Formation, Deformation and Chertification of Systematic Clastic Dykes in a Differentially Lithified Carbonate Multilayer
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JAHRBUCH DER GEOLOGISCHEN BUNDESANSTALT Jb
JAHRBUCH DER GEOLOGISCHEN BUNDESANSTALT Jb. Geol. B.-A. ISSN 0016–7800 Band 149 Heft 1 S. 61–109 Wien, Juli 2009 A Revision of the Tonnoidea (Caenogastropoda, Gastropoda) from the Miocene Paratethys and their Palaeobiogeographic Implications BERNARD LANDAU*), MATHIAS HARZHAUSER**) & ALAN G. BEU***) 2 Text-Figures, 10 Plates Paratethys Miozän Gastropoda Caenogastropoda Tonnoidea Österreichische Karte 1 : 50.000 Biogeographie Blatt 96 Taxonomie Contents 1. Zusammenfassung . 161 1. Abstract . 162 1. Introduction . 162 2. Geography and Stratigraphy . 162 3. Material . 163 4. Systematics . 163 1. 4.1. Family Tonnidae SUTER, 1913 (1825) . 163 1. 4.2. Family Cassidae LATREILLE, 1825 . 164 1. 4.3. Family Ranellidae J.E. GRAY, 1854 . 170 1. 4.4. Family Bursidae THIELE, 1925 . 175 1. 4.5. Family Personidae J.E. GRAY, 1854 . 179 5. Distribution of Species in Paratethyan Localities . 180 1. 5.1. Diversity versus Stratigraphy . 180 1. 5.2. The North–South Gradient . 181 1. 5.3. Comparison with the Pliocene Tonnoidean Fauna . 181 6. Conclusions . 182 3. Acknowledgements . 182 3. Plates 1–10 . 184 3. References . 104 Revision der Tonnoidea (Caenogastropoda, Gastropoda) aus dem Miozän der Paratethys und paläobiogeographische Folgerungen Zusammenfassung Die im Miozän der Paratethys vertretenen Gastropoden der Überfamilie Tonnoidea werden beschrieben und diskutiert. Insgesamt können 24 Arten nachgewiesen werden. Tonnoidea weisen generell eine ungewöhnliche weite geographische und stratigraphische Verbreitung auf, wie sie bei anderen Gastropoden unbekannt ist. Dementsprechend sind die paratethyalen Arten meist auch in der mediterranen und der atlantischen Bioprovinz vertreten. Einige Arten treten zuerst im mittleren Miozän der Paratethys auf. Insgesamt dokumentiert die Verteilung der tonnoiden Gastropoden in der Parate- thys einen starken klimatischen Einfluss. -
UC Davis UC Davis Previously Published Works
UC Davis UC Davis Previously Published Works Title Molluscan marginalia: Serration at the lip edge in gastropods Permalink https://escholarship.org/uc/item/2mx5c6w9 Journal Journal of Molluscan Studies, 80(3) ISSN 0260-1230 Author Vermeij, GJ Publication Date 2014 DOI 10.1093/mollus/eyu020 Peer reviewed eScholarship.org Powered by the California Digital Library University of California Journal of The Malacological Society of London Molluscan Studies Journal of Molluscan Studies (2014) 80: 326–336. doi:10.1093/mollus/eyu020 Advance Access publication date: 16 April 2014 Molluscan marginalia: serration at the lip edge in gastropods Geerat J. Vermeij Geology Department, University of California, One Shields Avenue, Davis, CA 95616, USA Correspondence: G.J. Vermeij; e-mail: [email protected] Downloaded from (Received 5 September 2013; accepted 10 February 2014) ABSTRACT The shells of many marine gastropods have ventrally directed serrations (serial projections) at the edge http://mollus.oxfordjournals.org/ of the adult outer lip. These poorly studied projections arise as extensions either of external spiral cords or of interspaces between cords. This paper describes taxonomic, phylogenetic, architectural and func- tional aspects of serrations. Cord-associated serrations occur in cerithiids, strombids, the personid Distorsio anus, ocenebrine muricids and some cancellariids. Interspace-associated serrations are phylo- genetically much more widespread, and occur in at least 16 family-level groups. The nature of serration may be taxonomically informative in some fissurellids, littorinids, strombids and costellariids, among other groups. Serrated outer lips occur only in gastropods in which the apex points more backward than upward, but the presence of serrations is not a necessary byproduct of the formation of spiral sculp- tural elements. -
