SMITHSONIAN CONTRIBUTIONS TO BOTANY NUMBER 12

An Introduction to the Botanical Type Specimen Register

Stanwyn G. Shetler

with Mary Jane Petrini, Constance Graham Carley, M. J. Harvey, Larry E. Morse, Thomas E. Kopfler, and Collaborators

SMITHSONIAN INSTITUTION PRESS City of Washington 1973 ABSTRACT

Shetler, Stanwyn G., with Mary Jane Petrini, Constance Graham Carley, M. J. Harvey, Larry E. Morse, Thomas E. Kopfler, and Collaborators. An Introduc- tion to the Botanical Type Specimen Register. Srnithsonian Contributions to Botany, number 12, 186 pages, 3 figures, frontispiece, 1973.--In the first part, the development of a computer-based system for storing and retrieving infonna- tion about botanical type specimens is described from its pilot stage to its present operational stage. The concept, purpose, and scope are explained, and the operational procedures are outlined. Ways of using and contributing to this computerized register of types, both in the short-run and in the longrun, are proposed. A statistical summary of the content of the Type Register as of 30 September 1972 is given. Over 13,000 specimens representing more than 10,000 taxa have been registered. The second part consists of a Catalog of more than' 1,000 specimens representing over 600 taxa of the genus Carex (Cyperaceae), which are deposited in ten major American herbaria, and the Catalog is cross- indexed five different ways: by author, publication date, collector, country, and herbarium. An introduction summarizes the preparation and editing of the Catalog. This Carex Catalog represents the first published installment of the Type Register and as such is intended to serve as an example.

COLLABORATORS Listed here by institution are the persons who have collaborated in the compiling of data from their respective herbaria for the Carex Catalog: California Academy of Sciences, Golden Gate Park, Sun Francisco, California 94118; Stanford University (Dudley Herbarium), Stanford, CaGifornia 94305; University of Cali- fornia (Herbarium), Berkeley, California 94720 Dr. John H. Thomas, Associate Professor of Biological Sciences and Curator of Dudley Herbarium (Stanford), Curator of Botany (Acad-my) , and compiler of data at Berkeley; assisted by Dr. Margaret Sharp Field Museum of Xatural History, Roosevelt Road at Lakeshore Drive, Chicago, Illinois 60605 Dr. Lt'illiam C. Burger, Associate Curator of Vascular Plants: assisted by Mr. Ronald L. Liesner Harvard University, 22 Divinity Avenue, Cam bridge, Massachusetts 02138 Dr. Richard 4. Howard, Director, Arnold Arboretum; Dr. Reed C. Rollins, Asa Gray Professor of Systematic Botany and Director of the Gray Herbarium Missouri Botanical Garden, 2315 Tower Grove Avenue, St. Louis, Missouri 63110 Dr. IValter H. Lewis, Senior Botanist, Professor of Biology (Washington University); assisted by Dr. Raymond F. Altevogt h'ew York Botanical Garden, Bronx Park, New York 10458 Dr. Howard S. Irwin, Director: Dr. Patricia Kern Holmgren, Associate Curator and Herbarium Administrator; Dr. Noel H. Holmgren, Associate Curator Smithsonian Institution, Washington, D. C. 20560 Dr. Mason E. Hale, Curator of Cryptogams; Dr. Dan H. Nicolson, Associate Curator of Phanerogams

OFFICIALPUBLICATION DATE is handstamped in a limited number of initial copies and is re- corded in the Institution's annual report, Smithsonian Year. SI PRESS NUMBER 4783. SERIES COVFR DESIGN: Leaf clearing from the katsura tree Cercidiphjlluin japonicum Siebold and luccarini.

Library of Congress Cataloging in Publication Data Shetler, Stanwyn G. An introduction to the botanical type specimen register. (Smithsonian contributions to botany, no. 12.) Bibliography: p. 1, Information storage and retrieval systems-Botany-Type specimens. 2. Carex. I. Title. 11. Title: Botanical tjpe specimen register. 111. Series: Smithsonian Institution. Smithsonian contributions to botany, no. 12. QKlS2747 no. 12 [Z699.5.B6] 58lt.08s[579] 72-11706

For sale by the Superintendent of Documents, US. Government Printing Office, Washington. D.C. 20402 Price $2.85 domestic postpaid or $2.50 GPO Bookstore Foreword

It always has been the policy of the United States National Herbarium to make its collections as easily available as possible. We have welcomed visitors and loaned specimens on request since the founding of our herbarium. Now, in an effort to make our collections even more accessible, we have undertaken a new project to compile a computerized catalog of our type collection of ap- proximately 65,000 specimens. Eventually, we hope to broaden this catalog, which we are calling the “Botanical Type Specimen Register,” to include the type collections of many other institutions so that it will serve as a union listing of types. Already we have enlisted the cooperation of other institutions, and the computer file presently includes records from more than a score of herbaria. The United States National Herbarium, a worldwide collection of plants now totaling some 3 million specimens, is administered by the Smithsonian Institu- tion’s Department of Botany, a unit of the National Museum of Natural History. The Department of Botany has played a pioneering role in the development of the Museum’s active program in data processing, The Type Register is the Museum’s first operational effort in cooperative, multi-institutional (network) data banking and, as such, is of special interest. If this approach to common data banking proves successful, it \vill point the way for many cooperative efforts in other branches of natural history. Although the Type Register is still very much in its infancy, we are zealous to demonstrate its potential to the botanical community with a tangible product so that we can receive advice and counsel from the community on the basis of concrete results while the data bank is still small and susceptible to modifica- tion. This publication should prove useful in itself as a catalog of type speci- mens of Cayex, particularly to specialists on the family Cyperaceae. The larger purpose, however, in issuing a preliminary catalog of limited scope at this time is to demonstrate the concept of the Type Register in concrete terms and thereby to solicit the collaboration of all plant systematists in molding the Register into an effective, scholarly tool for future generations of the profession. The computer file presently registers over 13,500 type specimens, representing some 10,500 vascular plant taxa. Thus the Cawx Catalog, with its 1,000 speci- mens and 600 taxa, is a printout of less than 10 percent of the current, rapidly growing file. Less formal and less expensive means of putting out the informa- tion will be tried with future installments, and it may become desirable or neces- sary at some point to begin publishing in microform. Perhaps the most common and economical mode of disseminating the accumulated information will be to provide computer printouts to individual users in response to queries for up-to- the-minute reports on specific taxonomic groups. Once the data bank is well established query service can be provided to any user for a modest fee. We welcome your reaction to the concept of the Botanical Type Specimen Register on the basis of this sample. Only with the backing of the botanical community can we continue to get the necessary financial support to carry on the work.

EDWARDS. AYENSU,Chairman Department of Botany 30 September 1972

iii

Con tents Page Foreword ...... iii THEBOTANICAL TYPE SPECIMEN REGISTER Stanwyn G. Shetler Introduction ...... 1 Acknowledgments ...... 3 Concept and Purpose of Type Register ...... , ...... 6

Scope of Register ...... , ...... , ...... I..... 10 Procedures and Standards ,...... 12 Source of Data ,,,,...... ,...... 12 Instructions for Contributors . . . , . , ...... , . . . , , . . 12

Record Format and Content ,..... , .,...... ,...... 15 Level I-Taxon Data ...... 16 Family , ,...... ,...... 16 Genus a ..... ,...... It...... 16 Species .,...... 16 Infraspecific Taxon , , . , . . . , ...... 17 Author ...... 17 Title ...... 17 Level 2-Collection Data , ...... 17 Collector(s) ...... 17 Collection Number . , ...... , , , . . . , . . . . , ...... , , . . , . . . . , ...... 18 Collection Date(s) ,...... , ...... 18 Geographic Data Fields ...... 18 Level 3Specimen Data , . . . , ...... , , , , . . . . , , ...... , ...... , . 19 Herbarium Abbreviation , . . . . 19 Herbarium Sheet Number , , . , . . . . , . . . , . . . , . , , . . . . . , ...... 19 Kind of Type , , , , . , . . 19 Data Source Code . . , ...... 21 Processing System . . . , . . . , . . , , . . . , . . , , . . . . . , , , ...... 21 Paper Tape System ...... 22

On-Line System , . . . , , , . . , , , , , , , . , , , , , , . , , , , . , . , . . , , , . . , , , . 22 Optical Scanning System , , , , , , , , , . . , . , . , , , , . , , . . . . . , , ...... 22 Magnetic Tape System , , , , , . , ,,...... 22

Computer Processing System , . , , ...... 22 Statistical Summary of Type Register Contents , . . . , , , . . , , , , , . , , . , . , , ...... 23 Use of Type Register , , , , . . . , . . . , ...... , . , ...... , ...... 25

A CATALOGOF THE GENUSCarex (CYPERACEAE) Stanwyn G. Shetler (Editor): Mary Jane Petrini, Constance Graham Carley, M. J. Harvey, Larry E. Morse (Assistant Editors) ; Thomas E. Kopfler (Programmer) : and Collaborators

Contributing Institutions , ...... 26 Data-Collection Procedure ...... 27 Editorial Process ...... 28 Milestone Events ...... 29 Use of Carex Catalog . . . . 31 Errata ...... 33 Statistical Summary of Catalog ...... , ...... , ...... 33 Catalog of Specimens ...... 34 Author Index ...... 105

Publication-Date Index . . . . . , ...... , ...... 118 Collector Index . , . . . . . , ...... I...... 131 Geographic Index . , . , . . . 149 Herbarium Index ...... 162 Bibliography . , . . . , , ...... 185 V

An Introduction to the Botanical Type Specimen Register’

Stanwyn G. Shetler, with Mary Jane Petrini, Constance Graham Carley, M. J. Harvey, Larry E. Morse, Thomas E. Kopfler, and Collaborators

The Botanical Type Specimen Register

STANWYNG. SHETLER

Introduction The Botanical Type Specimen Register is a The Type Register was conceived by Mason E. computer-based system for recording information Hale, who organized a pilot project in 1968 while about type specimens of plant species and infraspe- he was chairman of the Smithsonian’s Department cific taxa, which is designed to become a union reg- of Botany. This was one of several projects that istry of type holdings in the world’s herbaria. It Hale initiated as chairman to introduce electronic introduces a new dimension to the management of data processing (EDP) to collection management herbarium collections. Through the use of advanced in the United States National Herbarium. (For a information processing methods, critical data are description of his automated system for recording compiled from the herbarium and library and specimen exchanges, see Hale and Creighton, 1970; merged into a common, machine-searchable file the pilot Type Register project is described in from which catalogs can be printed or special Shetler et al., 1971.) At the outset, the author and queries, defined by complex selection criteria, can Flora North America (FNA) personnel assisted in be answered rapidly on demand. As a result, future the development of the pilot system, and the proj- taxonomic investigators will be able to learn the ect, though separate, has continued until the pres- whereabouts of type specimens of particular con- ent to be associated closely with the FNA program cern to them and obtain answers to certain basic (Shetler and Meadow, 1971). After launching the questions without painstaking research or time- pilot effort, Hale, while continuing his involvement consuming travel or correspondence. on an advisory level, passed the supervisory re- sponsibility to the author, who continues to direct Stanwyn G. Shetler, Department of Botany, Smithsonian the work. Institution, Washington, D.C. 20.560; same address: May The project was initiated with special funds. In Jane Petrini; Constance Graham Carley (nee Graham) fiscal year 1969 and 1970, limited allocations of (present address: 275 Collier Rd., Atlanta, Georgia 30309) ; M. J. Harvey (present address: Department of Biology, Dal- regular funds of the National Museum of Natural housie University, Halifax, Nova Scotia) ; Larry E. Morse History were made through the Department of (present address: Biological Laboratories, Harriard University, Botany, but regular budgeting did not begin until Cambridge, Massachusetts 02138) ; Thomas E. Kopfler. FY-71 (1 July 1970-30 June 1971) when the Mu-

~~ ‘This is No. 70 in the Flora North America Report seum organized a new, Museum-wide program in series. data processing and incorporated the Type Regis- 1 ShIITHSOSIAN COSTRIBUTIOSS TO BOTANY ter as one of the charter projects. These projects mation Systems Division, and since the project has are controlled by the respective departments, but become operational this work has been performed the overall program is coordinated and managed on a similar basis by FNA personnel so as to keep by James F. Mello, Assistant Director. the Type Register system compatible with the FNA The response to the several requests for participa- system. tion during the pilot phase was excellent, showing From the beginning Hale planned for the in- general interest among botanists and convincing clusion of data from an indefinite and constantly Hale and others at the Smithsonian of the potential growing number of other institutions and actively value of a type-specimen register and of the bo- sought such collaboration. In one test of the feasi- tanical community’s willingness to collaborate to bility of multi-institutional input, he distributed the fullest extent possible in creating a register a computer-printed set of 52 cards, representing a along the lines conceived. From the preliminary card catalog of the National Herbarium’s complete experiments much was learned about the logistical type holdings in the genus Mimulus (Scrophu- problems and the costs and manpower requirements lariaceae), to each of 50 large herbaria in the of network data banking. The Smithsonian neces- United States and abroad and solicited their co- sarily is concentrating present operations on its own operation in providing similar data from their own type collections ot about 65,000 specimens. The type collections, if any, of Mimulus. A second ma- system is designed, however, to accept data from jor test involved sending a computer-printed card any institution at any time, and as long as the catalog of the National Herbarium’s complete type inflow remains on a small scale, data from other holdings in the family Lamiaceae (Labiatae) to the institutions can be added to the central file by the University of California, Los Angeles, where Carl paresent staff more or less as they are received. It Epling’s extensive t) pe collections in this family are is hoped that as interest in collaboration grows deposited, and later to the Missouri Botanical Gar- support commensurate to the interest will be forth- den and the New York Botanical Garden. All three coming both for the central operation and for institutions cooperated in providing data from participating institutions. their own collections of types in this family. The The second part of this paper represents the first latter two institutions continued thereafter to col- published installment of the Type Register. It is laborate as much as possible on other taxonomic a provisional union catalog of the type specimens groups, and, apart from the Smithsonian itself, of the genus Carex on deposit in ten major Ameri- they have been the institutions with the greatest can herbaria (see “Contributing Institutions”) and involvement in the Type Register project. a concrete example of cooperative data banking. Certainly, no claim to completeness can be made In FY-72, the year that ended on 30 June 1972, for a catalog that concerns a single genus and only the Type Register project finally was put on a solid one percent (10/1000) of the world’s public insti- footing. Midway through this year it was possible tutional herbaria (Shetler, 1969). “Usefulness” is for the first time in the four-year history of the the pragmatic criterion for compilation and pub- project to staff adequately by employing three full- lication of the Type Register, however, and useful- time persons (two assistant editors and a data con- ness is dependent on critical mass, not absolute version operator) and also to have adequate funds coverage. Clearly it is unrealistic to think that the to process regularly. Prior to January 1970, the Type Register could ever achieve absolute com- pilot project was carried on intermittently as funds pleteness, registering all type specimens for all pub- were available by one to three part-time employees, lished taxa in all of the world’s herbaria, and the and from January 1970 to December 1971 the proj- Register has not been conceived on this false prem- ect advanced on a more or less continuous, opera- ise. The Register is being created on the assump- tional basis with the assistance of one, two, and tion that some information is better than no occasionally three full-time persons, the number information and that a catalog of ten type collec- depending again on available money. During the tions is more useful than a catalog of one type pilot effort, the computer analysis, programming, collection. Although it must be admitted that the and file processing were done on a part-time basis, coverage is very uneven among the ten herbaria as needed, by personnel of the Smithsonian Infor- contributing to the Carex Catalog, for example, NUMBER 12 nevertheless this Catalog tells us more than we have always share his vision nor his optimism for its ever known before about the Cal-ex type collections potential value. The continuing support of Rich- of the participating institutions and provides a ard S. Cowan, former Director, National Museum solid framework to which new information can be of Natural History, and of Hale’s successor as chair- added as it becomes available from these or any man, Edward S. Ayensu, have been crucial in put- other institutions. ing the project on a stable footing. The Assistant The concept of a type register is not new. Al- Director, James F. Mello, and his assistant, David ready in the mid-l930s, A. S. Hitchcock of the Bridge, deserve much credit for laying the budget- Smithsonian Institution, in his capacity as chair- ary groundwork that has brought the project to its man of the Committee on Nomenclature of the present viable and relatively healthy state, and for Botanical Society of America, coordinated the com- facilitating its administration. From his vantage pilation of information on the location of type point as overseer of all EDP projects in the Na- specimens. Lists of authors of new names, indicat- tional Museum of Natural History, including the ing the major group(s) of plants they described Type Register, RIello has provided stimulating and (e.g., phanerogams) and the herbaria where they \vise counsel, as well as constant encouragement and deposited their types, were compiled (Hitchcock help. et al., 1934, 1935). Other members of the Com- Many curators have contributed in some way to mittee in 1934 were L. R. Abrams, J. C. Arthur, the development of the Type Register thus far, A. TV. Evans, J. &I. Greenman, $1. A. Howe, E. D. especially in the course of the multi-institutional Nerrill, F. TY. Pennell, and C. L. Shear. “The Com- data-collecting experiment with MimuEus, con- mittee is not attempting to decide what specimens ducted by Hale, and their cooperation is hereby are types nor to determine the identity of types,” gratefully acknowledged. Those curators and their Hitchcock wrote in 1934; “it is attempting only to assistants who participated directly in the compila- aid botanists in their search for types.” His words tion of data for the Carex Catalog are listed earlier can only be reiterated in the present context. Other as “Collaborators.” Among them, Hale, Irwin, recent efforts to catalog types have been made in Lewis, and Nicolson have taken a dedicated per- connection with specimen-data retrieval projects in sonal interest in the success of the Type Register the herbarium or museum (e.g., Beschel and Soper, from the beginning and have in effect constituted 1970; Collier, 1971; Crovello, 1972). a standing advisory editorial board. In their re- Information processing technology has advanced spective institutions, they have played a role in all far since the days of the first applications in biol- of the data compilation that has been done for the ogy, when the limitations of the computer led to Register thus far, regardless of the taxonomic some unfortunate consequences, as thoughtfully group. As a Smithsonian colleague, Nicolson has analyzed, for example, in reviews by Wood et al. been a steadfast supporter of the project, showing (1963) and Rollins (1966) of some early applica- deep interest in the work itself and sharing his time tions in plant taxonomy. We make no pretense of and seemingly inexhaustible nomenclatural knowl- having avoided all the pitfalls cataloged by these edge willingly and unselfishly whenever there has reviewers, but we have tried to make good use of been need, which often has been daily. John H. their advice. If we have learned anything so far, Thomas raised enthusiasm for the Type Register we have learned that no one can design the perfect to a new level wrhen, as a result of his collabora- system on the first trial. Every operational system tion on the Cal-ex Catalog, he began to ask special- is at the same time a pilot system for an even more advanced and refined, next-generation system. ists borrowing from the collections he curates to compile data for the Register from type specimens kKsOWLEDChfENTs.-Without the inspiration and genius of Rlason E. Hale the Type Register loaned to them and to affix an annotation label of would not exist. 1Yhile chairman of the Smith- his own design to each, which reads, “The written sonian’s Department of Botany (1967-69), he had itiformation on this specimen has been abstracted not only the foresight to inaugurate this computer for the TYPE REGISTER PROJECT by - application but also the fortitude to persist with on administrative support for it when others did not 4 SMITHSOSIAS CONTRIBUTIONS TO BOTANY

