department of Biology, Bursa, Turkey 2 Istinye University, Faculty of Science and Literature, Department of Molecular Biology and Genetics, Istanbul, Turkey 3 University of York, Department of Chemistry, Heslington, York, United Kingdom 4 Istinye University, Faculty of Medicine, Department of Medical Biochemistry, Istanbul, Turkey Abstract There is a limited number of studies about the constituents ofHypericum olympicum subsp. olympicum and its genotoxic and cytotoxic potency. We examined the possible antigenotoxic/genotoxic properties of methanolic extract of H. olympicum subsp. olympicum (HOE) on human lymphocytes by employing sister chromatid exchange, micronucleus and comet assay and analyzed its chemical composition by GCxGC-TOF/MS. The anti-growth activity against MCF-7 and MDA-MB-231 cell lines was assessed by using the ATP viability assay. Cell death mode was investigated with fluorescence staining and ELISA assays. The major components of the flower and trunk were determined as eicosane, heptacosane, 2-propen-1-ol, hexahydrofarnesyl acetone and α-muurolene. HOE caused significant DNA damage at selected doses (250–750 µg/ml) while chromosomal damage was observed at higher concentrations (500 and 750 µg/ml). HOE demonstrated anti-growth activity in a dose-dependent manner between 3.13–100 µg/ml. -
Response of Two Chrysolina Species to Different Hypericum Hosts
Seventeenth Australasian Weeds Conference Response of two Chrysolina species to different Hypericum hosts Ronny Groenteman', Simon V. Fowler' and Jon J. Sullivan2 ' Landcare Research, Gerald Street, PO Box 40, Lincoln, New Zealand 2Bio-Protection Research Centre, PO Box 84, Lincoln University, Lincoln, New Zealand Corresponding author: [email protected] SummaryChrysolina hyperici and C. quadrigemina introduced in 1963 but, for many years was thought (Coleoptera: Chrysomelidae) were introduced to to have failed to establish (reviewed by Hancox et al. New Zealand for biological control of St John's wort 1986). Chrysolina quadrigemina was rediscovered in (SJW), Hypericum perforatum, following successful the late 1980s (Fraser and Emberson 1987). It is now biological control in Australia. In other parts of the abundant in mixed populations with C. hyperici (R. invaded range of SJW worldwide C. quadrigemina is Groenteman personal observations). generally accepted as the more significant contributor St John's wort beetles are not strictly restricted to to SJW successful biocontrol. Their ability to feed and H. perforatum, and are known to be able to develop on develop on indigenous Hypericum species was not other Hypericum species (some examples are reviewed tested. Chrysolina hyperici established well while C. by Harris 1988). New Zealand hosts 10 naturalised quadrigemina was initially thought to have failed to es- (Healy 1972) and four indigenous (Heenan 2008) tablish in New Zealand, although it is now widespread. Hypericum species. Little is known about the suit- Thus, identifying differences between Chrysolina spe- ability and impacts of either Chrysolina species on cies in host preference and performance would have these Hypericum species. -
Food: Just Grow It!
Food: Just Grow It! Developed with funding support from the Healthy Hawai`i Initiative State of Hawai`i Department of Health __________________________________________________ PROJECT LEADERS: University of Hawaii College of Tropical Agriculture and Human Resources Cooperative Extension Hawaii State Department of Education Food: Just Grow It! … a supplementary compendium of teaching-learning activities designed to enhance secondary students’ thinking and reasoning skills … __________________________________________________ University of Hawaii College of Tropical Agriculture and Human Resources Cooperative Extension Hawaii State Department of Education February 2004 FFoooodd:: JJuusstt GGrrooww IItt!! TABLE OF CONTENTS OVERVIEW: 1-6 ACTIVITIES: “Rot for Your Plot” Introduction to Theme Units 7 Creating Soil (Weathering Effects) 9 Hot Spots (Warming and Cooling) 16 Porous or Poor-Us (Soil Characteristics) 25 Taste of Dirt? (pH) 36 Dirt Rich (Soil in the Food Cycle) 45 Under-Cover Critters & Creatures (Composting) 54 Compost Cook-Off (Making Compost) 63 “Why Organic Growing?” Introduction to Theme Units 71 Malama i ka `Aina (Hawaiian Culture) 73 Victory Gardens (WW II Oral History) 83 What Goes Down Stays Around (Water Cycle) 92 OG-What? (Organic Farming Certification) 105 People’s Perceptions (Organic Farming Survey) 114 The Great Debate (Organic vs. High-Intensity) 125 WOG It! (Growing Organically) 133 “Know Your Pests” Introduction to Theme Units 141 Pest-iness (Informal Classification) 143 Least “Wanted” (Local Pest / Disease Problem) -
Division Properties 2 Ccr 406-7
DEPARTMENT OF NATURAL RESOURCES Division of Wildlife CHAPTER 9 - DIVISION PROPERTIES 2 CCR 406-7 [Editor’s Notes follow the text of the rules at the end of this CCR Document.] ARTICLE I - GENERAL PROVISIONS #900 REGULATIONS APPLICABLE TO ALL DIVISION PROPERTIES, EXCEPT STATE TRUST LANDS A. Public Access to State Wildlife Areas 1. Only properties listed in this chapter are open for public access. The Director may open newly acquired properties for public access for a period not to exceed nine (9) months pending adoption of permanent regulations. In addition, the Director may establish and post restrictions based upon consideration of the following criteria: a. The location and size of the area. b. The location, type and condition of roads, vehicle parking areas and the number and type of sanitary facilities available. c. The number of users and vehicles the area will tolerate without significant degradation to wildlife resources, and public or private property. d. Opportunity to assure public safety, health and welfare. 2. If a property is opened for public access pursuant to this provision, the property shall be posted with a list of applicable access restrictions. It shall be unlawful for any person or vehicle to enter any such property except in accordance with its posting and the applicable restrictions. B. Prohibited Activities Except as specifically authorized by contractual agreement, official document, public notice, permit or by posted sign, the following activities are prohibited on all lands, waters, the frozen surface of waters, rights- of-way buildings, and other structures or devices owned, operated, or under the administrative control of the Division of Wildlife. -