Gradual Miocene to Pleistocene Uplift of the Central American Isthmus: Evidence from Tropical American Tonnoidean Gastropods Alan G
J. Paleont., 75(3), 2001, pp. 706±720 Copyright q 2001, The Paleontological Society 0022-3360/01/0075-706$03.00 GRADUAL MIOCENE TO PLEISTOCENE UPLIFT OF THE CENTRAL AMERICAN ISTHMUS: EVIDENCE FROM TROPICAL AMERICAN TONNOIDEAN GASTROPODS ALAN G. BEU Institute of Geological and Nuclear Sciences, P O Box 30368, Lower Hutt, New Zealand, ,[email protected]. ABSTRACTÐTonnoidean gastropods have planktotrophic larval lives of up to a year and are widely dispersed in ocean currents; the larvae maintain genetic exchange between adult populations. They therefore are expected to respond rapidly to new geographic barriers by either extinction or speciation. Fossil tonnoideans on the opposite coast of the Americas from their present-day range demonstrate that larval transport still was possible through Central America at the time of deposition of the fossils. Early Miocene occurrences of Cypraecassis tenuis (now eastern Paci®c) in the Caribbean probably indicate that constriction of the Central American seaway had commenced by Middle Miocene time. Pliocene larval transport through the seaway is demonstrated by Bursa rugosa (now eastern Paci®c) in Caribbean Miocene-latest Pliocene/Early Pleistocene rocks; Crossata ventricosa (eastern Paci®c) in late Pliocene rocks of Atlantic Panama; Distorsio clathrata (western Atlantic) in middle Pliocene rocks of Ecuador; Cymatium wiegmanni (eastern Paci®c) in middle Pliocene rocks of Atlantic Costa Rica; Sconsia sublaevigata (western Atlantic) in Pliocene rocks of Darien, Paci®c Panama; and Distorsio constricta (eastern Paci®c) in latest Pliocene-Early Pleistocene rocks of Atlantic Costa Rica. Continued Early or middle Pleistocene connections are demonstrated by Cymatium cingulatum (now Atlantic) in the Armuelles Formation of Paci®c Panama. -
Early Pliocene Molluscs from the Easternmost Mediterranean Region (SE Turkey): Biostratigraphic, Ecostratigraphic, and Palaeobiogeographic Implications
Turkish Journal of Earth Sciences Turkish J Earth Sci (2017) 26: http://journals.tubitak.gov.tr/earth/ © TÜBİTAK Research Article doi:10.3906/yer-1705-2 Early Pliocene molluscs from the easternmost Mediterranean region (SE Turkey): biostratigraphic, ecostratigraphic, and palaeobiogeographic implications 1, 2 3 4 5 2 Yeşim BÜYÜKMERİÇ *, Erdoğan TEKİN , Erdal HERECE , Koray SÖZERİ , Nihal AKÇA , Baki VAROL 1 Department of Geological Engineering, Faculty of Engineering, Bülent Ecevit University, İncivez, Zonguldak, Turkey 2 Department of Geological Engineering, Faculty of Engineering, Ankara University, Gölbaşı Campus, Gölbaşı, Ankara, Turkey 3 Department of Geological Research, General Directorate of Mineral Research and Exploration (MTA), Balgat, Ankara, Turkey 4 Natural History Museum, General Directorate of Mineral Research and Exploration (MTA), Balgat, Ankara, Turkey 5 Research Centre, Turkish Petroleum Corporation (TPAO), Esentepe, Ankara, Turkey Received: 04.05.2017 Accepted/Published Online: 05.12.2017 Final Version: 00.00.2016 Abstract: The mollusc faunas from Pliocene deposits of the Hatay-İskenderun region were investigated at nine localities and complemented with three localities from earlier studies. The Pliocene units were deposited in three adjacent subbasins, Hatay-Samandağ (HS), Altınözü-Babatorun (AB), and İskenderun-Arsuz (İA); the first two are also known as the Hatay Graben. Basin configurations and shape, environmental evolution, and faunal compositions were affected by differential tectonic histories since the Late Miocene. In total 162 species (94 gastropod, 61 bivalve, and 7 scaphopod) are recorded, 80 of which are recorded for the first time from the region. The occurrence of tropical stenohaline benthic taxa (such as Persististrombus coronatus and some conid gastropod species) and a number of chronostratigraphically well-constrained mollusc species shows a Zanclean age. -