\Vitli respect to the editorship of the Carex Cata- laboration in the Type Register project. Mildred log, Carley, who served the FNA program and the E. klathias, Director, Botanical Gardens-Herba- Type Register project from December 1969 to April I ium, supervised the compilation of data on types 1971, and Petrini, who is the senior technical editor of the family Lamiaceae at the University of Cali- of the Type Register project, assisted Shetler with fornia, Los Angeles, early in the project. Bruce the clay-today technical editing. Harvey and Morse RIacBrJde assisted in the compilation of the data assisted him with the botanical editing during the lor this family and for the genus MimuZus at the )ear that each spent working at the Smithsonian on Missouri Botanical Garden. Other botanists who the FNA program-1969-50 and 197 1-72, respec- deserve mention for playing a part in the project ti\elj. Harvey provided botanical supervision of at the Smithsonian are Amy Jean Gilmzrtin, Mon- the data-capture operation during the initial input terey Peninsula College, Monterey, California, and of Cn?ex records from the Missouri Botanical Gar- Milosla\ Kovanda, Czechoslovak Academy of Sci- den, New York Botanical Garden, and the U.S. ences, Prague, .izho spent the years 1969-70 and National Herbarium, He directed the two-week, 1970-5 1, respectively, with the FNA program. on-site input effort at the New York Botanical Gar- The FNX Editorial Committee (John H. Bea- den in June 1970, Tvhich involved other genera man, M’alter H. Lewis, John McNeill, John T. besides Carex. Cynthia N. Ostroff of the Index ;\lickel, Peter H. Raven, Stanwyn G. Shetler, Roy Sominuin Generzcoi urn project assisted part-time L. Taylor, John H. Thomas) has taken a deep at the Smithsonian in the technical editing of the interest in the Type Register project from the out- Cai ex data from Harvard University. Marilyn An- set and provided financial and material support draeson helped with the data compilation at the through the FNA program, as well as invaluable I\lissouri Botanical Garden, while similar assistance advice and encouragement. A report on progress was rendered by Zella Ellshoff, Robert Helliwig, has been given at every meeting of the Committee and Gail Johnson at the New York Botanical Gar- since the Register was organized, and much time den. has been devoted to evaluation of the present and At least part of the Carex Catalog was examined future development of the Register. in near final form by Frederick J. Hermann of the The pilot processing system was designed by U.S. Forest Service Herbarium at Fort Collins, Reginald Creighton, Manager of Information Stor- Colorado, and Tetsuo Koyama of the New York age and Indexing, and programmed by Willard Botanical Garden, specialists on Carex and the Handley, both of the Smithsonian’s Information family Cyperaceae, respectively. Although both Systems Division. Creighton worked closely with provided helpful comments, they should not be Hale to develop the pilot system, which served the held responsible in any way for the final Catalog, original purposes well, and the Type Register because neither was able to devote the enormous would not exist if it had not been for Creighton’s amount of time that would have been required to pioneering insight and dedication. When the proj- check the file authoritatively. This responsibility ect became more closely associated with the FNA rests with the editors and collaborators. systems effort, Harriet R. Meadow, Systems Devel- Several specialists at the New York Botanical opment Manager of ENA, designed the present op- Garden have provided data to the Type Register erational sjstem in its general outlines. Kopfler for groups other than Carex: Caroline Allen (Lau- has been responsible for the detailed design and raceae: a few records of selected taxa), Patricia Kern Holmgren (Brassicaceae: Draba, Thlaspi), FIGURE1.-Stages in the preparation of records for the Type Rcgister: a, Constance Carley checks nomenclatural data in Tetsuo Koyama (Cyperaceae: a few records of se- the Gray Herbarium Card Index: b, Mary Jane Petrini lected taxa), John T. Mickel (Schizaeaceae: Ane- enters corrections into the computer via a remote typewriter mia subgenus Coptophyllum and segregate genera), terminal connected by telephone (photos a and b by Walter and Ghillean T. Prance (Chrysobalanceae, Di- G. Peter 111); c, type specimens of the genus Viola (Violaceae) chapetalaceae). (See also “Statistical Summary of are examined and discussed with respect to the problems of recording them in the Type Register by (left to right) M. J. Type Register Contents.”) Holmgren has been the Harvey, John T. Mickel, and Harriet R. Meadow in the one chiefly responsible for coordination at the work- herbarium at the New York Botanical Garden (photo by ing level of the New York Botanical Garden’s col- New York Botanical Garden). NUMBER 12 5 SMITHSONIAN COSTRIBUTIONS TO BOTANY programming and for the maintenance and query- describing new plant taxa and establishing their ing of the machine files. Morse wrote the COBOL types are myriad and scattered through the world's program for concatenating the data into the para- literature. The taxonomist who wishes to make a graph form used in the Carex Catalog. Meadow's scholarly study of a group of species, including an analysis and design resolved some basic unforeseen investigation of their typification, faces the for- problems that arose after using the pilot system for midable task of locating the relevant original de- a time, and her efforts resulted in the operational scriptions and type specimens. Fortunately for him, system that continues to serve very well. She has two standard indices, Index Kewensis (Rouleau, played a crucial role in the success of the project 1970; Meikle, 1971) and the Gray Herbarium and continues to provide advice and guidance on (Card) Index (Shaw, 1971), and various standard matters of systems development. Several botanists library catalogs and union listings are readily avail- who also have experience with computer applica- able to guide him to the pertinent literature. No tions have provided valuable advice trom time to similar indices exist, however, to guide the taxono- time: Theodore J. Crovello, University of Notre mist to the pertinent type specimens. To find types Dame; David J. Rogers, University of Colorado; he first must search the original literature case by and James H. Soper, National Museum of Natural case for indications or clues and then, through Sciences, Ottawa. travel or correspondence, continue his search in Technical help in editing and capturing data herbaria among the specimens themselves. Even has been given by the following persons, listed in with the original descriptions in hand the specialist the order in which they were hired, some as em- often faces great difficulties in trying to determine ployees of FNA and others as employees of the where the types are deposited. The modern litera- Type Register project: John Bolduc, Nancy How- ture still shows an astonishing lack of standardiza- ard, Barbara Bryant, Grace Rickard, Edna Mont- tion in the way types are designated and their ford, Mary Beth Moore, Barbara Halter, Gudrun herbarium deposition indicated. Christenson, and Rita Abessinio. Julia E. Taylor 'IVhat the taxonomist needs, therefore, is a find- and Liliosa Mangosing Evangelista have been cheer- efs guide to the type holdings of at least the world's fully indispensable in discharging the myriad major herbaria. This guide should be indexed pri- clerical and administrative tasks without which a marily by taxon but also cross-indexed several ways, project of this nature could never succeed. and it should include citations of the original pub- In addition to receiving regular budgetary sup- lications and basic collection data as provided by port on an increasing scale from the Smithsonian the specimen label and/or published description. Institution, the Type Register project has been Such an index ultimately would incorporate and funded in part by the National Science Foundation enhance the functions of Index Kewensis and the through grants made to the American Institute of Gray Hcibar.ium Index. This type of registry could Biological Sciences (GB-844 1, GN-8 12, GB-26 173) become effective at once for newly published taxa and to the Smithsonian Institution (GB-31715 and if taxonomists would agree to require registration contract C-720) for the FNA program, the Smith- of all new taxa and type depositions at the time of sonian Research Foundation (grants Sg062 1054, publication as a condition for effective publication. Sg0621054/Cl and Sg0621054/C2), and the Smith- Furthermore, the registry, if computerized, would sonian Office of Systematics. be the logical central repository for specialists' an- notations on typification, especially with respect to lectotypification and neotypification. Plant taxon- Concept and Purpose of Type Register omy desperately needs an effective central place and Perhaps 200 million specimens are on deposit in straight-forward procedure for registering lecto- the more than one thousand public herbaria in the types, neotypes, and specialists' conclusions about world, and scattered among these vast collections other kinds of types. may be as many as 4 million type specimens (Shet- The Botanical Type Specimen Register, as con- ler, 1969; Shetler et al., 1971). North American ceived, therefore, is to serve primarily as a finder's herbaria alone probably contain more than a half- guide for locating type specimens. The secondary million types. Likewise the original publications function, however, is to serve as a guide to the SUMBER 12 7 original descriptions of the registered taxa and to card file or published index is used, however, there provide collection data for the registered specimens. is a need to verify the information subsequently by The specimen information necessarily is organized checking the specimens and original descriptions. by taxonomic name. Thus the file is not purely a Two examples will illustrate. (1) Index Kewensis specimen register; the data are organized so as to and the Gray Herbal ium Index constitute indis- constitute a three-level hierarchy: taxon, collection, pensable secondary sources of references to original specimen. A full entry (record) in the Register is, publications, but the original publications them- therefore, a synthesis of data from the herbarium selves must be examined in order to verify both and the library which cannot be completed with- the existence of the descriptions and the accuracy out examining the original specimen and the origi- of the citations. (2) Present-day revisions and nal publication, as well as other specimens and monographs customarily indicate where the key publications as necessary. A record can be initi- type specimens are deposited, but this information ated with data from either the herbarium or the is not considered verified, for purposes of the Regis- library and supplemented later by data from the ter, until the specimens have been seen in the other. To a degree this happens naturally as other process of compiling the data for the Register, be- institutions contribute to the initial record, but in cause the data must be verified in the context of any event years may elapse before all records of a the specific requirements of the Register. This is given taxonomic group can be brought to relative a critical point. completeness. It should be emphasized here that a taxonomic The development of the Type Register ulti- revision or monograph, no matter how carefully mately involves three stages: registration, verifica- and authoritatively executed, constitutes a second- tion, and validation. ary source of information for all taxa treated except The immediate objective is to initiate the com- those being described to science for the first time. puterized working file without getting bogged down In tact, as synthetic works, these treatises often in time-consuming researches to resolve challenges present only the barest details on type specimens, of the validity of the data. Such scholarly research especially tor previously described but even for is the province of specialists who may need years newly described taxa, and the author’s own nomen- to resolve particularly difficult nomenclatural and clatural interpretations frequently are not clearly typification questions, and the preparation of the distinguished from the original data. It is not un- kind of index envisaged here could never be ac- usual, for example, to discover in the process of complished if all questions had to be answered first verifying a record in the Type Register that the and the compiled data had to be “perfect.” In the author of a revision or monograph, in identifying first stage, therefore, the primary goal is to record what he believed to be the holotype, unwittingly or register the facts more or less at face value as designated a lectotype or even a neotype by strict they are given by the available primary and/or application of the international rules of botanical secondary sources. Research and editing are kept ncmenclature. Therefore, except as a reference to to a minimum except where obvious discrepancies the original sources, for which it is of course in- can be resolved without extensive investigation. valuable, the monographic treatise has proved to be The editing is restricted largely to formatting the a disappointing starting point for compiling the data according to the technical standards of the Type Register; it seldom provides all necessary data processing system, and to standardizing the use of and often presents summaries of the original facts names, titles, and terms in key fields. which are telegraphic to the point of being impre- The second stage is to verify all data by firsthand cise or even inaccurate. On the other hand, the examination of the pertinent specimens and origi- monographic treatise is indispensable in the third nal publications. When the data are compiled in or validation stage, because it deals in a systematic the first place from the original sources expressly way with the typification of the taxa covered and for the Type Register, verification is accomplished establishes authoritative precedents that must be in the process of preparing the data for registra- considered in the interpretation of the information tion, and the two steps merge into a single opera- in the Type Register. Furthermore, short of hav- tion. When a secondary source such as an existing ing all the original references and specimens in S\lITHSOSIAN COETRIBUTIONS TO BOTANY hand, the monograph, which brings together all the The long-term goal of the third and ultimate data for a taxonomic group into one place, is by stage in the development of the Type Register is far the best single source of data for the given to ualzdate the data according to the rules of the group. Zntu natzonal Code of Botanical Nomenclature Verification is not a simple procedure that can (ZCBIY; Stafleu et al., 1972) and thereby to estab- be accomplished once for all time but an involved, lish the Register as a wholly reliable, authoritative virtually never-ending process, which seems to ex- index of types of plant species and infraspecific pand in direct proportion to the number of speci- taxa. Validation involves typification, specifically mens and publications examined. Comparison of the designation of kind or status of type, and such the original description with data from one or more matters as rank, priority, synonymy and homonymy, specimens rarely can be made without uncovering authorship, and orthography. Up to a point, the at least minor discrepancies that must be reconciled. records can be validated by any botanist or tech- As types from additional herbaria are registered it nical person skilled in the strict application of the often becomes necessary to reexamine the original proLisions of the ZCBN, because many of the prob- publication and secondary references again and lems are purely technical or legal. Indeed, experi- again to resolve new discrepancies, and such dis- ence with the Type Register has proved that a crepancies frequently multiply faster than they can trained technical editor frequently makes decisions be resolved as publication after publication is con- more consistent with the ZCBN than the specialist, sulted. Gazetteers, atlases, biographies and bio- at least insofar as the objectives of the Register are graphical dictionaries, personal fieldnotes and concerned. Such technical validation, while it letters, and even new correspondence with current greatly increases the reliability of the data, never- specialists, in addition to the obvious taxonomic theless is without the force of authority that can treatises and reference works, may be employed be gained only through the sanction of the taxo- eventually in the course of trying to verify the data nomic authorities themselves. As in all taxonomic of an entry in the Register. research, many of the questions that arise regarding Apart from the facts themselves is the matter of tjpification have no absolute answer but require interpretation and judgment. The compilation of good judgment by an experienced specialist on the any highly condensed, formatted, and standardized basis of all available evidence, and no amount of file of data such as the Type Register is bound to technical expertise could suffice. This type of au- involve much interpretation of fact and judgment thoritative kalidation is needed in the long run if of what to include and what to exclude. The com- the Type Register is ever to take its place as an puter imposes the additional problem of judging indispensable and thoroughly accepted tool of how best to format and standardize the data for plant systematists, and it is hoped that the special- search and retrieval. As new data are provided ists will cooperate in validating the information or brought to light, there is a constant need to re- in the Register as it becomes available group by evaluate prior interpretations and judgments, and group in preliminary form. In the short run, how- this in turn may require reexamination of previ- ever, the most that can be achieved is some degree ously consulted literature and specimens. The prob- of technical validation. The important point to stress here is the dynamic state of the file which lem is one not only of accuracy and completeness can be updated at any time to accord with current rvithin a given record but also of consistency among knowledge and understanding. records. How the geographic information is stand- Like verification, validation is a continuous proc- ardized in the record for Taxon A, for example, has ess that never really ends, because the light of new a direct bearing on how the. geographic informa- information often requires important reevaluations tion is standardized for Taxon B, and decisions and appropriate changing of the computer file. At made for the first case without knowledge of special the same time, a basic threshold can be achieved. problems to be faced in the second case may have A record is considered verified at least on an initial to be reevaluated and changed when the two rec- basis once the original description and all registered ords are considered together. In short, there is no specimens have been seen in person by someone n prion' way to set standards for all time. compiling and editing data expressly for the T ALVBER 12 9