Attachment A
PLANT EXPLORATION IN THE REPUBLIC OF GEORGIA TO COLLECT GERMPLASM FOR CROP IMPROVEMENT August 26- September 14, 2007 Participants: Joe-Ann H. McCoy, USDA/ ARS/ NPGS, Medicinal Plant Curator North Central Regional Plant Introduction Station G212 Agronomy Hall, Iowa State University, Ames, IA 50011-1170 Phone: 515-294-2297 Fax: 515-294-1903 Email: [email protected] / [email protected] Barbara Hellier, USDA/ ARS/ NPGS, Horticulture Crops Curator Western Regional Plant Introduction Station 59 Johnson Hall, WSU, PO Box 646402, Pullman, WA 99164-6402 Phone: 509-335-3763 Fax: 509-335-6654 Email: [email protected] Georgian Participants: Ana Gulbani Georgian Plant Genetic Resource Centre, Research Institute of Farming Tserovani, Mtskheta, 3300 Georgia. www.cac-biodiversity.org Phone: 995 99 96 7071 Fax: 995 32 26 5256 Email: [email protected] Marina Mosulishvili, Senior Scientist, Institute of Botany Georgian National Museum 3, Rustaveli Ave., Tbilisi 0105 GEORGIA Phone: 995 32 29 4492 / 995 99 55 5089 Email: [email protected] / [email protected] Sandro Okropiridze Mosulishvili, Driver Sandro [email protected] (From Left – Marina Mosulishvili, Sandro Okropiridze, Joe-Ann McCoy, Barbara Hellier, Ana Gulbani below Mt. Kazbegi) 2 Acknowledgements: ¾ The expedition was funded by the USDA/ARS Plant Exchange Office, Beltsville, Maryland ¾ Representatives from the Georgia National Museum and the Georgian Plant Genetic Resources Center planned the itinerary and made all transportation, lodging and guide arrangements ¾ Special thanks -
Tidal Creek Buffer Information Packet
Dear New Neighbor, Welcome to the neighborhood! We hope that you find the Lowcountry as beautiful and enjoyable as we do. We are the Lowcountry Stormwater Partners, or LSP for short. The LSP is a partnership between the Clemson University Cooperative Extension Service, local government, and other organizations that work to educate communities on ways to keep our waterways healthy, fishable, and swimmable. We’re reaching out to discuss the gem of our region, the salt marsh. The salt marsh provides us with many benefits including a thriving commercial fishery, wildlife habitat, and water filtration. It also protects us from storm effects and flooding. To continue enjoying these benefits, we must protect the salt marsh from polluted stormwater runoff. Stormwater runoff becomes polluted when it flows over materials like excess fertilizer, pet waste, and litter. Stormwater runoff can pick up these pollutants and bring them to the nearest waterway. One of the best ways to protect our salt marshes is to maintain a vegetated buffer (land preceding the salt marsh filled with native plant life). Buffers can benefit the saltmarsh and your home by: • reducing stormwater runoff and shoreline erosion • providing increased privacy while maintaining view corridors and wildlife habitat • saving money as this area requires little to no water, fertilizers, or pesticides Many local governments in our area have ordinances related to activities in the buffer area. To help you understand and protect your buffer, we’ve created and attached a small information packet. This packet contains: • The Life Along the Saltmarsh: Protecting Tidal Creeks with Vegetative Buffers factsheet • Backyard Buffers from the SC Department of Health and Environmental Control • An additional native plant list for expanded design options • An example buffer design with plant cost information • Contact information for local buffer ordinances If you have any questions about buffers or protecting water quality, please do not hesitate to reach out to us. -
Best Practice Management Guide 7 BEST PRACTICE MANAGEMENT GUIDE for ENVIRONMENTAL WEEDS
best practice management guide 7 BEST PRACTICE MANAGEMENT GUIDE FOR ENVIRONMENTAL WEEDS ISSN 1442-7192 St Johns wort, Hypericum perforatum Taxonomy and status Description Botanical name: Hypericum perforatum L. - Habit/lifeform: St Johns wort is a perennial Family Clusiaceae (previously Guttiferae or herb with two growth stages - in autumn and Hypericaceae). winter as a flat low rosette, diameter 10-60 cm, with spindly non-flowering stems and a dense Standard common name: St Johns wort. mat of leaves, and in spring and summer as an Aculus hyperici mite, a biocontrol Relationship to other species in Australia: erect twiggy form which produces one or more agent. There are two indigenous native species of woody flowering or non-flowering stems, 30- Photo: CSIRO. Hypericum which may co-occur with St Johns wort 120 cm high. and with which it could be confused. Both Description: Mature plants have a central indigenous species may be distinguished by the woody crown. In late autumn, winter and absence of black gland dots on the petals and early spring, horizontal, pale green or reddish leaves, the presence of 4 longitudinal ridges on the stems with bright green, elongate leaves grow stem (young stems of St Johns wort are 2-ridged) from the crown to form a rosette. One to many and by the stamens not being fused into bundles. upright flowering stems are produced from this Hypericum gramineum, small St Johns wort, is an crown in spring. Clusters of bright yellow indigenous species usually smaller (10-430 cm flowers (1-2 cm in diameter, with 5 petals and high) than St Johns wort which can be black glands on the margins) develop in distinguished by its petals being less than 8 mm summer (Figure 1).