A Review of Fossil Bursidae and Their Use for Phylogeny Calibration
geodiversitas 2019 ● 41 ● 5 DIRECTEUR DE LA PUBLICATION : Bruno David, Président du Muséum national d’Histoire naturelle RÉDACTEUR EN CHEF / EDITOR-IN-CHIEF : Didier Merle ASSISTANTS DE RÉDACTION / ASSISTANT EDITORS : Emmanuel Côtez ([email protected]) ; Anne Mabille MISE EN PAGE / PAGE LAYOUT : Emmanuel Côtez COMITÉ SCIENTIFIQUE / SCIENTIFIC BOARD : Christine Argot (MNHN, Paris) Beatrix Azanza (Museo Nacional de Ciencias Naturales, Madrid) Raymond L. Bernor (Howard University, Washington DC) Alain Blieck (chercheur CNRS retraité, Haubourdin) Henning Blom (Uppsala University) Jean Broutin (UPMC, Paris) Gaël Clément (MNHN, Paris) Ted Daeschler (Academy of Natural Sciences, Philadelphie) Bruno David (MNHN, Paris) Gregory D. Edgecombe (The Natural History Museum, Londres) Ursula Göhlich (Natural History Museum Vienna) Jin Meng (American Museum of Natural History, New York) Brigitte Meyer-Berthaud (CIRAD, Montpellier) Zhu Min (Chinese Academy of Sciences, Pékin) Isabelle Rouget (UPMC, Paris) Sevket Sen (MNHN, Paris) Stanislav Štamberg (Museum of Eastern Bohemia, Hradec Králové) Paul Taylor (The Natural History Museum, Londres) COUVERTURE / COVER : Aquitanobursa tuberosa (Grateloup, 1833) n. comb., MNHN.F.A70285, Burdigalian of le Peloua, Staadt coll. Geodiversitas est indexé dans / Geodiversitas is indexed in: – Science Citation Index Expanded (SciSearch®) – ISI Alerting Services® – Current Contents® / Physical, Chemical, and Earth Sciences® – Scopus® Geodiversitas est distribué en version électronique par / Geodiversitas is distributed electronically -
Management Plan for Antarctic Specially Protected Area No. 126 BYERS PENINSULA, LIVINGSTON ISLAND, SOUTH SHETLAND ISLANDS
Measure 4 (2016) Management Plan for Antarctic Specially Protected Area No. 126 BYERS PENINSULA, LIVINGSTON ISLAND, SOUTH SHETLAND ISLANDS Introduction The primary reason for the designation of Byers Peninsula (latitude 62°34'35" S, longitude 61°13'07" W), Livingston Island, South Shetland Islands, as an Antarctic Specially Protected Area (ASPA) is to protect the terrestrial and lacustrine habitats within the Area. Byers Peninsula was originally designated as Specially Protected Area (SPA) No. 10 through Recommendation IV-10 in 1966. This area included the ice-free ground west of the western margin of the permanent ice sheet on Livingston Island, below Rotch Dome, as well as Window Island about 500 m off the northwest coast and five small ice-free areas on the south coast immediately to the east of Byers Peninsula. Values protected under the original designation included the diversity of plant and animal life, many invertebrates, a substantial population of southern elephant seals (Mirounga leonina), small colonies of Antarctic fur seals (Arctocephalus gazella), and the outstanding scientific values associated with such a large variety of plants and animals within a relatively small area. Designation as an SPA was terminated through Recommendation VIII-2 and redesignation as a Site of Special Scientific Interest (SSSI) was made through Recommendation VIII-4 (1975, SSSI No. 6). The new designation as an SSSI more specifically sought to protect four smaller ice-free sites on the peninsula of Jurassic and Cretaceous sedimentary and fossiliferous strata, considered of outstanding scientific value for study of the former link between Antarctica and other southern continents. Following a proposal by Chile and the United Kingdom, the SSSI was subsequently extended through Recommendation XVI-5 (1991) to include boundaries similar to those of the original SPA: i.e. -