Register. Likewise a record is considered validated and onl) when professional botanists or other spe- at least initialiy once the designation of types has cialists skilled in the application of the ZCBN per- been worked out in accordance with the lCBN ex- form this function is it possible to perform the pressly within the framework of Type Register k~lidationfunction at the same time. specifications and format, These thresholds must be The present computer file of some 13,000 speci- attained before the second and third stages of de- men entries, constituting the entire Type Register, velopment can be said to have been achieved on a 15 a registry ot largely unverified and unvalidated minimal basis, Authoritative validation as de- records of apparent or piesiimptiue types, and for scribed above, on the other hand, will require the the next several years, at least, effort will continue input of many specialists in the years to come and to be concentrated on the rapid compilation and is a very long-term proposition. input of similar preliminary data from many other The three stages of development may be sum- taxonomic groups and institutions, starting with marized as follows: the Smithsonian’s own type collection. The strategy Stage 1. Registration.-Creation of the initial is to register the greatest number of taxa and speci- file, ivhich involves basic standardization of cita- mens in the shortest possible time so as to achieve tion and geographical fields. quickly a critical mass of data for producing cata- Stage 2. Verification.-Editing file against pri- logs and answering queries. Clearly the usefulness mary sources in the light of the accumulated data. of such a data base will be directly proportional Stage 3. Validation.--Shaping the Register as an to its taxonomic and institutional comprehensive- authoritative tool on typification, fully in accord- ness. Unless efforts to verify and validate the data ance with the ZCBN, by getting input from spe- are kept to an essential minimum as new records cialists and by incorporating information on lec- are being processed, there is little chance that a totypification and neotypification, as well as other comprehensive data base can be created in the critical annotations. foreseeable future. The manpower and resources In practice, registration, verification, and valida- simply are not available at present for the massive tion certainly are not sharply delimited phases and searches in the herbarium and library that would often merge into each other as a single process. be required to bring every new record to the Stage Once the original description and specimens are in 2 or Stage 3 level of refinement as it is being en- hand one attempts to accomplish as much of the tered into the file. To a large extent, therefore, entire three-stage process as possible. Verification the Stage 1 Type Register will have to be verified and validation, in particular, tend to overlap; it is and validated through use, through feedback from in fact impossible to accomplish the one without the specialists Lvho discover its shortcomings in the to a degree accomplishing the other. From the course of their research. point of view of the daily operation, however, reg- Under no circumstances is the Type Register istration, verification, and validation represent dis- being used or is it intended to be used as a place to tinct working stages in the creation of the computer designate lectotypes and neotypes and thereby to data base, involving different procedures and per- set nomenclatural precedents. If the Register is ever sonnel. Editorially, each stage results in a more to be used in this manner, which as indicated ear- refined, reliable, and authoritative data base. In lier may prove desirable eventually, the taxonomic the first stage, the data can be compiled and regis- fraternity will have to make a conscious decision to tered entirely from secondary sources, if necessary, do so. although this is not recommended, but neither veri- When the scope of the task is considered, it is fication nor validation can be accomplished without not surprising that no one has attempted to compile consulting the original sources. Regarding the op- a union catalog of type specimens before now. The erational distinction between verification and vali- task can be cut down to size, however, because rela- dation, it should be realized that a technical person tively few of the world’s public herbaria are large may be quite competent to verify the accuracy of enough to have a significant concentration of type the data but not to validate the type designations specimens. Only about a score of the world’s her- even with the original sources in hand. Ordinarily, baria, for example, contain over two million speci- technical editors are responsible for verification, mens each, and a published index, including 10 S!vlITHSONIAK CONTRIBUTIONS TO BOTANY literature citations, to any one of these collections should be reversed, proceeding from the taxon to would be enormously useful in itself. Each new the specimen. No doubt this publication will stim- institution to be added to such a base would en- ulate debate on these alternatives; meanwhile, it hance the catalog greatly and move it one step should be made clear that primarily the first ap- closer to the goal of a worldwide union registry. proach is being taken. The U. S. National Herbarium is one of the \Vitlz one exception, only the original names of score of major herbaria with more than two million newly described taxa, i.e., taxa being described to specimens, and its type collection of 65,000 or more science for the first time, are included. The one specimens certainly constitutes a significant initial exception is a wholly new name for a previously data base. Furthermore, not only is this type col- described taxon necessitated because all other possi- lection separate from the main herbarium and ble names and combinations would violate the easily accessible, but it also has an associated file international rules. New combinations involving of cards on which are recorded pertinent data from previously published epithets are excluded rigor- the original publication (see “Source of Data”). ously insofar as they are Enown to be combina- Without this large, ready-made card file and with- tions; in such cases, only the basionym is entered out computer technology, which permits the crea- into the Register. In one sense, therefore, the Type tion of a union register on a much more flexible Register is a basionym file. This approach has been and dynamic basis than would otherwise be pos- taken because it is the only feasible way in the sible, the Type Register doubtlessly would never foreseeable future to create a stable file with fixed have been conceived or started. The Botanical points of reference. Eventually, viewed in the long- Type Specimen Register is in the first instance, est terms, it will be necessary to link the Type therefore, an index (catalog, register) of the U. S. Register to a much vaster name list that shows all National Herbarium’s own type collections. The possible synonymy connections among basionyms thousands of man-hours that have gone into the and combinations and thus makes it possible to creation and maintenance of the National Her- trace the nomenclatural history of a particular spe- barium’s type collection and card file have paid off, cies, for example, from modern usage back to origi- of course, to the many who through the years have nal usage. This is far too much to expect of the used the type herbarium on the Smithsonian prem- Type Register in itself, however, and for this rea- ises. By computerizing this information the Smith- son the design of the Register allows for no syn- sonian’s Department of Botany now makes it onymy except for orthographic variants. If a possible for taxonomists at large to benefit from taxon originally was published under a generic or thl:, accumulated data and enormous manpower specific name with a spelling that later was cor- in iestment. rected, then the original spelling is indicated in a special field, while the accepted spelling is shown in the main taxon field; for example, many species Scope of Register have been published in the genus Penstemon under The Register is designed to handle taxa typified the spelling “Pentstemon,” and this spelling is in- by specimens, namely, taxa of the rank of species dicated in the orthographic synonym field, as neces- or below, and it encompasses all infraspecific taxo- sary. IVithout this approach, the same genus would nomic levels recognized by the ZCBN. In the fu- alphabetize in different parts of the file (e.g., ture, modifications in design may be desirable if Aplopnppus vs. Haplopappus). not essential to accommodate cases in which the Only validly published names are included, but type is not a specimen but a description or a figure. the names need not be legitimate, as defined by the For the present, however, the object is to register ZCBN (see also hfcVaugh et al., 1968). specimens, and for this reason data collection usu- Taxonomically and geographically, the Register ally begins with the specimens and proceeds to the is limited only by the availability of data and op- literature rather than the other way around. There erational resources. The present machine file in- are good reasons for arguing on the one hand that cludes only vascular plants and primarily flowering registration should proceed from the specimen to plants, but it could be expanded at any time to the taxon and on the other hand that the process include cryptogamic groups if the data and the NUMBER 12 resources to input the data, especially personnel, a type specimen?” not “Where can I find a type were available. The geographic scope already is fragment?” The latter question will interest him worldwide because the initial source of most rec- only if all efforts to see a type specimen fail or ords, the U.S. National Herbarium’s type collec- prove impractical. For these reasons, type frag- tion, is worldwide in scope although particularly ments have been registered sparingly in the present strong in New World areas. The input is further file and only when the circumstances seem to war- biased geographically at present by the fact that rant doing so. the other herbaria which have cooperated thus far Lectotypes and neotypes present a special prob- on the Register also are North American institu- lem that cannot be handled properly with the cur- tions with principally New IVorld collections (ex- rent system design. The system allows for only a cept Arnold Arboretum). single bibliographic citation, namely, the citation Inclusion of type photographs has been sug- of the original publication where the taxon was first gested several times, but so far this has not been described and the name proposed. In cases of lec- done because they present special problems requir- totypification or neotypification, however, it is ing careful study before the system can be modified necessary to cite also the second, later reference to accommodate them. Whereas type specimens are where the lectotype or neotype was designated. The unique and, even in the case of isotypes and syn- problem has been largely ignored in this initial types, are distributed to a relatively limited set of phase because lectotypification and neotypification herbaria among the total, type photographs are not cannot be documented properly without the direct unique, and in theory every herbarium can have a participation of specialists. Identification of lecto- photograph of any type. The Register soon could types and neotypes is part of the Stage 3 validation be overloaded with references to photographs, and process described earlier, and by the time this level no purpose would be served. While there is a clear of documentation is possible the system will be need, especially on the part of floristic workers who modified to include a separate file, linked to the may be able to satisfy their requirements with pho- basic file, for recording lectotypes, neotypes, and tographs and thus avoid a massive borrowing of other pertinent taxonomic or nomenclatural anno- type specimens, for a central index of negatives on tations. Such a file for “remarks” will provide a deposit at major centers from which type photo- way of recording the names of authorities who have graphs could then be purchased, this problem calls validated the data. for separate attention. A word is necessary about the relationship of the To an extent, the same reasoning applies to type Type Register project to the Index Nominum fragments because many institutions potentially can Genericoyum (ZA‘G) project (Cowan, 1970). The have fragments of the same specimen. A type frag object of ZArG is authoritative typification of all ment has no standing in the ZCBN unless it can generic plant names. Thus it deals with genera, be interpreted as a formal type of some kind (e.g., not species, except for type species, and it is not isotype), and most fragments cannot be dignified concerned with type specimens or collection data by such interpretation. In the modern era when of any kind. Emphasis is placed on achieving at travel and communication are easy, making the re- once, before input, the level of validation that the motest corners of the earth accessible, the informa- Type Register is expected to achieve only in the tional value of the type fragment in one’s own long run. The Register, which is not concerned herbarium has diminished greatly because the type with the typification of genera, and ING are com- specimen itself can be borrowed or examined by plementary, therefore, and do not duplicate each personal visit. Thus only in the case of types that other in any way (see also p. 16). have been destroyed or of types that for political IVith respect to Flora North America, the closest or other reasons are still inaccessible can importance links are maintained between it and the Type Reg be attached to a register of information on the ister project on the one hand and ZhrG on the whereabouts of type fragments. In other words, the other hand, to ensure that the work of each project taxonomist wants to know, “Where can I find will complement rather than duplicate the others. 12 SMITHSONIAN CONTRIBUTIONS TO BOTANY

Procedures and Standards and infraspecific taxa, and the other approximately 10,000 are types of cryptogamic taxa-ferns, mosses, SOURCEOF DATA and lichens. Other institutions can contribute to the Type The principal source of data at this stage is the Register in any of a number of ways, as explained permanent card file associated with the type col- in the next section. Basically, there are two ways: lection of the U.S. National Herbarium (US). The (1) annotation of a printou, listing records already card records are converted into machine-readable registered in the machine file, and (2) submission form genus by genus in alphabetical order. During of completed data forms or some equivalent pro- the pilot phase cards were pulled from the file by cedure for new records not presently registered in family (e.g., Scrophulariaceae), but this approach the machine file. These are complementary pro- is impractical for the file as a whole, which is ar- cedures which must both be used. Institutions with ranged alphabetically by genus. While that ap- ready-made card files like the Smithsonian’s are in proach was being taken, the cards were being the best position to contribute in a significant way compared with the specimens in the type collec- quickly, and their contributions will spare the tion, which are arranged systematically (modified smaller herbaria from repeating costly bibliographic Englerian sequence), prior to input. Now that an research that already has been done somewhere alphabetical rather than systematic approach to the else. The cumulative Register provides a basis for file is being taken the specimen-comparison step checking rapidly for isotypes and other “duplicate” is being postponed until the whole file is in the type material, leaving bibliographic research to be computer and can be sorted systematically by performed only for those cases where new taxa are family. to be added to the Register. In other words, to Curators of the U.S. National Herbarium have conserve effort maximum advantage should be followed the practice of segregating type specimens taken of the existing file in the process of adding from the general collection since the early part of new data, especially bibliographic data, and of the present century. The practice was first estab- course the larger the machine file becomes the lished about 1918 by then-curator Paul C. Standley. greater can be the economy of scholarship on the At the same time an associated card file was started part of newly collaborating herbaria. to supplement the specimen data with information AZonographs and the personal manuscripts or from the literature. The file includes a card for files of monographers are obvious sources of au- every taxon (species, subspecies, variety, form) rep- thoritative data for the Register and have been resented in the type collection, and generally the used in a few instances, although there are some responsible curator has had the original publica- distinct disadvantages in using the monograph as tion in hand while preparing the card and the the starting point (see p. 7). Future monographers standard folder for filing the type specimen (s). should register data routinely for type specimens of Each card includes the original taxonomic name new taxa prior to, or simultaneously with, publica- (basionym in cases of later transfer), author, origi- tion. Likewise, it is hoped that graduate students nal reference, basic collecting data, and designation in plant taxonomy will be advised to submit data of kind of type. To re-create this file today from on type specimens examined by them in the course the specimens and the literature tvould require at of their research. least 10 and more likely 20-30 professional man- For every specimen registered in the file, a code years, and there is no reason to suppose that the is appended at the end of the record which indi- file could be re-created with any higher professional cates the source of the data according to a broad standards or greater degree of accuracy on the av- classification of source categories, summarized later erage than the first time. In short, it is scientifically under “Data Source Code.” sound as well as eminently practical to create the preliminary edition of the Type Register from the INSTRUCTIONSFOR CONTRIBUTORS Smithsonian file as it stands. Of the 65,000 specimens in the US type collec- Any herbarium interested in contributing to the tion, about 55,000 are types of phanerogamic species Register is advised to consult with the staff at the NUMBER 12 13

FIGUREP.-Standard Data Collection Form of Botanical Type Specimen Register. 14 SMITHSONIAN CONTRIBUTIONS TO BOTANY

Smithsonian well in advance of initiating a project mitting duplicate information for taxa already re- so that a compatible way of compiling data can be corded. devised before the first record is collected. At pres- The procedural details will differ from institu- ent the Type Register staff has its hands full with tion to institution, but these general guidelines the internal Smithsonian file but will do its best should be followed, unless other special arrange- to cooperate with other potential contributors to ments have been made beforehand: set up procedures for submitting data. The most (1) All new records-taxa new to the file-should efficient way of collaborating under present cir- be submitted on the standard data collection form cumstances is to submit the records on the standard (Figure 1) and according to the technical data spec- data form (Figure 2) to the Smithsonian for proc- ifications used by the Smithsonian. Be sure that essing, but with adequate planning it will be possi- the original name and not a later combination is ble for another institution to convert its own data being used. into machine-readable form in-house before submit- (2) All additions to existing records-supple- ting the data to the Smithsonian and thereby to save mentary publication or collection data, and collec- time and effort in the overall process, at least where tions or specimens new to the file-should be large herbaria are concerned. Complete records, submitted as annotations to the appropriate records i.e., with all essential data present, submitted by in a copy of the working catalog; or, if they are other institutions are placed in the queue and proc- submitted on standard data forms or in a separate essed in due course as time permits. Incomplete typewritten listing, each addition should be prop- records, particularly those without proper biblio- erly referenced to the record in the file to which graphic citations, are set aside in an inactive file it belongs. Care should be taken to distinguish where, for lack of staff to complete them, they may between specimens that belong to a collection al- remain indefinitely. Potential contributors should ready registered in the file and specimens that keep in mind, therefore, that any effort that falls introduce a new collection but to an already regis- short of providing complete records risks becoming tered taxon. Both constitute additions but on dif- a wasted effort. ferent levels. The latter situation arises frequently A collaborating institution should always work when syntypes are involved, but slight discrepan- from the current catalog in the Type Register, if cies in collection data can easily be overlooked by one exists, of the genus or other taxonomic group the compiler, leading him to the conclusion that concerned. The Carex: Catalog, for example, pro- the former situation obtains. If the collection data vides a checklist of taxa already registered which recorded in the Register are not identical or at least can be used as a tool by any herbarium wishing to reconcilable with the collection data of the speci- search its own collections for type specimens of this men in hand, then the discrepancy should be re- genus. Working catalogs for other registered groups solved appropriately or the specimen excluded. In can be provided at cost by the Smithsonian as they annotating the catalog, it is important to be sure are needed. The purpose of this procedure is to that the additions are clearly associated with the reduce unnecessary effort on the part of both the proper collection where two or more collections compiler and the editor. In the system, all speci- are registered. mens pertaining to a given taxon are registered un- (3) All proposed changes to existing information der a single entry, and thus there is one unit record in the file should be submitted with documentation per taxon. It is the editor’s responsibility to pre- as annotations to the appropriate records in a copy vent duplications of the same taxa in the first place of the working catalog; or, if they are submitted and in the second place to discover and delete the on standard data forms or in a separate typewritten occasional duplication that inevitably creeps into a listing, each addition should be properly referenced file of the size and complexity of the Type Register. to the record in the file to which it pertains. With- Individual compilers can do much to assist the out documentation, proposed changes raise more editor in preventing duplications, however, by questions than they answer and complicate the keeping themselves informed, through working work of the editor. catalogs, of the current status of the file, and by The most common errors, experience has shown, using procedures that minimize the chances of sub- are mistaking (1) a later combination for an ongi- SUhIBER 12 15 nal name, (2) the type specimen of a variety or be bothered with all of the technical details of the form for the type of the species itself, and (3) a system. For purposes of contributing records, it is syntype (or isosyntype) for an isotype. The second essential to know only the main conventions and of these mistakes often turns out to be the ex- standards that govern the content and format of planation for the situation in which the specimen the data fields. These basic standards are explained appears to have been collected after the species was below by field, and contributors are strongly urged described, i.e., the collection date is later than the to follow them closely so that the work of the publication date. An undetected name transfer editor will be simplified. The rules of standardiza- may lead to puzzling discrepancies or to unnoticed tion have been applied more rigorously and con- duplication. Frequently, for example, a type co:. sistentl) to the Carex Catalog than to any other lection passing under an undetected later combina- part of the Register, and potential contributors are tion becomes the basis for introducing a new taxon asked to study this Catalog carefully for specific to the file, while at the same time the collection examples of how standards have been applied to already is properly registered under its basionym. govern content, form, and style. Implicit in this A final note should be added about the use of the Catalog are the answers to many specific questions Carex Catalog or any similar working catalog in about standardization which cannot be answered herbaria where type specimens have never been here. It should be kept in mind, however, that the identified and segregated into separate folders or a typical edit format is different from the format of separate collection. By means of the collector in- this published Catalog. In the typical format, the dex, it is possible for curators who wish to begin information is not strung together (concatenated) segregating types to use such a catalog as a means in paragraph form, but each field is labeled and of identifying type specimens within their herbaria printed by itself with room for annotations. Fur- which belong to collections recorded in the Type thermore, certain fields of data (e.g., source code) Register. have not been printed out in the Carex Catalog. The reference works used most frequently in the course of compiling and editing data for the RECORDFORMAT AND CONTENT Type Register are cited in the bibliography. Some The data content of a unit record in the Regis- of these works have been adopted for editorial pur- ter was established largely according to the conven- poses as the standard references and authorities for tions long used for the US card file. With the US verifying and standardizing new data during initial card format as a standard, the data form shown input. An “authority” (authority file) is an index, in Figure 2 was devised for use by contributors at thesaurus, or dictionary of terms, names, or titles other institutions. This form shows what fields of tvhich is used to standardize some category of data, data should be included, distinguishing between eg., B-P-H (Lawrence et al., 1968) for titles of essential and nonessential fields and indicating cer- botanical periodicals. Published standards have tain of the basic standards. The fields tagged as been adopted as authorities whenever possible, but “essential” constitute the minimum number of data in some instances it has been necessary to begin elements which the processing system is designed creating authority files expressly for use in the Type either to require or expect. From the botanical- Register project. A “standard reference” is an content point of view, however, none of the fields authoritative and reliable secondary source that should be regarded as optional. Every effort should provides the editor with a practical means of be made by the contributor to provide data for all quickly verifying some category or categories of fields. Contributors can obtain blank forms at cost incoming data, e.g., a name index such as Index from the Smithsonian or use facsimiles. Kerciensiy or LVillis’ Dictionary of Flowering Plants Before input to the system, all records are edited nnd Fems (7th edition, revised by Airy Shaw, 1966). to conform with the field-by-field technical specifica- Verification in this editorial sense of double- tions that have been established to standardize con- ( herking in secondary sources is not to be confused tent and format in the system. A copy of these Tvith verification in the primary scholarly sense dis- specifications can be made available to collaborators cussed earlier in connection with Stage 2 develop- on request, but most contributors will not want to ment of the Type Register (p. 7). 16 SXITHSOSIAN CONTRIBUTIONS TO BOTANY

The Internotional Code of Botanical Nomencla- irregular names with their adopted regular equiva- tu)e is, of course, the final authority on all matters lents are: Compositae/ASTERACEAE, Cruciferae/ of typification and nomenclature. (The 1966 edi- BRASSICACEAE, Graminae/POACEAE, Guttife- tion, prepared under the chief editorship of Lan- rae/CLUSIACEXE, Labiatae/LAMIACEAE, Legu- jouw and Stafleu, has been used thus far, but the minosae/FABACEAE, Palmae/ARECACEAE, Um- rietver 1972 edition of Stafleu et al. is now availa- belliferae/APIACEAE. The use of regular family ble.) Of the essential desk-top references, the one names conforms with practice in the FNA program. most frequently consulted, perhaps, is the seventh IVillis’ Dictionary is used in conjunction with the edition of JVillis’ Dictionary. Also invaluable as Syi;obus to determine the family to which a genus general reference works are Stearn’s Botanical Latin belongs. (1966) and Stafleu’s Taxonomic Literature (1967). GEXS AND GENUSSYsoNuhr.-Index Nominum Though hardly desk-top references, Index Kewensis Gcne,’icorum, insofar as it is completed, is the final and the G?ay He)bai ium Index are indispensable, authority for generic names, to determine their ac- and the Type Register editorial staff is fortunate cepted spelling and whether they are validly pub- in haking available to it an integrated version of lished. IVillis’ Dictionary, which in any case is an the lormer and both the card and book forms of indispensable authoritative handbook on these the latter. Insofar as possible, the same procedures matters, is consulted for genera not yet covered by and standards are being applied in both the Type I*YG. Il’henever the data are being compiled di- Register project ant1 the Flora North America pro- rectiy from the primary sources, the generic name gram, so that the data bases will be compatible. is entered on the data form exactly as it was spelled .luthority files developed especially for the one in the original description of the particular species project are being used also for the other as ap- or other taxon in queution. If for some reason this propriate. spelling is a variant of the currently accepted or- The main editorial procedures that have been thography of the name, then both spellings are en- adopted to verify and standardize the ingoing data tered into the machine file, the accepted spelling are summarized below field by field. Collaborators in the “Genus” field and the orthographic variant can do much to increase the reliability and stand- in the “Genus Synonym” field. In no case is more ardization of their own data by using these same than one spelling permitted in the “Genus” field procedures in the process of compilation. The for the species and infraspecific taxa of any par- editorial burden is eased greatly when the editor ticular genus of plants. knows in advance that the essential standards have SPECIES.-Attempt is made to record the specific been upheld consistently by the contributor, epithet exactly as it was spelled originally, except where a minor change is required by the provisions Level 1-Taxon Data of the ICBhTgoverning orthography. (A two-word epithet, for example, is hyphenated to form a single The following fields of data are recorded only word.) To verify the spelling given on the data once each for every species or infraspecific taxon form, the editor relies upon the Gray Heybarium registered in the machine file because these data Index and/or Index Kewensis insofar as possible, are unique for each taxon. Furthermore, the taxon is the unit record, and every taxon is entered into because to check the primary publications in all the file only once. cases is impractical at this stage. These two indices, FA%fItu.-The latest edition of Engler’s Syllabus which overlap considerably in coverage, frequently der Pflanzenfonizlien (vol. 1 : Melchior and Werder- provide a check on each other. Unless the taxon in mann, 1954; vol. 2: Melchior, 1964) is the authority question is by definition outside the limits of one for the system of families with the exception that of the two indices, the second index is checked the accepted family name with a regular ending routinely whenever the first does not confirm the is used in all cases, including the eight cases where spelling given on the data form. TVith respect to the Syllabws, following the traditional practice sanc- species regarded as being of hybrid origin, the tioned by the ICBA‘ (see list of h’omina Familia- standard practice of placing an “X” followed by a rum Coriservandn), uses irregular names. These blank space before the epithet is followed. SUMBER 12 17

INFKASPECIFICTAxos.--An infraspecific taxon of countered in dealing with author names, but the any rank recognized by the ICBN can be accom- details are beyond the scope of this introduction. modated in the Type Register by entering the in- Examples of specific solutions can be found in the fraspecific epithet and the appropriate rank Carex Catalog. designator in this field, e.g., VAR GRACILIS or TiTLE.-The title of the periodical, monograph, SFXI CRASSA. Infraspecific names are entered in or book in which the name of the taxon in question the Register in the form of trinomial combinations ivas first validly published is recorded in this field. Ivith the appropriate rank designator, because this The title of the article in the periodical or of the form is adequate for nomenclatural purposes; and chapter in the monograph or book is never included quatlrinomials create problems in the system. The or given in lieu of the title of the periodical or Gray Herbarium Index is used to verify infraspe- work. In cases where a name was introduced into cific epithets as to rank and spelling, but it covers the literature before it was validly published, refer- onl) infraspecific taxa of the New World published ence is made only to the place of valid publication. during the past 100 years. For this reason, many A name published first without description (nomen of the infraspecific names must go unverified at this nzidzim), for example, often is later published valid- stage when extensive literature research is imprac- ly with description. Nomina nuda are not included tical. Following is a list of the infraspecific ranks in the Type Register. All titles are abbreviated con- and their standard abbreviations used in the Regis- sistently. B-P-H is the authority for abbreviating ter: titles of periodicals. For titles of monographs and subspecies SSP books, the Type Register project is developing its variety VAR own authority file, using the principles of abbrevia- subvariety SVR tion set forth in B-P-H, and, insofar as possible, form FOR taking advantage of the title abbreviation file de- subform SFM veloped by the ZNG project. An author’s or editor’s nothomorph NM. iiame is not included in this field unless it actually .4L1THoK.-This field carries the full last name is part of the title; otherwise, it is included in the and all initials of each author of the binomial or previous field as the publishing author or editor. trinomial name under which type specimens are being registered. Whether single or multiple au- Level 2-Collection Data thors, the last name is always placed before the initials. When the use of initials only is certain to Because there may be multiple type collections lead to confusion, the full first name also is in- (e.g., syntypes) for any taxon, the collection data cluded, e.g., MACOUN, JOHN vs. MACOUN, fields may repeat as a set any number of times. The JAAIES hi., not hIACOUN, J. vs. hIACOUh’, J. Xi. folloiving set of fields is recorded for each collection, The publishing author or the author of the work, insofar as the data exist and are available. if different from the author of the name, is always COLI.EcToR(S).-The names of all collectors of the included in this field and separated from the author type collection being registered are recorded in this of the name by “IN” or “EX” in accordance with field exactly as author names are formatted in the the rules and recommendations of the ZCBN. No “Author (s)” field. The name of a collecting expedi- authority exists for author names, but an author authority file has been started for the Type Register tion may be recorded here when individual collec- on the basis of the Carex Catalog. Further, the tors cannot be determined. The Type Register FNA Author File is well underway, and it is project is developing its own authority file for col- planned that the two be compatible and that ulti- lectors, but meanwhile the following biographical mately they be merged. Meanwhile, the standard indices, in addition to those already mentioned references are Barnhart’s Biographical Notes Upon under the “Author (s)” field, are being used as Botanists (1965) and Stafleu’s Taxonomic Litera- standard references: “Zndex Herbariorum, Part 11: ture (196’i), but all available biographical refer- Collectors” (Lanjouw and Stafleu, 1954, 1957; ences are consulted as necessary. (See also under Chaudhri et al., 1972-completed for letters A to “Collector[s].” Many specific problems are en- L); “Index to Principal Collections Represented in SMITHSOSIAN CONTRIBUTIOSS TO BOTANY the U. S. National Herbarium” (compiled by U. S. the smallest units, and unimportant words are omit- National Herbarium staff for internal use, 1965). ted; latitude and longitude, if given, are placed last COLLECTIONNuMBER.-ordinarily this will be the in this field; and ecological terms, except where re- collector’s own number, but when there is no trust- quired to clarify the geographic location, are omit- worthy means of determining his number, or if he ted. Because locality data from the specimen and had none, a serial collecting number assigned by an the original description often are merged into a institution or expedition may be recorded instead. single telegraphic statement, the reference given in The field is regarded pragmatically as the place for this fourth field cannot be assumed to be a direct a number, any number, that has been associated quotation. Every effort is made, however, to stay with the collection and which, when combined with close to the words of the original collector and/or the name (s) entered in the “ (Collector (s)” field, author of the description, and substantive addi- normally will form a unique reference to the col- tions or interpolations by the compiler or editor are lection. If a distinction can be made between the indicated appropriately. It is important to the number of the collector and the number of his etli tor, therefore, that compilers set apart clearly expedition or institution, then the name of the their own comments from the original information. expedition or institution assigning the serial num- Quotation marks are used only when the context ber should be placed in the “Collector (s)” field requires that the exact original words be identified, unless one or more collectors’ names already have as, for example, when some part of the locality been entered there; otherwise, this name should be statement is so archaic, confusing, or general as to prefixed to the collection number to make clear appear to contradict the rest of the geographic in- that the number is not the collector’s own. When formation. there is absolute evidence that the collection never The three, higher level geographic data fields are has been numbered in any series, then the abbre- used to place the locality in its proper geopolitical viation S.N. (sine numei-o), meaning “without hierarchy. IVhereas the vocabulary used in the number,” should be entered in this field. If on the “Locality” field is standardized for sorting purposes other hand the number is merely unknown or there but not controlled, the vocabulary used in these is doubt about the existence of a number, then three fields is controlled as well as standardized for dashes (- - -) should be entered in the field. purposes of search and retrieval. Insofar as possible, COLLECTIONDATE(S).-collection date is recorded current official political units are used in all three just as accurately as it is known, and if necessary fields so as to avoid overlapping and inconsistent two dates or a range of dates are given. All dates, terminology. As the term “geopolitical” implies, whether single or in ranges, take the form: 28 Sep concessions to age-old geographic designations, as in 1928. On the data form, dashes should be entered the case of certain islands, are made in a few in- to indicate that the collection date is unknown, and stances, and “country” is not always an independent the abbreviation S.D. (sine dato) should be used to political unit in the strictest or most modern sense. indicate that the collection is known to be without Such changes are made only within the structure of a collection date. In using the latter designation, the controlled vocabulary, however, as explained the compiler should be absolutely certain that the below. Island names present a particular problem, collection is undatable; otherwise, he should use because often they have long been used in the bio- da4ies (- - -). geographic literature but do not fit into a consistent GEOGRAPHICDATA FIELDS.-Four geographic fields geopolitical hierarchy (e.g., Borneo, Madagascar). are used to pinpoint hierarchically the collecting Various stratagems, mostly involving comments in locality: (1) country; (2) state, province, depart- ment, or equivalent; (3) county or equivalent; (4) the “Locality” field, have been devised to cope with locality. The specific place is recorded in the fourth the problem of identifying well-known biogeo- or lowest field more or less in the terms of local graphic areas within the file structure of the Type reference given by the collector himself and should Kegister. If the name used in any of the three include a town, post office, or other place name higher level fields is not obviously equivalent to that can be found in an atlas. The locality terms the name used originally by the collector, then his in the fourth field are ordered from the largest to original designation is included parenthetically NUMBER 12 19

with appropriate annotation at the end of the HERBARIUMAABBREvIATION.-The standard inter- “Locality” field. national abbreviations established in the fifth edi- A11 available atlases and gazetteers are used as tion of “Index Heibariorum, Part I: The Herbaria standard references, the most valuable being The of the SVorld” (Lanjouw and Stafleu, 1964) are Columbia Lippincott Gazetteer of the World (Selt- used to designate the herbaria. zer, 1962) and the desk-top Webster’s New Geo- HERBARIUM SHFET NuniBER.-Many herbaria graphical Dictionary (1972). The latter, to the stamp a serial number on every sheet to which an extent that it covers the geographic units and prob- herbarium specimen is attached, and that number is lems encountered, has taken on the force virtually entered into this field. The field may be left blank of an authority for geographic standardization. The when the sheet in question lacks a serial number. authority adopted for the system of classification Because a sheet number represents the single most and names of the world’s countries and equivalent effective and reliable means of uniquely identifying political units is the National Bureau of Standards’ n specimen, any collaborating herbarium which Federal Information Processing Standards Publica- presently does not number its sheets is strongly tion (FIPS Pub) 10: Countries, Dependencies and urged to number the sheets of type specimens as the Areas of Special Sovereignty (1970). With slight data are compiled for the Type Register. From the modifications for the purposes of Type Register, standpoint of the Register, the serial number is a this publication is used to control the vocabulary completely arbitrary datum and need not belong to used in the first or “Country” field. Any new name any general numbering system within the collabo- must be fitted into this system before it can be rating institution provided that it is part of a used. unique series. SVhenever there is any choice on the From the data-processing point of view, the pur- matter, a totally numerical series, not a mixed pose of the geographic information is to make alphabetical/numerical (alphanumeric) series, search and retrieval possible at least by country and should be used, to facilitate proper numerical sort- state or province. Thus if the first two fields are ing b) machine. left blank or if the names are not carefully control- Kmn OF TYPE.-This small field represents the led and standardized the retrieval aim is clearly purpose of the Type Register and is certain to evoke thwarted. Collaborators should make every effort more discussion and controversy than any other to provide data in all four geographic fields, but the data field in the unit record. For this reason it is most important ones are the first, second, and vital that every user of the Carex Catalog or any fourth. other part of the Register understand from the outset the limitations of the data recorded in the “Kind of Type” field. Level 3-Specimen Data .is emphasized repeatedly, the initial aim of the Because there may be multiple type specimens Type Register project is to record the facts just as (e.g., isotypes) in any type collection, the specimen they exist in the presently available sources so as to data fields may repeat as a set any number of times. put before the taxonomic user community the Ordinarily, an institution will be represented by a greatest amount of information in the shortest pos- single type specimen under a given collection, but sible time, leaving until later stages the objective of there is no limit to the number of specimens that methodical, authoritative verification and valida- may be registered per collection as long as each tion. Once comparative data on type specimens are specimen is uniquely identified. In practice, this available group by taxonomic group on a large means that two or more specimens will be cited for scale, the specialists themselves, who alone are truly the same institution onty if they have different qualified to render authoritative decisions on mat- herbarium sheet numbers or represent different ters of nomenclature and typification, can help kinds of types. The data fields in the specimen enormously to refine the Type Register data base citation are: Herbarium Abbreviation/Herbarium through feedback arising from actual use of the Sheet Number/Kind of Type/ Data Source Code. file. The consequence of this register-now-validate- The typical specimen citation takes this form: US later approach is seen most often in the imprecise 1727345 HOLOTYPE CF. if not incorrect terms by which the different kinds 20 SMITHSONIAN COSTRIBUTIOSS TO BOTANY of types are designated. Because the nomenclatural for purposes of lectotypification. Fragments of holo- rules have changed through the years and the well- types, isotypes, or syntypes may be included at the developed modern terminology is of relatively re- discretion of the editor (see under “Scope of Reg- cent origin, it is natural that types have not been ister”). designated according to any consistent standards When a holotype has not been designated, as in through the years. all the older literature, one usually is faced with a When a type specimen is first registered, its typi- “syntype situation,” which often is difficult to re- fication is designated by whatever term is indicated solve precisely on the basis of the ZCBN’s terminol- in the data source, which usually is a secondary ogy. A syntype, according to the ZCBN (Article 7, source (card file, specimen file, monograph), unless Note 3), is “any one of two or more specimens cited there is firm evidence to indicate otherwise. Thus by the author when no holotype was designated, or any type designation, whether legal or illegal by any one of two or more specimens simultaneously present nomenclatural rules, may appear in the designated as types,” and an isosyntype is a dupli- Type Register. cate of a syntype (see “Guide for the Determination At the United States National Herbarium, it was of Tjpes” in the ZCBN). “Duplicate” in this con- customary for many years to designate two basic text is defined as “part of a single gathering made categories in the segregated collection of type by a collector at one time.” In other words, a “dup- specimens: “type” and “type collection.” Usually, licate” is one of two or more specimens constituting “type” has meant what would now be called “holo- a single “collection,” as this term ordinarily is used type,” while “type collection” has embraced syn- by plant taxonomists and is being used in the con- types, isotypes, and even paratypes according to text of the Type Register. present terminology. “Types” often prove not to be The distinction betxveen syntype and isosyntype holotypes, however, and it would be very wrong to hinges on such relatively subjective criteria as draw simple equations between the older and newer “specimen citation” and whether or not the original terms. The collection of type specimens at the New author had the specimens in hand (cf. definitions York Botanical Garden provides another example of McVaugh et al., 1968), which are matters for of the problems with archaic terminology. Here the specialists to determine. The older literature, where categories “type” and “cotype” were used for many the problem of syntypes arises, is well known to be years, and now “type” often but certainly not always less than precise in the manner of citing specimens. translates to “holotype,” while “cotype” may desig- The editorial staff of the Type Register must re- nate any of the kinds of types masquerading under strict its interpretations to the letter of the ZCBA‘, “type collection” at the US. and for the most part such fine distinctions as be- The authority for designation of kind of type tween syntype and isosyntype necessarily are defer- is the ICBN. An auxiliary, highly authoritative red for the proper specialists to make at a later standard reference is “An Annotated Glossary of time. To do otherwise would be to assume the Botanical Nomenclature,” by McVaugh et al. specialist’s ro!e and responsibility and to introduce (1968). LVhenever the original specimens and liter- false precision at this stage. Accordingly, the term ature can be examined and the kind of type vali- syntype is used for both syntypes and isosyntypes dated in accordance with the rules, proper termi- except in the rare cases where the evidence for the nology is used. By this terminology, the Register isos) nt) pe designation is clear and convincing. is designed to include primarily lzolotypes, isotypes, A final point on the use of the term syntype con- syntypes, and isosyntypes. As explained earlier, the cerns the distinction between single collections and present file structure is not designed to handle lec- multiple collections. Throughout the older litera- totypes and neotypes (or isolectotypes and isoneo- ture there are numerous cases where a single col- types), although these are entered sometimes by lection has been designated as the type collection, using the fourth geographic field (“Locality”) as a either explicitly or implicitly by virtue of being the remarks field for the second bibliographic citation. only collection cited, even though a holotype has Paratypes are excluded unless other, higher order not been set apart. hiany specialists would single types cannot be located and there is reason to be- out a presumptive holotype in these cases on the lieve that the paratypes will become important later hsis of the rpecimen (s) which the original author SUMBER 12 21 is presumed to have examined firsthand, but the specimen is likely to be more reliable than a record Type Register editors cannot and should not make based solely on secondary sources. Following is the authoritative selections in such cases and must re- classification of data source codes: gard them all as syntypes. The ZCBN does not seem to provide term for distinguishing this common 0s Original publication and type Specimen examined by syntype situation from the other common syntype person compiling data for Register. situations in which two or more type collections are OP Original Publication examined by person compiling data for Register, but type specimen not seen: designated simultaneously. Because it is useful to supplementary information about the specimen, if know Lvliether one or more than one type collection any, derived from secondary source (s) . is cited, in the Type Register project the term “type TS T\pe Specimen examined by person compiling data for collection” has been given a proper meaning for Register, but original publication not seen: citation the purpose of distinguishing these two syntype and other publication data, if any, derived from secondary source (s) , including standard indices (In- situations. Type collection, in this proper sense, dex Kewensis, Gray Herbarium Index) , monographs designates a specimen from a single type collection, and revisions, annotations on specimen sheet, card while “syntype” is reserved for designating a speci- files, original dacriptions removed from context of men from any one of two or more simultaneously publication without exact citation and necessary designated type collections. prefatory matter, and the Type Register catalog itself. IMG Data derived from most recent MonoGraph of taxo- The catchall term type material is used to desig- nomic group in question without reference to any nate any specimen presumed for some reason to be other source (s) of information. a type but for which there is no basis at the time of ss Data derived entirely from Secondary Sources. data input to assign a more precise classification. CF Data transcribed directly from a card in the Card Many situations arise in the course of compiling File of the type collection of the U. S. National Herbarium without verification against the original and editing data for the Type Register in which publication or type specimen. it would be useful to have a collective term for co Data from US Card file verified by examination of designating a collection as a counterpart to Ihe Original publication. singular term given in the ZCBN for the specimen. cs Data from US Caid file verified by examination of In fact, the terms “holotype collection” and “syn- t!pe Specimen is) . CM Data from US Card file verified or supplemented by type collection” often are used informally within consulting latest Monograph of taxonomic group in the project as collective counterparts to the singular question. terms holotype/isotype and syntype/isosyntype. UIL Source of data UnKnown. DATASOURCE Com.-Source of data is indicated according to a classification of source categories, by PROCESSINGSYSTEM appending the appropriate code at the end of the specimen citation. It is impossible to document in The first step is to convert the data to machine- detail the source of every datum, and any categori- readable form, i.e., to “automate” or “capture” the zation of sources is certain to have many imperfec- data, so that they can be processed by computer. tions. The present classification is only a rough first No data conversion (automation, capture) system is approximation of the kind of documentation need- perfect, and none is capable of handling all appli- ed, but presumably it is a strike in the right direc- cations equally well (Shetler, 1972). Several meth- tion. It is drawn up largely from the point of view ods and media have been tried thus far in the Type of the central staff and their internal Smithsonian Register project in an effort to find the data con- operation, and other categories will have to be version system best suited to this data-processing added as other institutions join in the effort. Prob- application. In general, the aim is to use the system ably codes or numbers will be assigned to individual that will get the data into the computer with the contributors in the future in the manner of Index least amount of error and effort. Because the devel- A’ominzrm Genericorurn. It should be noted that opment of data conversion devices and procedures this is more than a classification of sources; up to a continues to evolve rapidly, a flexible approach has point it also is a classification of degree of verifica- been taken; the data conversion system is kept as tion/validation. Obviously, a record based on exam- independent as possible from the rest of the proces- ination of the original publication and the original sing system so that a new conversion system can be 22 ShiITHSOSIAN CONTRIBUTIONS TO BOTANY adopted at any time with minimal impact on the COMPUTERPROCESSING SYSTEM.-During the pilot overall Type Register operation. phase of the Type Register project, this system con- The following data conversion systems have been sisted of a set of specially written COBOL pro- used in the chronological order given: grams, designed to run on the Smithsonian’s PAPERTAPE SYsTEM.-Data were mechanically Honeywell 1250 machine, to create and maintain a encoded on paper tape with a tape-punching type- machine file, and to account for collaborating insti- writer. The tapes then were read by the computer tutions to which data-collection cards were sent or which converted the holes in the paper tape (i.e., from which data were received. It was not designed mechanical codes) to electronic codes on magnetic as an information retrieval system in the strict tape and thus transferred the machine-readable sense, and in terms of output the system was capa- data from the medium of capture to the medium of ble chiefly of producing catalogs, either in book or computer processing. This system was used for the card form. The pilot system proved with use to be duration of the pilot project, and several thousand highly specialized with too few capabilities and records, a third of the present file, were captured serious weaknesses in file structure such as inability with it. During the pilot phase, while paper tape to handle syntypes. It served the purpose of a pilot lvas being used, corrections to the machine file system, however, to get a file started and thereby were made by means of the standard 80-column define through experience the problems to be punch card. solved in the operational system. ON-LIKESYsTE&t.-with a typewriter terminal, After the pilot phase, the specialized COBOL data were entered via telephone directly on disk programs were abandoned, and the processing sys- storage of a remote time-sharing computer under tem was completely redesigned around the com- the on-line control of a sophisticated text-editing mercially available, IBM-supported program pack- program package. This text-editing software per- age known as the “Generalized Information Sys- mitted the terminal operator to direct the computer tem” (GIs), which runs on the larger models of in making any of a whole series of deletions, IBRl’s System/360 and System/370. This general- changes, and additions during the input process so ized software performs all the normal functions of that maximum editorial accuracy could be achieved creating, maintaining, and querying files and of in the machine-readable data base immediately, be- generating reports. It is an information retrieval fore it was output onto magnetic tape for subse- system, in the proper sense, with the full capability quent processing by the information retrieval sys- to search, select, and print answers to specific tem. queries on demand in addition to the capability for OPTICALSCANNING SYsTEM.-Data were typed on producing various types of tabulations, tallies, and standard forms with an ordinary 10-pitch IBM Se- catalogs. A COBOL preprocessing program (“pre- lectric Typewriter equipped with a head having processor”) and a COBOL concatenating program, a special optical scanning font. Completed forms by which, respectively, the data are prepared for were scanned by an optical character reader (OCR) processing by GIS and the data are joined field by which encoded the data directly on magnetic tape field into publishable paragraph output after pro- for further computer processing. cessing by GIS, form a part of the total operational ~IAGNETICTAPE SYsTE;M.-In the system currently system. GIS runs only on IBM equipment and is being used, a typewriter encodes data electronically offered by several computer service bureaus in the on a magnetic cartridge which is compatible with Washington, D.C., area with IBM machines. Type computer tape. The typewriter unit also serves as a Register processing with this system so far has been communication terminal to transmit the data cap- carried out successfully at several different service tured via telephone directly to the computer, where bureaus, and the project basically is independent of the data are transferred to disk or tape for further the computing center. processing. Processing programs can be controlled One of the most important initial tasks in de- from this same remote terminal. Of the several veloping the Type Register is to build a file of data conversion systems used, this one seems to offer sufficient size to make reliable studies concerning the best compromise of advantages and disadvan- such matters as record comprehensiveness, record tages in the context of the operation as a whole. format, field format, need for authority files, and SUMBER 12 23