Revision of the Subterranean Genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae)
A peer-reviewed open-access journal ZooKeys 769: 13–48 (2018) Revision of the subterranean genus Spelaeodiscus 13 doi: 10.3897/zookeys.769.25258 RESEARCH ARTICLE http://zookeys.pensoft.net Launched to accelerate biodiversity research Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae) Barna Páll-Gergely1, Tamás Deli2, Zoltán Péter Erőss3, Peter L. Reischütz4, Alexander Reischütz4, Zoltán Fehér5 1 Centre for Agricultural Research, Hungarian Academy of Sciences (MTA), Herman Ottó út 15, Budapest, H-1022, Hungary 2 Móricz Zsigmond u. 2, Gyomaendrőd, H-5500, Hungary 3 Bem u. 36., Budapest, H-1151, Hungary 4 Puechhaimg. 52, A-3580, Horn, Austria 5 Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H-1088, Hungary Corresponding author: Barna Páll-Gergely ([email protected]) Academic editor: Martin Haase | Received 25 March 2018 | Accepted 16 May 2018 | Published 26 June 2018 http://zoobank.org/C31B0F6B-D3C2-42CD-BAED-8CE9D5769E8A Citation: Páll-Gergely B, Deli T, Erőss ZP, Reischütz PL, Reischütz A, Fehér Z (2018) Revision of the subterranean genus Spelaeodiscus Brusina, 1886 (Gastropoda, Pulmonata, Spelaeodiscidae). ZooKeys 769: 13–48. https://doi. org/10.3897/zookeys.769.25258 Abstract The Balkan genus Spelaeodiscus Brusina, 1886 is revised based on museum collections and newly collected samples from Montenegro and Albania. The following species and subspecies are introduced as new to sci- ence: Spelaeodiscus albanicus edentatus Páll-Gergely & P. L. Reischütz, ssp. n. (southern Montenegro and northern Albania), Spelaeodiscus densecostatus Páll-Gergely & A. Reischütz, sp. n., Spelaeodiscus hunyadii Páll-Gergely & Deli, sp. n., Spelaeodiscus latecostatus Páll-Gergely & Erőss, sp. n. -
Cephalopod Id Guide for the Mediterranean Sea
CEPHALOPOD ID GUIDE FOR THE MEDITERRANEAN SEA Christian Drerup – University of Algarve, Portugal Dr Gavan Cooke – Anglia Ruskin University, UK PREFACE The intention of this guide is to help identifying cephalopod species in the Mediterranean Sea which you may find while SCUBA diving, snorkelling, a boat trip or even while walking along a rocky shore. It focuses on shallow water and subsurface-inhabiting species or those which at least partially spent their life in depths less than 50 meters. As you may encounter these animals in the wild most likely just for a short glance, we kept the description of each cephalopod rather simple and based on easy-to-spot external features. This guide was made within the scope of the project “Cephalopod Citizen Science”. This project tries to gather scientific information about the “daily life” of cephalopods by analysing pictures of those animals which were posted in several, project-related facebook groups. For further information about this project, please follow the link below: https://www.researchgate.net/project/Cephalopod-Citizen-Science We hope this guide will provide useful information and help you to identify those cephalopods you may encounter soon. Christian Drerup – University of Algarve, Portugal Dr. Gavan Cooke – Anglia-Ruskin-University, UK Common octopus (Octopus vulgaris). © dkfindout.com 2 CONTENTS HOW SHOULD WE INTERACT WITH CEPHALOPODS WHEN DIVING WITH THEM? ............... 4 SIGNS THAT A CEPHALOPODS IS UNHAPPY WITH YOUR PRESENCE .................................... 5 OCTOPUSES (OCTOPODA) -
BAP377-378 Text.Indd