FIGURE%-Two oversimplified schematic representations of the three-level record structure in the Botanical Type Specimen Register (from Meadow, 1973a,b).

report types and formats, This requires a flexible ously for correlated data. With respect to Type information processing system that allows for data Register, for example, the hierarchical feature per- to be restructured, reports to be reformatted, tallies mits subordination of two or more specimen collec- to be made, and edits and mass updates to be made tions to a single taxon or, in turn, two or more without reprogramming. From the output stand- specimens to a single collection within a taxon. The point, it must be possible to select records according three-level hierarchy of the Type Register is out- to the content of any data field, to relate records to lined in Figure 3. each other on the basis of selected fields, and to sort The second-generation processing system for the and format selected fields as desired. GIS affords Type Register by no means represents the ultimate all these capabilities. system, but it does handle the vast majority of cases The FNA program is using GIS, and it was for very well and solves the most bothersome problems this reason particularly that the decision was made encountered during the pilot phase with the special- to use GIS in the Type Register project so that ized first-generation system. New problems have these botanical data bases would remain fully cont arisen, however, and with the experience gained by patible. The rationale for using a generalized infor- using the present system it will be possible to de- mation processing system and a description of the sign a third-generation system in due course that use of GIS in the FNA program have been set forth will accommodate all of the special cases that con- in separate papers recently by Harriet R. Meadow tinue to be troublesome, e.g., the case of a lectotype (1973a, b), who is responsible for the basic design or neotype that requires a second bibliographic of the Type Register processing system. The two citation. most important features of GIS, from the point of view of biological data retrieval, are its capability Statistical Summary of Type Register Contents of handling hierarchical data structures and its Following is a statistical summary of the records capability of querying two or more files simultane- on the machine file as of 30 September, 1972: 24 SMITHSOXIAX CONTRIBUTIONS TO BOTANY

No. Families 135 of famil) monographs for the Flora Neotropica se- No. Genera 950 ries. All type specimens seen by him in the course No. Taxa* 10,525 So. Collections 10,625 of his research on these families as of June 1970 are So. Specimens 13,535 recorded, which means that many herbaria are rep- *Species, subspecies, \ arieties, forms, and nothomorphs. resented. With respect to these two families, the Type Register is relatively “complete” in the com- The figures show that the number of collections prehensive, monographic sense in that the file cites averages just slightly more than l/taxon, while the the significant types, regardless of where they are on number of specimens averages about 1.3/taxon. The deposit, as they will be cited in the published number specimens will grow rapidly relative to of monograph. (The monographs of these families the number of taxa as additional herbaria register have since appeared-see Prance 1972a, b.) At the their tjpe holdings of taxa already in the file. same time the coverage may not be as thorough for This is a tally largely of records input from the any individual herbarium as in the case of the tjpe collection of the U. S. National Herbarium other three families-Lamiaceae, Scrophulariaceae, (US); i.e., the vast majority of the families and Violaceae-for which the data were compiled di- genera are represented only by taxa, collections, rectly from type collections or card files in the her- arid specimens in the US type collection. None of baria indicated. Within the Scrophulariaceae, con- the 135 families is present solely on the basis of tributions to the genus Mimulus have been regis- tjpes 1 egistered from another herbarium, which is tered by more than a dozen herbaria (CAN, COLO, to say that at least one taxon in the collection is US DXO, F, GH, JEPS, RlICH, NY, OSC, PH, uc, registered under every family. The inclusion of a LA, LTS, 1171s) as a result of Hale’s experi-ent family does not mean, however, that all US types (see “Introduction”), and there are miscellaneous belonging to that family have been recorded. Quite other contributions to this family recorded from the contrar), the project has only begun, and, as in- 110, NY, and a few other herbaria. dicated earlier, it now is proceeding alphabetically Apart from the families listed and the genus by genus and is still in the letter “C.” This means Carex, for which the catalog is appended, several that for the vast majority of families only genera other groups have been completed in some sense. starting with “A” or “B” are recorded thus far. If a genus is present, however, then all type ma- Nearl) half of the US types of Asteraceae (Compos- terial in the US collection belonging to that genus itae) had been recorded when the switch from a is registered. In other words, the file is complete to systematic to an alphabetical approach was made, the genus level with respect to taxa, collections, and and the file for this family contained as of 30 specimens in the US type collection. September 1972 the following: 255 genera, 2,600 Before the alphabetical approach was started, reg- taxa, and 2,650 specimens. Several important genera istration of US types was essentially completed for of the Brassicaceae (Cruciferae) have been complet- several families, and in each case the types of one ed at least for the US, and the tallies for these or more other herbaria also had been recorded. genera as of 30 September 1972 were: Following is a list of these completed families, show- ing for each the number of genera, taxa, and speci- Arabis (97 taxa, 106 specimens), US Draba (119 taxa, 195 specimens), NY, US mens registered and the abbreviations of the her- Lepidium (29 taxa, 32 specimens), US baria for which the file is relatively complete: Lesquerella (38 taxa, 43 specimens), US Chrysobalanaceae (12 genera, 216 taxa, 1,110 specimens), Thlaspi (15 taxa, 59 specimens), F, GH, MO, XY,RM, S, UC, many herbaria US, WTU Dichapetalaceae (7 genera, 43 taxa, 153 specimens), man\ The data for Thlaspi were provided by Patricia herbaria Lamiaceae (84 genera, 1,234 taxa, 1,851 specimens), LA, MO, Kern Holmgren (1971) of the New York Botanical XY, us Garden from her revision of the genus, during Ecrophulariaceae (85 genera, 943 taxa, 1,073 specimens) , V5 which she saw types from the above-indicated ten Violaceae (17 genera, 232 taxa, 321 specimens), NY, US herbaria. She also provided the data from NY for The data for the Chrysobalanaceae and Dichapet- the genus Draba. alaceae were provided by Ghillean T. Prance of the Finally, John T. Rfickel of the New York BO- Kew York Botanical Garden from his manuscripts tanical Garden provided significant type data on NUMBER 12 25 the genus Anemia subgenus Coptophyllum (Schizae- one who is familiar with the file. Otherwise he will aceae) and its three segregate genera Aneimiaebo- make meaningless or impractical requests. trys, Coptophyllum, and Trochopteris from his By request the file will be queried at cost for monograph of the Anemia (Mickel, 1962), and the anyone. Acy kind of query is welcome, and guid- statistics are: 4 genera, 35 taxa, 81 specimens. ance can be provided in framing queries. It is im- portant at this stage to have feedback from poten- tial users in the form of requests for file queries so Use of Type Register that all needs are taken into consideration as the Some of the main uses of the Type Register will Type Register system undergoes further test and be obvious from the discussion in the foregoing refinement, particularly with respect to report for- sections if not from the concept of the Register mats. Persons wishing to make extensive use of the itself, and other uses will become apparent through Register should plan to spend time in Washington, study of the Carex Catalog and its indices. It D.C., working with the project staff at the Smith- should be emphasized that a catalog of this type sonian. The costs and other requirements of such with the same or other types of indices can be pro- an undertaking should be worked out in advance duced for any taxonomic group, large or small, by consultation with the staff. presently registered, although the data have not The Type Register can be queried or sorted by been edited to the same high degree in any other taxonomic name, author, book or journal title, group thus far. The Carex Catalog and each of its year of publication, collector, collector’s number, indices represent outputs to particular queries. date of collection, country, state or province, county, Many other types of queries are possible, and the herbarium, and kind of type or any combination of amount of output depends on the scope of the these. IVith a query or sort on any of these fields query and the depth and comprehensiveness of the can be printed other selected fields from the same data base at the time of querying. For example, the records, as illustrated by the Carex Catalog, which request “Print all records of ferns,” would yield a is sorted by taxonomic name, and its five indices, relatively small printout at this stage, because only which are sorted in the lead field by author, collec- a few fern types are registered, but eventually such tor, country, publication date, and herbarium, re- a request could yield an overwhelming printout. In spectively. With the cooperation of specialists willing to de- querying the file, the user always must exercise dis- vote time to editing of groups of interest to them- cretion in framing his requests, and to do this he selves, other catalogs can be published. Anyone must have a reasonable knowledge of the limits of is invited to propose collaborations in publishing the machine file beforehand or be guided by some- from the Type Register. A Catalog of the Genus Carex (Cyperaceae)

STANWYNG. SHETLER(Editor); MARYJANE PETRINI,CONSTANCE GRAHAM CARLEY, M. J. HARVEY,LARRY E. MORSE(Assistant Editors); THOMASE. KOPFLER (Programmer); AND COLLABORATORS

Contributing Institutions eral of the above institutions. The Carex project was undertaken in the first place at the suggestion of The ten American herbaria that have collaborat- ed in the Carex project to provide data on their re- the FNA Editorial Committee. The Committee saw in Carex a good model of a large genus with a pre- sective collections of types in this genus are listed ponderance of North American species which pre- here in descending order of number of specimens sents a full gamut of nomenclatural and taxonomic registered. The name of each herbarium is preceded problems to cope with in an information system. by its standard international abbreviation as estab- (The Carex project, insofar as FNA was concerned, lished in the fifth edition of “Index Herbariorum, went beyond the Type Register. While the data Part I, The Herbaria of the World” (Lanjouw and were being gathered for the Register, other morpho- Stafleu 1964). logical data were also being assembled by FNA- NY Herbarium, New York Botanical Garden, Bronx Park, unpublished Carex data on Sections Montanae and New York Ovules collected by A. J. Gilmartin and M. B. GH Gray Herbarium, Harvard University, Cambridge, Moore-to test the matrix techniques of Morse, Massachusetts 1, for computer-assisted identification and key US United States National Herbarium, Smithsonian In- 197 stitution, Washington, D. C. construction.) Once the Carex Catalog was estab- CAS Herbarium, California Academy of Sciences, San Fran- lished on the basis of several large herbaria, other cisco, California major herbaria were able to add their data with a F John G. Searle Herbarium, Field Museum of Natural surprisingly low investment of man-hours. The History, Chicago, Illinois Field Museum, for example, reported just under 35 MO Herbarium, Missouri Botanical Garden, St. Louis, Missouri man-hours-less than a man-week-to compile its DS Dudley Herbarium, Stanford University, Stanford, data on Carex types, at a cost of under $175 for California technical labor, but by the time the Museum came UC Herbarium, University of California, Berkeley, Cali- into the picture it had the benefit of a basic catalog fornia A Herbarium, Arnold Arboretum, Harvard University, already including the contributions of A, GH, MO, Cambridge, Massachusetts NY, and US. JEPS Jepson Herbarium, University of California, Berkeley, Each institution determined its own extent of California coverage, and without exception each tried to in- Each institution has contributed to the Carex clude all types in the general herbarium which al- Catalog voluntarily and with little or no outside ready had been segregated or otherwise identified financial support for work on the Type Register. or which could be identified quickly on the basis of The FNA program has made use of the Type Reg- the available computer printout. At the same time, ister as a pilot project to evaluate various ways of none of the herbaria makes any claim to absolute collecting, editing, capturing, and processing hierar- completeness or thoroughness relative to the num- chically structured taxon/collection/specimen data. ber of types in the herbarium, including those still In the course of this work, therefore, small sums of buried in the general collection. A pragmatic phi- “seed” money were allocated from FNA funds to losophy prevailed, with each herbarium doing the support data-gathering for the Carex project at sev- best it could under the constraints of its own local