NEOGENE TONNOIDEAN GASTROPODS OF TROPICAL AND SOUTH AMERICA: CONTRIBUTIONS TO THE DOMINICAN REPUBLIC AND PANAMA PALEONTOLOGY PROJECTS AND UPLIFT OF THE CENTRAL AMERICAN ISTHMUS A. G. Beu GNS Science, P. O. Box 30368, Lower Hutt, New Zealand 5040, email [email protected]. ABSTRACT Th e 142 species of tonnoidean gastropods recorded from the Neogene to Recent faunas of tropical America (Mexico, and a few taxa from Florida and California, south to Ecuador and Brazil) are revised, along with the 12 species of Neogene to Recent Personidae and Ranellidae occurring in Chile and Argentina. Taxa included are: (1) BURSIDAE: Bursa, 11 species, including the eastern Atlantic species B. scrobilator (Linnaeus, 1758) in the Pliocene and Pleistocene of tropical America; Aspa marginata (Gmelin, 1791), an eastern Atlantic species recorded in the Pliocene-Pleistocene of Limón, Costa Rica; Crossata, with one eastern Pacifi c species (California to Peru); Marsupina, fi ve species, including M. judensis n. sp. (Miocene, Punta Judas, Costa Rica). (2) PERSONIDAE: Distorsio, 12 species, including D. biangulata n. sp and D. jungi n. sp. (both Miocene, Cantaure, Venezuela); Personopsis, one Recent species. (3) RANELLIDAE, RANELLINAE: Argobuccinum, one species; Fusitriton, three species; Halgyrineum, one species; Priene, one Chilean species; Ranella, three species, including R. chilena n. sp. (Oligocene-Miocene, Chile); Ameranella, one species. (4) RANELLIDAE, CYMATIINAE: Cabestana, one living southwestern Atlantic species; Charonia, three species; Crassicymatium crassicordatum n. gen., n. sp. (Oligocene- Miocene, Chile); Cymatium, four species; Gelagna, one species; Gutturnium, one species; Linatella, one species; Monoplex, 31 species, including M. gatunicus n. sp. (Miocene, Panama), M. jackwinorum n. -
Tonnoidea, Gastropoda) Del Pliocè Del Mediterrani Nord-Occidental
Butll. Inst. Cat. Hist. Nat., 68: 50-71. 2000 ISSN: 1133-6889 GEA, FLORA ET FAUNA Sistemàtica i descriptiva de les famílies Ranellidae i Bursidae (Tonnoidea, Gastropoda) del Pliocè del Mediterrani nord-occidental Marta Solsona i Masana* Rebut: 02.09.99 Acceptat: 07.06.99 Resum MOTS CLAU: Ranellidae, Bursidae, Tonnoi- dea, Gastropoda, sistemàtica, Pliocè, Medi- Es revisa la sistemàtica de les espècies de les terrani, protoconquilla. famílies Ranellidae i Bursidae (Gastropoda) presents als dipòsits pliocens marins del Me- diterrani nord-occidental. Les espècies tractades són les següents: Cymatium corrugatum (La- Abstract marck, 1822), C. doderleini (D’Ancona, 1872), C. parthenopeum (von Salis, 1793) i Charonia Systematics and Description of the Ranellidae apenninica (Sassi, 1827) de la família Ranellidae i and Bursidae (Tonnoidea, Gastropoda) from Bufonaria marginata (Martini, 1777) de la família Bursidae. Aquesta revisió s’ha dut a terme a partir the Pliocene of the North-western Mediterra- d’exemplars recollits als sediments pliocens de les nean conques del Baix Llobregat i l’Alt Empordà (Catalunya) i del Rosselló i el Var (sud de França). A review of the systematics of the species of the Així mateix, s’ha examinat material comple- families Ranellidae and Bursidae (Gastropoda) mentari que prové de les principals conques from the Pliocene marine sediments of the north- miocenes i pliocenes del Mediterrani occidental i western Mediterranean is presented. The species l’Atlàntic nord-oriental, la cual cosa ha permès treated are the following: Cymatium corrugatum d’establir amb precisió la distribució (Lamarck, 1822), C. doderleini (D’Ancona, cronostratigràfica i geogràfica de les espècies 1872), C. -
Marine Vertebrates from Pliocene Shell Beds from Tuscany (Italy): Prospecting, Taphonomy, Palaeoecology and Systematic Palaeontology