26 SUMBER 12 27 circumstances. It might be assumed in general out (personal correspondence) that CAS has a that the coverage is more complete for the herbaria complete set ot H. P. Sartwell’s Carices Arnericanae like NY and US which had separate type collections SeptentiionaliJ, Part 1 (nos. 1-70, 1848) and Part in the first place and had long-established tradi- 2 (nos. 71-143, 1850), and a “rather good set, but tions of segregating type specimens from the main not complete,” of S. T. Olney’s Carices Boreali herbarium as they are recognized, than for the her- Ametzcunae, distributed in 1870 and 1871. Accord- baria without segregated types, but even this as- ing to Thomas, many CAS specimens bear the an- sumption can be questioned. At CAS, a folder-by- notations of J. W. Stacey, who was connected with folder search of Carex specimens in the herbarium CAS and published on western sedges. with printout in hand turned up as many unrecog- Thomas points out these additional interesting nized type specimens as had previously been recog- facts about the other herbaria in the San Francisco nized and segregated into the type collection. A Bay area. Mackenzie, in the course of doing the similar search of North American folders at F re- treatment of Carex for Abrams’ An Zllustrated sulted likewise in doubling the number of recog- Flom of the Pacific States (Mackenzie, 1923), de- nized types. If one pass through the herbarium can termined all the Carex specimens at DS from the double the number of types, it certainly cannot be Pacific States and a number from other regions concluded that all types have now been found. From a purely curatorial point of view, therefore, sometime prior to 1920. The Parish Herbarium, the Carex project was very useful in these instances one of the best early collections of southern Cali- because, as a result, the collection of recognized fornia plants, is housed at DS. The I. W. Clokey types was increased by 100 percent. Herbarium, including his sedge types, are at UC, Clearly it would be unfair to judge the relative and of course W. L. Jepson’s material is at JEPS. size or value of the type collection or even just the In general, every curator who collaborated in Catex type collection of any of these ten herbaria the Caiex project felt that his effort had beneficial on the basis of the present catalog. Much more consequences in the herbarium, and there was a extensive development of the Type Register is re- nearly unanimous opinion that the results were quired before such judgment will be warranted. well worth the effort from the curatorial stand- At the same time, the present ranking is almost point alone. predictable. An herbarium that is the major de- pository for the types of a monographer naturally Data-Collection Procedure is expected to have a disproportionate number of types in the group (s) on which the monographer The Carex Catalog was initiated with data from worked. The extensive work of monographer K. K. the US and hfO type collections. For the US, rec- Mackenzie (1 93 1-35, “Cyperaceae-Cariceae,” in ords were converted from the existing card file hrorth American Flora) clearly might be expected, (see “Source of Data”), while the MO records were therefore, to have put NY in the first position, and captured from data forms filled out at MO in the no one will be surprised to find GH and US close course of a special search of all the Carex type behind. Comparison of the NY and US type col- folders, which are distributed through the herbar- lections of Carex provides a good example of the ium in association with the main collections. The influence of a monographer on a type collection. special search was undertaken specifically as part Together, NY and US have type specimens for over of the Carex project of FNA. 400 taxa. Of these taxa, half are represented by After a common catalog of the US and MO col- type specimens only at NY, while about 40 are represented only at US. Thus, NY has five times lections was compiled, members of the Type Regis- more “unique” Carex taxa in the Type Register ter staff moved their data-conversion operation to than US. By contrast comparisons in some other the New York Botanical Garden for two weeks, groups show NY and US to have about an equal where, with the assistance of NY personnel, the number of “unique” representatives in the Regis- NY Carex data were compiled and input to the ter. CAS is expected to be strong in types of west- system in a matter of days. The data were com- ern taxa. In addition, John Thomas has pointed piled from the specimens in the separate type col- 28 SMITHSOXIAN CONTRIBUTIONS TO BOTANY lectioii, which by and large does not include the tollectoi and collector’s number to the US + h10 literature citations, and supplemented with bibli- + NY catalog. ographic data from reference works, primarily The collaborators in the San Francisco Bay area Mackenzie’s (193 1-35) monograph of Carex. weie provided with a new catalog incorporating It should be pointed out that the type collec- the A and GH data along with the US, MO, and tion at NY was first segregated for security reasons NY data, and thia catalog also included an index during World War 11, and the selection had to be by collector and collector’s number. This catalog done in haste under less than ideal conditions. was annotated by them with new data from CAS, Consequently, many nontype but historically im- DS, JEPS, and UC on taxa already registered, and portant specimens, constituting perhaps as much as data for taxa new to the file were compiled on 25 percent of the total type collection, deliberately standard forms. The annotated catalog and com- were pulled along with the known or presumptive pleted forms then were returned to the Smithsonian type specimens in the process of going through the Institution for input to the system. main herbarium. The person who compiled Cnrex ’it CAS, DS, F, JEPS, and UC the data were data attempted to sort out the nontype material, compiled in the first instance from the specimens and further culling was done during the editorial in the herbarium, but in most cases the original process. Undoubtedly some nontype material still publications also were checked when taxa new to remains in the present Catalog, although it seems the file were involved. Otherwise the bibliographic unlikely that the percentage of such specimens is data were obtained from secondary sources. The much if any higher for NY at this stage than for original literature was checked for taxon additions any of the other nine herbaria. In any event, it without exception at F. will be a simple matter to delete nontypes from In summary, a round-robin procedure of sorts the file as they are discovered and brought to the was used to collect the data. To the extent possible, attention of the Type Register editors in the future. each new collaborating institution was given the benefit of the latest cumulative catalog incorporat- Once the NY data were merged fully with the ing the contributions of previous collaborating US and MO data, a union catalog was delivered to herbaria. In this way, maximum advantage could Harvard University for additions from A and GH. be taken of previous herbarium and library re- .4t Harvard, data on types of taxa already regis- search, and duplication of effort was kept to a tered were annotated in the catalog, and data for minimum. Once the combined data of CAS, DS, taxa new to the catalog were compiled on standard TEPS, and UC were returned to the Smithsonian, forms. In general, Harvard provided only specimen the data-collection phase was closed out and final data, and the bibliographic data were looked up editing began. and supplied later by the Type Register staff at the Smithsonian before annotations and new records Editorial Process were captured ant1 merged with the US + MO All entries were edited in accordance with the + h’Y machine file. principles and procedures set forth in the first part, When the annotated catalog was returned from and the editors take final responsibility for the Harvard, the X and GH annotations were tran- form and style and all other editorial matters of scribed to the main working catalog at the Smith- the present Catalog. To integrate new contribu- sonian, and then the Harvard catalog, as annotated, tions into the accumulating data base, differences was sent to tkField Museum of Natural History. between supposedly identical records from different Sent with the annotated catalog were photocopies institutions constantly had to be reconciled by turn- of the data forms for the new taxa added by A ing to standard references and the original litera- and GH. In this way it was possible to give the ture. Whenever possible, record content was verified collaborators at F the benefit of the new data sup- by checking the original publication. In the end, plied by A and GH immediately, while the capture nearly every original description cited in the Cata- of these new data was still in progress at the Smith- log was seen at least once by the editors and in sonian. As a further aid to their work, the col- many cases several times. In many cases, further- laborators at F were provided with an index by more, the designation of kind of type was validated SUMBER 12 29 according to the ZCBh’; however, because much of was begun. This iterative process continued until the checking of original literature was done by a the editing began to yield diminishing returns and technical editor, untrained in the application of it became necessary to bring the never-ending proc- the type method and terminology, many of the ess to a reasonable stopping point, which the pres- records were verified without being validated, to ent Catalog is believed to represent. In such a use the distinction defined in an earlier section dynamic system, editorial perfection is relative at (p. 00). According to this distinction, the develop- best, and at this stage the editors certainly make ment of the Cuiex file of the Type Register can be no claim to perfection in any sense of the word, said to have attained the Stage 2 “verification” aithough every effort has been made to be thorough level overall, with some records still at the Stage 1 and consistent. “registration” level and with a substantial number of others having been “validated” more or less ac- Milestone Events cording to Stage 3 standards. Of course the careful scrutiny of specialists, with appropriate feedback Computerized data banks are a new development from them, is needed over a period of years before in biology, and there is still much to be learned the present Cmex file can be said to have truly about the mechanics of creating and maintaining attained Stage 3 development. them. The process is all too easily underestimated, While the bibliographic data could be double- especially with respect to manpower requirements, checked or supplied (if not provided in the first and overly optimistic timetables are the rule. An place) by examining the original publication, the enormous effort on the part of many individuals specimen data could be verified in this way only to \vent into the creation of the Cal-ex Catalog over the extent that the publications gave corroborating a period of more than two years. During this time details. Ultimately, therefore, each contributing in- the tedious editorial work seemed to go on end- stitution is responsible for the reliability of its own lessly, while technical problems with the systems specimen data. In the case of NY, however, the development and processing also came in a steady editors, who assisted in the data collection itself, stream. The following chronology of milestones in share responsibility for the reliability of the speci- the more than two-year process has a two-fold men data. In any event, citations always were purpose-first, to provide a practical example of checked against Zndex Kewensis and/or the Gray Herbni !urn Zndpx if the original publications could the laborious steps involved in creating a data not be examined. bank, and, second, to caution against overenthusi- Editing of this type of open-ended file, in which asm and oversimplification on the part of others It considerable subjective judgment is required, is a contemplating similar efforts. must be stressed, neyer-ending process, and a reasonable degree of however, that all during the two and one-half thoroughness is achieved only after many editorial years other taxonomic groups were being input to “passes” through the file. The editing proceeded the Type Register along with Curex, and it never in a series of phases and cycles in which all records was possible to work exclusively on Carex. were examined a field at a time, by means of in- 1970 verted listings or indices, for consistency and ac- January Decision made to begin work on genus as part curacy, and updated printouts were obtained for of FNA Carex project. Editing of data in US another editorial round. Thus, for example, title card file begun. citations were standardized in one editorial phase, February Collection of data begun at MO and first records returned to US for input. while names of authors were standardized in an- Photocopies of all Carex records in US card file other phase. The editorial corrections for all fields sent to MO. then were merged into a common working hard- March h’ew data-capture procedure organized using on- copy of the file before the process of updating the line, text-editing system. machine file was begun. After one cycle of such April ,411 US and MO records input, totaling just over 200 taxa and about 250 specimens. editing was finished and the file was thought to be May First printout of US + MO records produced “clean,” a new catalog and set of indices were and edited, and machine file updated. printed out. Then a whole new cycle of editing Second printout produced, and copy sent to NY. 30 SMITHSOSIAN CONTRIBUTIONS TO BOTANY

June Second printout of US + MO records edited, antl Report on progress to meeting of FNA Program machine file updated. Council in Miami, Florida. Members of Type Register staff worked at NY for So\ ember Arrangements made for CAS, DS, JEPS, and uc last two weeks during which time all SY to contribute, and necessary materials supplied. Cares records (200-k taxa, 300 specimens), cop) Of collector index sent to MO. along with records of other groups, were input Critique Of Sample catalog of 100 Carex received to the system on-line via telephone to computer from F. J. Hermann. in Washington, D.C. Editing of A and GH data completed, and First printout of new taxa added to file by NY records input to system, produced. December Data collection at F completed, Report on NY work prepared. July Sew cumulative catalog incorporating A, GH, August Printout of NY additions edited, antl machine hfO, NY, and US produced, with 537 taxa, 542 file updated. collections, and 776 specimens. First printout of US + MO + NY records pro- duced. Four indices in preliminary format generated. COBOL paragraphing (concatenating) program ceptember rS + MO + NY printout edited, antl machine designed, written, and debugged: GIS/COBOL file updated. interface programmed and tested. cecond printout of joint records produced, total- ing 415 taxa and 544 specimens. 1972 Decision made to publish a Carex catalog as the sample installment of Type Register. January All data returned from F to US for input. Novemtier Preliminary report formats defined for catalog A11 data returned by C.4S, DS, JEPS, and UC. and indices. Limited input with magnetic cartridge system December Use of on-line system discontinued, and paper begun. tape system reorganized and used again. Computer program for listing and tallying taxa 1971 in Type Register tested. Data collection phase essentially brought to January- Intensive editing and updating of preliminary close. Carex Catalog (US MO NY) continued. May + + Sample catalog produced using COBOL con- OCR data-capture system inaugurated, and paper catenation program. tape system phased out. Februar) Annotations from catalog used by F and from Sample catalog of 100 Carex records with 4 cross- indices produced and distributed to a limited catalog used by CAS, DS, JEPS, and UC trans- cross-section of taxonomists. ferred to working copy. June Preliminary catalog of Carex displayed at botani- Editing and final data capture continued apace. cal meetings in Edmonton, Alberta, and copies COBOL concatenating program, including GIS of sample of 100 records handed out. interface, tested satisfactorily. Decision made to hold off publication of Catalog March- Xew curnulatit e catalog printed, including 600 at least until GH could be included. April taxa, 607 collections, and 854 specimens. July Final printout of US + hIO + NY catalog Nine indices printed for final field-by-field edit- generated for GH. ing. August Copy of latest catalog delivered in person to lt'riting of introductory sections of this paper Harvard University for use in compiling data begun. from A and GH; specimen data compiled and May New catalog printed, including 609 taxa, 615 returned to US. collections, and 1,050 specimens. Copy delivered also to SY for final prepublica- Xew set of nine indices also produced. tion check: check made and printout returned. Editing and updating of file continued. Copy sent to MO for final prepublication check. Final prepublication edition of Catalog printed, Copy displayed at annual meeting of American Institute of Biological Sciences, Ft. Collins, along with indices, which included 606 taxa, Colorado. 612 collections, and 1,059 specimens. Contributions solicited by letter from F, Sari Final format of Catalog established after various Francisco Bay area herbaria. tests with COBOL program. Feptember F indicated willingness to contribute Carey Several qpes of computer paper tested. records. June Camera-ready copy of present Catalog and in- October Arrangements made for F to contribute records. dices produced for Smithsonian Institution and necessary materials supplied, including Press, including same data base as final catalog index by collector to preliminary catalog. in May. Data collection begun at F. August Manuscript and camera-ready copy of Catalog Editing of A and GH data begun. and indices sent to press. SUMBER 12 31

Tlie decision to use Carex for the trial publica- The indices, in addition to providing access to tion was made in September 1970, exactly two years the Catalog by other criteria than the taxonomic before the manuscript finally went to press. What name, represent data files in their own right which is obvious from this chronology is that for every may serve a user’s purpose without his ever taking adtli tional collaborating institution the prepara- recourse to the main Catalog. They are independ- tion ot a final catalog is delayed at least a few ent data files because they all include the name of months. Under the best of circumstances, turn- the taxon and one or more other fields that place around time between c)cles of data-collecting, edit- the key indexing field in the context of related ing, and processing quickly add up to days, weeks, data. The data source code has not been included and months. Clearly, the task of creating a data record by record because it is usually 0s; further- base of this type is limitless ultimately, and definite more, the classification of sources has been in use bounds must be established if the dynamic process for only a short time and was not used throughout is ever to stand still long enough to yield mean- the development of the Carex Catalog (see “Data ingful products. Desirable as it would have been, Source Code”). therefore, to include many more herbaria in this The Catalog provides citations of original au- Catalog, tlie line had to be drawn finally. If the thors ant1 publications, data on type collections, Catalog had been closed off after the NO, KY, and and a list of type specimens known to exist in the US contributions, as originally planned, it would ten herbaria surveyed-all subordinated to the tax- have gone to press a year earlier, in mid-1971. The onomic name. No taxa are included that are not addition of seven more herbaria, including several represented by at least one specimen in at least key ones, surely justifies the year’s delay, but it is one of the ten herbaria. The organization of the doubtful that further delajs could be justified at data in the paragraphed unit-entry is as follows: this time for any other herbaria. We believe that SPECIFIC EPITHET/RANK/INFRASPECIFIC EPITHET/ it is vitally important now to get the concept of AUTHOR OF NAME, CITATION OF ORIGINAL PUB- tlie Type Register arross to the botanical com- LICATION. DATE OF PUBLICATION. munity through the medium of the Cnrex Catalog COUNTRY: STATE OR PROVINCE: COUNTY: LOCAL- without further delay. In(COMMENT IF ANY) (COLLECTOR, COLLEC- TOR’S NUMBER. DATE OF COLLECTION) HERBARILX ABBREVIATIOS/SHEET NUMBER/ Use of Carex Catalog KIND OF TYPE HERBARIUM ABBREVIATION [etc., for additional The Carex Catalog consists of the “Catalog of specimen] Specimens” and cross-indices to the “Catalog of COUNTRY [etc., for additional collection] HERBARIUM ABBREVIATION [etc.] Specimens” by five different fields (descriptors): (1) “Author Index,” (2) “Publication-Date Index,” The five indices are all cross-referenced to the (3) “Collector Index,” (4) “Geographic Index,” Catalog by means of the specific epithet instead of and (5) “Herbarium Index.” (Hereafter the term a page or record number because the unit records “Catalog” is used for the “Catalog of Specimens” in the Catalog are alphabetical by epithet, allowing proper.) The Catalog is arranged alphabetically by for fast look-up. Space did not permit printing in- taxon and includes the full unit-record for each fraspecific epithets in four of the indices; therefore, taxon, as it is stored in the computer, except for the specific epithet is prefixed by an asterisk (*) if the family and genus names, the data source codes, the record being cross-referenced is not the species and several file-control dates and numbers. The itself but one of its infraspecific taxa. This device family name and genus name, Cyperaceae and should permit the user to get to the desired entry Carex, respectively, have been omitted because almost as quickly as if the infraspecific epithet had they are the same for all taxa and, printed at the been printed, as in the “Herbarium Index.” Con- top of each entry, would constitute unnecesary tent and format are more or less self-evident in each words that would tend only to hide the key words of the indices. It should be emphasized that these for alphabetization, the epithets. The Catalog is are only five of an almost infinite number of pos- alphabetized, therefore, by the specific and infra- sible indices. Furthermore, they all were generated specific epithets. directly from the exact same data base from which 32 SMITHSONIAN CONTRIBUTIOXS TO BOTANY

the Catalog itself was produced and have identical ing the “Collector Index” with the “Author In- data wherever content overlaps. dex.” It becomes clear at once, for example, that The purpose of the “Author Index” is to point while some authors collected almost as many new to all taxa in the Catalog described by a particular taxa as they described, others described many more author or combination of authors, and, therefore, than they collected. Further examination indicates the Index is alphabetized primarily by author and that this difference may reflect the difference be- secondarily by specific epithet. Combinations of tween a floristic taxonomist like M. L. Fernald, who authors are alphabetized as combinations, not as tended to describe his own species, and a mono- individual authors, owing to present system lim- graphic worker like K. K. Mackenzie, who described itations, A person interested in a particular taxon taxa from among specimens of many collectors described by a particular author or author combina- amassed for a study of the genus. tion can learn quickly from the Index whether The “Geographic Index” provides access to the there is an entry in the Catalog for this taxon and collections of the Catalog primarily by country of then go to it. Persons wishing to study an author’s origin and secondarily by state, province, or equiva- descriptions chronologically, by year of publication, lent. The specific epithet, which is the cross- would want to have this Index resequenced with the reference to the Catalog, is the tertiary sort-key. The date rather than the epithet being the secondary value of this Index will be obvious at once to sort-key. floristic workers who wish to know which taxa in The primary sort-key of the “Publication-Date their region are typified by specimens collected Index” is the year of publication, from the oldest within the region, e.g., within the State of Cali- to the youngest, the actual range being from 1803 fornia. to 1971. Within each year the secondary sort-key The “Herbarium Index,” alphabetized in the is the specific epithet, which references an entry in the Catalog. This Index should be useful to those first instance by institution, lists alphabetically by who are interested in nomenclatural priority in the specific and infraspecific epithets under each of the genus or in tracing the historical development of ten institutions the taxa in the Catalog for which knowledge about Cnrex. By itself, the Index, which they hold type specimens. The herbarium sheet includes the name of the author or author combi- number, if there is one, and the kind of type are nation, is a chronological summary of the botanists listed also. The sheet number is the tertiary sort- who have described species in the genus as regis- key. Many of the type designations have not been tered in the Catalog. validated, and the problems of validating kind of The “Collector Index” is in effect a list by col- type have been discussed at length earlier in this lector of collections cited in the Catalog and paper (pp. 8 and 19-21). For this reason many provides a convenient means of checking any her- of the type specimens registered still carry the origi- barium for type material of taxa recorded thus far nal designation of the herbarium submitting the in the Carex file of the Type Register. Under the data. name of each collector or combination of collectors The following informal or archaic terms, which are listed the numbers and dates of all collections appear in the Catalag and in the “Herbarium In- in the Catalog, cross-referenced by the specific epi- dex” but which are not sanctioned by the ZCBAI, thets. The name is the primary sort-key, and the require brief definition of their meaning or ap- specific epithet is the secondary sort-key. A varia- parent meaning in the Catalog. It should be tion of this Index would include a list of all her- pointed out, however, that the presence of a cor- baria in which the collector’s type specimens are rect term (e.g., holotype, isotype) is no guarantee deposited. In other words, the kind of guide to that it has been used correctly. A large number of the location of types that A. S. Hitchcock and his the designations have been validated, and where colleagues were trying to compile directly in the the terminology seems to be used consistently 1930s (see “Introduction”) can be produced as a within a collection, e.g., where one holotype and by-product of the Type Register data base. Some one or more isotypes are indicated, it can be as- interesting observations can be made by compar- sumed that the designations were validated. SUMBER 12 33 rotype Presumptive syntype or isosyntype, but may No. 2. ABLATA BAILEY, L. H. The citation following be isotype or paratype, if a type at all. the author’s name is to the place where the neotype is type Presumptive holotype, but may be isotype, designated, because the original publication did not designate syntype, isosyntype, or paratype, if a type at a type specimen, and the citation of Bailey’s original publica- all. tion is cited as a parenthetic remark at the end of the type collection Term used in a proper sense for a specimen geographic locality. To be consistent with the rest of the of a single collection cited by original authoi Catalog, however, where the citation after the author is without designating a holotype-in this sense, (I~N\Sthe citation of the original publication, the editors a syntype or isosyntype according to ICBN should have reversed the two citations in this record. As a (see pp. 19-21); otherwise term flags a pre- result of this editorial error, Bailey in the “Author Index” and sumptive syntype, isosyntype, paratype, or “Publication-Date Index” appears to have described C. even isotype, if a type at all. ablata in 1935, when Mackenzie designated the neotype, type fragment Presumably fragment of the holotype, but rather than in 1888. This error does point up the problem of may be fragment of isotype, syntype, isosyn- dealing with neotypes, however, and a certain logic can be type, paratype, or other kind of type, if of advanced for either way of handling the two citations. type at all. type materiul Presumptive type specimen of some kind- No. 99. CHIHUAHUAENSIS. Spell CHIHUAHUENSIS, omitting second “A.” catchall term. No. 418. PIRCHINCHENSIS. Spell PICHINCHENSIS, The Catalog and Indices were printed by com- omitting “R.” puter directly from the data base, and not a single No. 424. PLUVICA. Spell PLUVIA, omitting “C.” So. 446. PURPUREOVAGINATA. Spell PURPUREO- change has been made. What is presented here is VAGINATA, inserting hyphen. exactly what was stored on the machine file as of No. 448. PYCNOTHYSOS. Spell PYCNOTHYRSOS, insert- 16 June 1972, with the exception of punctuation ing “R.” between fields which may have been added in the No. 544. TENERA VAR. RICH11 FERNALD, M. L. At process of concatenation. Because the system pro- the end of the geographic locality, MIDDLESEX FALLS should read MIDDLESEX FELLS. vided only for printing in upper case letters, possi- No. 549. TERRAE-NOVAE FERNALD, M. L. The hy- bilities for variation of typography were limited. phen should be removed from the collector’s name GIL- Boldfacing by overprinting the same words two or BERT-JR., F.A. The practice of joining Jr. to the end of the more times was used for the specific and infraspe- collector’s or author’s last name by a hyphen in this manner cific epithets, and in the process it was necessary was required by the specifications of the pilot processing also to boldface rank designators connecting epi- s\steni, but this requirement no longer obtains. thets. Thus the taxon names stand out from the Note: Changes in spelling of specific epithets also rest of the text and facilitate searching the Cata- apply wherever these epithets have appeared in log. Insofar as practical, the standard conventions the indices. of punctuation in nomenclatural literature were used. Errata Statistical Summary of Catalog 606 taxa (species, subspecies - SSP, varieties - VAR, Several errors were detected in the Catalog after forms-FOR, nothomorphs-NM.) the camera-ready copy had been produced. By rec- 612 collections ord number in the Catalog, these are: 1,059 specimens (sheets) CATALOG OF SPECIMENS

-A-

10 ABDITA BICKNELL9E.P.9 BULL. TORREY BOT. CLUB 35:492. 19080

USA: NEW YORK: LONG ISLAND9 RICHMOND HILL (BICKNELLTE.P*T -0-0 11 MAY 19@4) NY TYPE

2. ABLATA BAILEYTL.H.T No AMER. FL. 18:314. 1935. CANADA: BRITISH COLUMBIA: VANCOUVER ISLAND, MOUNT MARK; ALTO 250C FT.; (ORIG. PUB.: BOT. GAZ. 13:82. 1888.1 (MACOUNTJOHNT 134@1. 26 JUL 1887) GH I SONEOTY P E NY I SONEOTYPE

30 ABORIGINUM JONES9M.E.9 BULL. MONTANA STATE UNIV.9 BIOL. SER. 15:69. 1910 USA: IDAHO: ADAMS COO: INDIAN VALLEY: ALTO 2300 FT. (JONES,M.E.T --- 12 JUL 1899) CAS 242617 ISOTYPE DS 149709 I SOTYPE NY ISOTYPE

4. ABRAHSII MACKENZIE,K~K.T BULL. TORREY B0To CLUB 36:482. 1909. USA: CALIFORNIA: SAN BERNARDINO C0.t SAN BERNARDINO MOUNTAINST BEAR VALLEY (ABRAMStL., 28160 31 JUL 1902) DS 55317 ISOTYPE F 186491 I SOTYPE MY HOLOTYPE

50 ABRUPTA MACKENZIE,K*K.9 BULL. TORREY 601. CLUB 43:6180 1916. USA: CALIFORNIA: BUTTE Coo: STIRLING CITY; ALTO 3000 FT. (HELLERTA*AoT 10820. 07 JUN 1913) DS 64125 ISOTYPE NY HOLOTY PE

60 ABSCONDITA VARo ROSTELLATA FERNALD9M.L.9 RHODORA, 4423860 1942. USA: VIRGIYIA: ISLE OF WIGHT COO: LEE'S MILL (FERNALD9M.L. AND LONGIB., 120120 08 JUN 1940) GH ISOTYPE YO TYPE COLLECTION NY TYPE COLLECTION us 2003161 TYPE COLLECTION 70 X ABSCONDITIFORHIS FERNACD9M.L. , RHODORA 44:387. 19420 USA: VIRGINIA: SUSSEX COO: NOTTOWAY RIVER9 HUSKE (FERNALDfMoLa AND LONGvB.9 12969. 13 JUN 1941) GH HOLOTY PE GH ISOTYP E MO 1306480 ISOTYPE 34 NUMBER 12 35

YY ISOTYPE us 200 3299 ISOTYP E

ACCEDENS HOLMTH.T.,T AMER. J. SCI. SER.49 16:457. 1903. USA: OREGON: MULTNOMAH COO: SAUVIE ISLAND (COLUMBIA RIVER AT MOUTH OF WILLAMETTE RIVER) (HOWELL9T.J. T --- -- MAY 1880) GH SY NTY P E YO SY NTYP E

9. ACROPHILA BLAKETS~T.~J. ARNOLD ARBOR. 28:114. 1947. INDONESIA: WEST NEW GUINEA: LAKE HABBEMA; (COUNTRY AS "DUTCH NEW GUINEA") (BRASSTL.J.~ 9515. -- AUG 1938) A ISOTYP E

10 ACUTA VAR, PALLIOA BOOTT9F.p ILL. GENUS CAREX 4:1669 PL.554. 1867. USA: OREGON: "FORT COLVILLE TO ROCKY MOUNTAINS9 WEST KOOTENAY" ( L Y AL L T DA V ID T --- . -- --- 1861) GH TYPE COLLECTION

11. ACUTINA BAILEYTLoH*T MEMO TORREY BOT. CLUB 1:52. 1889. USA: OREGON: DESCHUTES RIVER (HOWELLTT.J.T 935. 09 MAY 1885) F 206585 TYPE MATERIAL GH ISOTYPE NY TYPE COLLECTION us 25164 TYPE COLLECTION us 817087 TYPE COLLECTION

12. ACUTINELLA YACKENZIETK.K* T N. AMER. FL. 18:407. 1935. USA: OREGON: -- (HENDERSONTL.F.~ 13. -- --- 1883 1 us 27286 HOLOTYPE

13 AOUSTA VAR. GLOMERATA OLNEY 9s.T EX BAILEYTL oHo 9 BOT. GAZ. 9: 139. 1884. CANADA: NEW BRUNSWICK: KENT COO: SALMON RIVER (FOWLER9J.q ---. __ --- 1872) GH SYNTYPE

14. AENEA FERNALD9M.L. 9 PROC. AMER. ACAD. ARTS 37:480. 1902. USA: NEW HAMPSHIRE: GRAFTON COO 2 FRANCONTAT FOREST HILLS HOUSE (FAXONTE. AND FAXONTC.E.~ --- 23 JUN 18881 GH SYNTYPE

15- AEQUA CLARKETC.B.T BULL. MISC. INFORM. ADD.SER.8:86. 1908. USA: CALIFORNIA: SAN MATE0 CO,: SAY MATE09 CRYSTAL SPRINGS LAKE (BAKERvC.F.9 811. 10 MAY 1902) GH TYPE COLLECTION NY TYPE COLLECT ION

16. AESTIVACIFORMIS MACKENZIETK~K.9 BULL. TORREY BOT. CLUB 37:238. 1910. USA: NEH JERSEY: PASSAIC COO: GREENWOOD LAKE (MACKENZIE,K.K. T 2676. 23 JUN 1907) GH ISOTYPE 36 SMITHSONIAN CONTRIBUTIONS TO BOTANY

YO TYPE MATERIAL

17. AESTIVALIS CURTIS9M.A. EX GRAY9A.i AMER. J. SCI. ARTS SER.1, 42:280 1841. USA: NORTH CAROLINA: -- ( CURTISTM.A.T --- -- JUL 1841) YO TYPE MATERIAL NY TYPE MATERIAL

18. AGGLOMERATA MACKENZIE,K.K.r BULL. TORREY BOT. CLUB 33:442. 1906. USA: MISSOURI: JACKSON COO: COURTNEY (BUSHTB.F.T 1718. 25 MAY 1902) YO TYPE COLLECTION us 440179 TYPE COLLECTION

19. AGGREGATA MACKENZIETK~KOTBULL. TORREY BOT. CLUB 37:246. 1910. USA: MISSOURI: JACKSON COO: COURTNEY (BUSHTBOF~T1718. 25 MAY 1902) NY TYPE

20 0 AGROSTOIDES YACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 34:607. 1907. USA: NEW YEXICO: SOCORRO COO: LUNA; ALTO 6500 FT. (WOOTONTE.O.T --- 28 JUL 1900) us 617798 TYPE us 694342 TYPE

21 0 ALATA TORREYqJ.9 ANN. LYCEUM NATO HIST. NEW YORK 3:396. 1836. USA: NORTH CAROLINA: CRAVEN COO: NEW BERN (CROOMTH.B.T ---. -- --- 1834) NY SYNTYP E

22 0 ALATA VARo FERRUGINEA FERNALO~Y.L.T PROC. AMER. ACAD. ARTS 37:477T PL.2. 1902. USA: OHIO: 0- (SULLIVANTTW.S.T --- . ---I GH HOLOTY P E

23. AtBIDA BAILEY7L.H.r MEMO TORREY BOT. CLUB 1:9. 1889. USA: CALIFORNIA: SANTA ROSA CREEK (BIGELOW9J.M. 9 -0-0 -- -*- 1853-1854) NY TYPE COLLECTION

24. ALBO-NIGRA MACKENZIE9K.K. IN RYDBERG,P.A. 9 FL. ROCKY MOUNT. 1379 1060. 1917. USA: WYOMING: PARK COO: NEEDLE MOUNTAIN (CARY9M.p 613. 11 JUL 19101 us 858947 TYPE COLLECTION

25. ALMA BAILEY,L.H.r MEMO TORREY ROT. CLUB 1:50. 1889. USA: CALIFORNIA: -- (PARRYTC~C. AND LEMMON*J.G.r 396. -- --- 1876 1 CAS 497554 ISOTYPE NY I SOTY P E

26, ALOPECOIDEA TUCKERMANTE~TENUM. CARIC. 18. 1843. NUMBER 12 57

USA: NEW YORK: YATES COO: PENN YAN (SARTWELL,H*P*, --- . ---I F 32699 ISOTYPE f 32700 ISOTYPE F 56916 ISOTYPE F 349624 ISOTYPE F 373673 ISOTYPE F 373679 ISOTYPE GH ISOTYPE MY ISOTYP E 21. ALOPECOIDEA VAR. SPARSI-SPICATA DEWEYtCor AMER. J. SCI. ARTS SER.2, 8:350. 1849. USA: MICHIGAN: MACOMB COO: WASHINGTON (COOLEY,D.q 74. ---I CAS 553879 ISOTYP E GH ISOTYPE MY ISOTYPE

28- AMPHIBOLA STEUDELIE~G.~ SYN. PL. GLUM. 2:234. 1855. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMOND,T., 437. ---) NY TYPE COLLECTION 29 . AHPHIBOLA VAR. TURGIDA FERNALDTM.L*~ RHODORA 44:311. 1942. USA: NEW YORK: TOYPKINS COO: ITHACA (WIEGANDTK.M. AND THOMASTC.C.T 1915. 15 JUN 1914) GH HOLOTY PE

30. AMPLISQUAHA HERMANN,F*J., RHODORA 57:158. 1955. USA: GEORGIA: GILMER COO: CHATSWORTH (PYRON,J.H. AND MCVAUGH~R.T 2951. 15 MAY 1938) us 2231424 TYPE

31. ANGUSTIOR MACKENZIETK~K. IN RYDBERG,P.A.q FL. ROCKY MOUNT. 124, 1060. 1917. USA: DISTRICT OF COLUMBIA: -- (STEELE,EoS.r --* . -- --- 1896) NY TYPE COLLECTION

32 ANGUSTIOR VARm GRACILENTA CLAUSEN9R.T. AND WAHLTH.A., RHODORA 41230. 1939. USA: PENNSYLVANIA: CENTRE COO: INGLEBY (CLAUSEN,R*T. AND WAHLqH.4.9 2532. 06 JUN 19371 GH TYPE

33. ANTHOXANTHERA PRESL,K.B.r REL. HAENK. 3:203. 1828. USA: ALASKA: NUTKA SOUNO ("SINUS NUTKA") (HAENKEqT. T --- . ---I us 865058 TYPE

34. APERTA BOOTT,F. IN HOOKERTW.J.T FL. BORo-AYER. 2:218. 1839 ( "1 840") USA: WASHINGTON: COLUMBIA RIVER (SCOULER, J. 9 -0- . ---I GH SYNTYPE

35. APERTA VAR. UMBROSA KUKENTHAL,G.T REPERT. SP. NOV. REGNI VEG. 38 SMITHSONIAN CONTRIBUTIONS TO BOTANY

26: 254. 1929. USA: WASHINGTON: KLICKITAT COO: BINGEN (SUKSD0RFtW.N.p 12347. ---) CAS 242962 ISOTYPE

360 APERTA VAR. VIRIOANS KUKENTHALt Got REPERT SP. NOV. REGNI VEG. 26 :254. 1929. USA: WASHINGTON: KLICKITAT COO: BINGEN (SUKSDORF9W.N.9 12348. 15 SEP-23 OCT 1927) CAS 242961 SYNTYPE USA: WASHINGTON: KLICKITAT C0.Z BINGEN 4 SUKSD0RFpW.Net 12359. 23 OCT 1927) CAS 242960 SYNTYPE

37. APODA CL0KEYtI.W.t AMER. J. SCI. SERo5t 3:88t PL.2. 1922. USA: IDAHO: CUSTER COO: MACKAY (NELSONgA. AND MACBR1DEtJ.F. 9 1533. 01 AUG 1911) NY ISOTYPE uc 90 543 9 HOLOTYP E