Università di Pisa, Dipartimento di Scienze della Terra Scuola di Dottorato in Scienze di Base “Galileo Galilei” Programma di Scienze della Terra MARINE VERTEBRATES FROM PLIOCENE SHELL BEDS FROM TUSCANY (ITALY): PROSPECTING, TAPHONOMY, PALAEOECOLOGY AND SYSTEMATIC PALAEONTOLOGY SETTORE SCIENTIFICO DISCIPLINARE: GEO (01) Candidato: Chiara Tinelli Tutore: Dott. Giovanni Bianucci XXV CICLO A.A. 2012-2013 INDEX Abstract 1 Riassunto 4 Chapter 1 – GENERAL INTRODUCTION 9 Chapter 2 – GEOLOGICAL AND PALAEOECOLOGICAL SETTING 16 2.1 Geological, stratigraphical and palaeontological context of the Arcille area 16 2.2 Geological, stratigraphical and palaeontological context of the Montalcino area 28 2.3 Taphonomy and Palaeoecology 36 2.3.1 Material and Methods 36 2.3.2 Taphonomy of the Arcille area 38 2.3.3 Taphonomy of the Montalcino area 44 2.4 Discussion and Conclusion 47 Chapter 3 – SYSTEMATIC PALAEONTOLOGY 59 3.1 Introduction 59 3.2 Material and Methods 59 3.3 Description 61 3.4 Comparison 85 Chapter 4 – GEORADAR AND PALAEONTOLOGY 93 4.1 Introduction 93 4.2 The palaeontological site 96 4.3 Material and Methods 98 4.4 Results 102 4.5 Discussion and Conclusion 108 CAPITOLO 5 – GENERAL CONCLUSIONS 111 REFERENCES 116 ANATOMICAL PLATES 137 APPENDICES 147 - Appendix 1 147 - Appendix 2 151 - Appendix 3 163 Abstract Marine vertebrates from early Pliocene shell-beds have been studied in two areas located in southern Tuscany (Italy). In particular these shell-beds observed in four sedimentary successions outcropping in Arcille and Montalcino localities, occur at the transition between sandstones and mudstones. Marine vertebrates were found in correspondence of these laterally continuous shell-beds: three nearly complete Metaxytherium subapenninum specimens occur in Arcille succession, whereas a nearly complete skeleton of a Mysticete, an incomplete Metaxytherium cf. -
An Ochered Fossil Marine Shell from the Mousterian of Fumane Cave, Italy Marco Peresani, Marian Vanhaeren, Ermanno Quaggiotto, Alain Queffelec, Francesco D’Errico
An ochered fossil marine shell from the Mousterian of Fumane Cave, Italy Marco Peresani, Marian Vanhaeren, Ermanno Quaggiotto, Alain Queffelec, Francesco d’Errico To cite this version: Marco Peresani, Marian Vanhaeren, Ermanno Quaggiotto, Alain Queffelec, Francesco d’Errico. An ochered fossil marine shell from the Mousterian of Fumane Cave, Italy. PLoS ONE, Public Library of Science, 2013, 8 (7), pp.e68572. 10.1371/journal.pone.0068572. hal-01842752 HAL Id: hal-01842752 https://hal.archives-ouvertes.fr/hal-01842752 Submitted on 11 Jun 2020 HAL is a multi-disciplinary open access L’archive ouverte pluridisciplinaire HAL, est archive for the deposit and dissemination of sci- destinée au dépôt et à la diffusion de documents entific research documents, whether they are pub- scientifiques de niveau recherche, publiés ou non, lished or not. The documents may come from émanant des établissements d’enseignement et de teaching and research institutions in France or recherche français ou étrangers, des laboratoires abroad, or from public or private research centers. publics ou privés. An Ochered Fossil Marine Shell From the Mousterian of Fumane Cave, Italy Marco Peresani1*, Marian Vanhaeren2, Ermanno Quaggiotto3, Alain Queffelec2, Francesco d’Errico2,4 1 Dipartimento di Studi Umanistici, Sezione di Preistoria e Antropologia, Università di Ferrara, Corso Ercole I d’Este 32, I-44100 Ferrara, Italy, 2 CNRS UMR 5199 PACEA-PPP, Université Bordeaux 1, Avenue des Facultés, F-33405 Talence, France, 3 Naturalistic-Archaeological Museum of Vicenza, Contrà S. Corona, 4, I-36100 Vicenza, Italy, 4 Department of Archaeology, Cultural and Religious Studies, University of Bergen, Postboks 7805, 5020 Bergen, Norway Abstract A scanty but varied ensemble of finds challenges the idea that Neandertal material culture was essentially static and did not include symbolic items.