38. APOOOSTACHYA OHW1,J.t JAP. J. BOT. 7:188. 1934. TAIWAN: --: MOUNT NANKO-TAISAN (0HWItJ.t 4182. -- JUL 1933) F 1464064 TYPE MATERIAL 39 . AQUATILIS WAHLENBERGtG.9 KONGL. VETENSK. ACAD. NYA HANDL. 24:165. 1803 USA: NEW YORK: SENECA CO.: JUNIUS (SARTWELLtHoPot 56. ---I NY TYPE COLLECTION

40 0 AQUATILIS VAR. SUBSTRICTA KUKENTHALtG. IN ENGLERtH.G.A.9 PFLANZENR. 49 FAM. 20: 309. 1909. USA: NEW YORK: SENECA COO: JUNIUS (SARTWELL,Ho\P.t 56. ---I CAS 554019 ISOTYPE GH ISOTYPE YO TYPE COLLECTION

410 ARAPAHOENSIS CLOKEYt 1.W.t RHODORA 21:83. 1919. U'SA: COLORADO: BOULOER COO: MOUNT ARAPAHOE; ALTO 11700 FT. (CLOKEYtI.W.9 3227. 29 JUL 1918) C AS 102030 ISOTYPE DS 109019 ISOTYPE GH ISOTYPE NY I S OTY PE uc 905436 HOLOTYPE

42 ARCTAEFORMIS YACKENZ1EtK.K. 9 No AMER. FL. 18:97. 1931. CANADA: BRITISH COLUMBIA: ELGIN (HENRY9J.K.t 9152. C4 JUN 1915) NY TYPE

43. ARCTICA DEHEYtC., AMER. J. SCI. ARTS SER.1, 27:239. 1835. CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51". 9 106 13'W. 1 (RICHARDSON9J.9 --- . --- b NUMBER 12 39

YY TYPE COLLECT ION 44 . ARGYRANTHA TUCKERMANtE. EX DEWEYTC., AMER. J. SCI. ARTS SER.29 29:346. 1860. USA: MASSACHUSETTS: HAYPSHIRE COO: AMHERST ( TUCKERMANTE. 9 ---. ---) GH HOLOTYP E GH ISOT YP E

45 0 ARISTATA VARo LONGO~LANCEOLATA DEWEYIC.T AMER J SCI ARTS SER.2, 18:102. 1854. USA: NEBRASKA: BAD LANDS ("MAUVAIS TERRES") (HAYDENTF.V., ---. -- --- 1853) GH HOLOTY PE

46 ARSENII KUKENTHALTG. T REPERT, SP. NOV. REGNI VEG. 8:3266 1910. MEXICO: MICHOACAN: YORELIA (ARSENETG. (FRERE) T 3054. 16 JUL 1909) GH ISOTYPE NY ISOTYP E us 1030011 TYPE COLLECTION

47. ARTITECTA VAR. SUBTILIROSTRIS HERMANNtFoJor RHODORA 40:79. 1938. USA: INDIANA: VERMILLION COO: CLINTON (DEAM,C.C.T 54764. 05 MAY 1934) GH HOLOTY PE

480 ASSINIBOINENSIS BOOTTTW.T BOT. GAZ. 9291. 1884. CANADA: MANITOBA: ASSINIBOINE RAPIDS (YACOUN9JOHNt 52. 14 JUN 18791 GH SYNTYP E

49 ATHABASCENSIS HERMANNTF.J.v LEAFL. W. BOT. 8:111. 1957. CANADA: ALBERTA: JASPER NATIONAL PARK, ATHABASCA RIVER, ATHABASCA FALLS ( HERMANNTF. J. T 13498. 28 AUG 1956 1 us 2265958 HOLOTYPE

50. ATHROSTACHYA OCNEYTS.T. IN GRAYvA.9 PROC. AMER. ACAD. ARTS 7:393. 1868. USA: CALIFORNIA: MARIPOSA COO: YOSEMITE NATIONAL PARK, YOSEYITE VALLEY (BOLANDERTH~N~T6213. 17 JUN 18631 NY SY NTY PE us 319165 SYNTYPE USA: CALIFORNIA: MARIPOSA COO: YOSEMITE NATIONAL PARK, YOSEYITE VALLEY (BREUER,W.H., 1650. 17 JUN 1863) MO SYNTYP E NY SYNTYPE

51. ATRACTOOES HERMANNTF. J.9 J. WASH. ACAD. SCI. 40:283. 1950. MEXICO: CHIAPAS: COMITAN (SHARPTA.J.T 45450. 29 APR 1945) NY ISOTYPE us 2133192 TYPE 40 SMITHSONIAN CONTRIBUTIONS TO BOTANY

520 ATROFUSCA VAR. DECOLORATA P0RSILDtA.E.t SARGENTIA 4:20. 1943. CANADA: NORTHWEST TERRITORIES: MACKENZ IE DISTRICT: GREAT BEAR LAKE9 CAPE MCDONNELL (P0RSILDtA.E. AND PORSILDt ROT., 5120. 02 AUG 1928) us 2096188 ISOTYPE

53. ATROSQUAUA MACKENZ1EtKoK.t PROC. BIOL. SOC. WASH. 25:51. 1912. CANADA: ALBERTA: SMOKY RIVER (H0LLISTERqN.t 14. 05 AUG 1911) NY ISOTYP E us 622651 HOLOTY PE

54 0 AUREA VAR, ANDROGYNA OLNEY9S.T. IN WATS0NtS.t BOT. U.S. GEOL. EXPLOR. 40TH PAR. 371. 1871. USA: PENNSYLVANIA: ERIE COO: ERIE9 PRESQUE ISLE (PENINSULA) (GAR6ERtAoP.t ---o 09 JUN 1869) NY ISOTYPE

55 0 AUROLENSIS STEUDEL, E.G.9 SYN. PL. GLUY. 2:223. 1855. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMM0NDtT.t 431. -s. --- 1832 1 NY TYPE COLLECTION

56 AUSTRO-CAROLINXANA BA1LEYtL.H.t BULL. TORREY BOT. CLUB 20:428. 1893. USA: SOUTH CAROLINA: PICKENS COO: TABLE MOUNTAIN (6UCKLEYtS.B. t __-. ---I YO 1834152 HOLOTYPE

57, AUSTROMONfANA PAR1SHtS.B.r BULL. So CALIF. ACAD. SCI. 4:108t PL.15. 1905 USA: CALIFORNIA: SAN BERNARDINO CO.: MILL CREEK FALLS; ALTO 6000 FT. (PARISHtS.B*t 2485. 03 JUL 1892) DS 489410 HOLOTYPE

58, AUTUUNALIS MACKENZIE,K.K. 9 No AMER. FL. 18:66. 1931. MEXICO: MEXICO: FLOR DE MARIA (PR1NGLEtC.G.t 4275. 03 OCT 1892) F 264169 ISOTYPE YO ISOTYPE us 817295 HOLOTYPE

59. AZTECICA MACKENZ1EtK.K.t No AMER. FL. 18:229. 1935. MEXICO: OAXACA! SIERRA DE SAN FELIPE; ALTO 2300 M. (PR1VGLEtC.G.t 4839. 19 AUG 1894) GH TYPE COLLECTION YO TYPE COLLECTION

60- AZUAYAE STEYERMARK, J.A. 9 PHYTOLOGIA 9:337. 1964. ECUADOR: AZUAY: TOREADOR (STEYERMARK9J.A.t 53105. 15 JUN 1943) F 1266184 TYPE MATERIAL NY ISOTYPE us 1933437 ISOTYPE NUMBER 12 41

- B-

61 BACKANA DEWEYTC., AMER. J. SCI. ARTS SER.19 29:250. 1836. CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51'N.r 106 13'W.l (RICHARDSONTJIT 417. ---I GH TYPE COLLECTION

62 BACKII BOOTTTF. IN HOOKER,W.J., FL. BORorAYER. 2:210. 1839 ( "1840") CANADA: SASKATCHEWAN: CARLTON HOUSE ( 52 51". q 106 13'W. 1 (RICHAROSONqJ.9 --- . ---I GH SYNTYPE NY SYNTYPE

63 BALTZELLII CH4PMANqA.W. EX DEWEYtC., AMER. J. SCI. ARTS SER.2, 3:335. 1847. USA: FLORIDA: BEAR CREEK (CHAPMAN PA. We 9 --- . ---I NY COTY P E

64 BAHBUSETORUH MERR1LLqE.D.r PHILIPP. J. SCI. 13:132. 1918. CHINAt KWANGTUNG: LOH-FAU-SHAN (MOUNTAIN) (MERRILLqE.D., 10985. 12 AUG 1917) us 2333748 ISOTYPE

65 BANKS11 BOOTTqF.9 TRANS. LINN. SOC. LONDON 20:119. 1846. ARGENTINA: T IERSA DEL FUEGO (TERRITORY) : TIERRA DEL FUEGO (BANKSqJ. AND SOLANDERTD.C.T --- . -- --- 1769) YO 1611724 SYNTYPE us 1232938 SYNTYPE

66 BARBARA€ DEWEYTC. IN TORREYqJ. IN EYORY,W.H. T REP. U.S. MEX. BOUND. SURV.9 BOT. 2(1):231. 1859. USA: CALIFORNIA: LOS ANGELES COO : SANTA BARBARA (PARRYqCoC. T -_-. -- --- 1850 1 GH HOLOTY PE NY I SOTYPE

67 BARRATTII SCHWEINIT2,L.D. AND TORREYqJ. T ANN. LYCEUM NATO HIST. NEW YORK 1:361. 1824. USA: NEW JERSEY: CAPE MAY COO: CAPE MAY (COLLINSTZ.T --- . ---I NY TYPE COLLECTION

680 BARTLETTII O'NEILLTH.T.T PUBL. CARNEGIE INST. WASH. 522:255. 1940. BRITISH HONDURAS: CAYO: MOUNT PINE RIDGE (BARTLETTTH.H.T 11718A. 24 FEB 1931) F 999642 TYPE MATERIAL GH ISOTYPE NY TYPE

690 BAYARDI FERNALD1M.L .q RHODORA 44371. 1942. 42 SMITHSONIAN CONTRIBUTIONS TO BOTANY

USA: VIRGINIA: SOUTHAMPTON COO: DREWRYVILLE (FERNALD,M.L.; LONGTB. AND SMART,R.F.p 5677. 22 JUN 1936) GH H01OTY PE

70. BILTHOREANA MACKENZIE,K.K. T BULL. TORREY BOT. CLUB 37:234. 1910. USA: NORTH CAROLINA: SATULA MOUNTAIN (---T 2688. 25 MAY 1897) GH ISOTYP E YY TYPE

71 BIPARTfTA VAR. AUSTROMONTANA HERMANNTF.J.T LEAFL. W. BOT. 10:16. 1963. USA: COLORADO: BOULDER COO: NEDERLANDV NOWIT RIDGE; ALTO 11500 FT. (HERMANNIF~J~T 17059. 15 AUG 1961) CAS 430881 ISOTYPE

72. BONANZENSIS BRITTON,N.L.9 BULL. NEW YORK BOT. GARD. 2:160. 1901. CANADA: YUKON TERRITORY: BONANZA RIVER ( WILLIAMStR. So T ---. 18 JUN 1899) NY TYPE 13. BONPLAND!! VAR. MINOR BOOTTTF. IN GRAYTA.~ PROC. ACAD. NATO SCI PHILADELPHIA 1863:77. 1863. USA: COLORADO: ROCKY MOUNTAINS; LAT. 39-41 No (HALLqE. AND HARBOURTJ.P.T 591. -- --- 1862) F 314869 ISOTYPE F 456934 I SOTYPE GH HOLOTYPE YO ISOTYPE

140 BRACHYPODA HOLMTH.T.T AMER. J. SCI. SER.4, 20:302. 1905. USA: OREGON: KLAMATH C0.t CRATER LAKE NATIONAL PARK, CATHEDRAL SPRING (COVILLETFOV-T 1455. 11 SEP 1962) us 415269 TYPE COLLECTION

75 BRAINERDII MACKENZIE,K.K. T BULL. TORREY BOT. CLUB 40:534. 1913. USA: CALIFORNIA: EL DORADO COO: SIERRA NEVADA RANGE, SLIPPERY FORD (BRAINERDTE., 1210 19 JUL 1897) us 964504 TYPE COLLECTION

76 BREVrCAULIS YACKENZIETK.K.T BULL. TORREY BOT. CLUB 40:547. 1913. USA: OREGON: LINCOLN C0.2 YAQUINA BAY (HDWELLTTOJO~2994. -- MAY 1886) h(Y TYPE COLLECTION

77. BREVIS BLAKEfS.T.9 J. ARNOLD ARBOR. 28:lll. 1947. PAPUA 4ND NEW GUINEA: PAPUA (TERRITORY): OWEN STANLEY RANGEt MOUNT ALBERT EDWARD: (COUNTRY AS "BRITISH NEW GUINEA") (BRASS,L.J., 4418. -- MAY-JUL 1933) A ISOTYPE

78. BREVISQUAMA MACKENZIETK.K.T BULL. TORREY BOT. CLUB 34:152. 19070 USA: WYOMING: SWEETWATER COO: RED DESERT9 ORENDO BUTTE (NELSON9A.q 7124. 11 JUN 1900) NUMBER 12 43

NY HOLOTYPE

19 BREWER1 B00fT~F.q ILL. GENUS CAREX 4:142~ PL.455. 1867. USA: CALIFORNIA: SISKIYOU COO: MOUNT SHASTA (BREWER,W.H.* 1422. -- --- 1863) GH I SOTYPE

80 0 BRONGNIARTI! VARo OENSA BAILEY,L.H., PROC. AMER. ACAD. ARTS 22:137. 1886 ( "1887") USA: CALIFORNIA: MARK WEST CREEK (BIGELOW,J.M., ---. -- --- 18 53 -1 854 1 NY SYNTYPE

810 BRUNNEA VARo SUBTEIOGYNA KUKENTHALTG. T REPERT. SPo NOV. REGNI VEG. 8:8. 1910. PHILIPPINES: BENGUET: LUZON (ISLAND), MOUNT PULOG (MERRILL,E.D.q 6505. -- MAY 1909) us 711129 TYPE

82. BUCKLEVI DEWEYqC., AMER. J. SCI. ARTS SER.1, 482143. 1845. USA: NORTH CAROLINA: MITCHELL COO: ROAN MOUNTAIN (BUCKLEY,S.B., --- . ---I NY TYPE COLLECTION

83 BULBOSTYLIS YACKENZIE9K.K. T BULL. TORREY BOT. CLUB 42:617. 1915. USA: TEXAS: TARRANT COO: FORT WCIRTH (RUTHqA., 360. 12 APR 1913) YO 7101 12 TYPE COLL ECT ION us 587668 TYPE COLLECTION

BURCHELLIANA BOECKELERqJ.O., LINNAEA 412234. 1877. SOUTH AFRICA: --:-- (BURCHELLqW.J.9 1911. ---I GH I SOTYPE

BUSH11 MACKENZ1EqK.K. 9 BULL. TORREY BOT. CLUB 37:241. 1910. USA: AQKANSAS: HEYPSTEAD COO: FULTON (BUSHqBeF. T 2514. 30 APR 1905) NY TYPE COLLECTION

- C-

86 CAESARIENSIS YACKENZ1EtKoK.q No AYER. FL. 18:440. 1935. USA: NEW JERSEY: CAMDEN COO: LAUREL SPRINGS (LONGTB.~ F23212. 15 JUN 1920) GH 1 SOTY PE NY TYPE

CALIFORNICA BA1LEYqL.H.q MEMO TORREY BOT. CLUB 1:9. 1889. USA: CALIFORNIA: MENOOCINO COO: YENDOCINO CITY (BOLANDER9H.N. 9 4741. 01 MAY 1866) CAS 383776 TYPE COLLECTION 44 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DS 49734 TYPE COLLECTION YO TYPE COLLECTION NY TYPE COLLECTION us 29741 TYPE COLLECT1ON us 319268 TYPE COLLECTION

88 CAMPLYOCARPA HOLM,H.T., AMER. J. SCI. SER.4, 20:304. 1905. USA: OREGON: KLAMATH COO: CRATER LAKE NATIONAL PARK, CATHEDRAL SPRING (COVILLE,F.V.q 1457. 11 SEP 1902) us 690937 TYPE COLLECTION

89. CAMPYLOCARPA SSPo AFFlNIS MAGUIRETB. AND HOLMGREN,A.H.( LEAFL. W. BOT. 42262. 1946. USA: UTAH: JUAR COO: DEEP CREEK RANGE, INDIAN FARM CREEK (MAGUIREvB. AND HOLMGREN9A.H. t 21947. 16 JUL 1943) CAS 334353 ISOTYPE NY HOLOTY PE us 1885701 I SOTYPE

90 CANESCENS VAR. DISJUNCTA FERNALD,Y.L. 9 PROC. AMER. ACAD. ARTS 37:4889 PL.5. 1902. CANADA: NEW RRUNSWICK: VICTORIA CO. : SERPENTINE RIVER (HAYIG~U.~ 84. 24 JUL 1900) GH PARATY PE

91 CANESCENS VAR. DUBIA BAILEYIL~H. t BOT. GAZ. 9:119. 1884. USA: UTAH: BEAR RIVER CANYON; ALTO 10000 FT. ( WATSONvS. T 1231A. -- AUG 18691 NY ISUTYPE

92 CANESCENS VARo SPHAEROSTACHYA TUCKERMAN,E., ENUM. CARIC. 19. 1843. USA: --: NEW ENGLAND (---, --- . -- --- 1843) GH I SOTYPE NY TYPE COLLECTION

93 (. CAREYANA TORREYvJ. EX DEWEY,C.r AYER. J. SCI. ARTS SER.1, 30:60. 1836. USA: NEW YORK: CAYUGA COO: AUBURN (CAREY,J.* --- -- MAY 1832) NY HOLOTY P E

94 0 CAROLINIANA BUCKLEY,S.B.v AMER. J. SCI. ARTS SER.1, 45:173. 1843. USA: SDUTH CAROLINA: PICKENS COO: TABLE MOUNTAIN (BUCKLEY,S.B., _--. ---I GH TY PE C OLL ECT ION NY TYPE COLLECTION

95 0 CEPHALOPHORA VAR. MAXIMA DEWEYqC., AMER. J. SCI. ARTS SER.19 43:92. 1842. USA: NEW YORK: YATES COO: PENN YAN (SARTWELL,H.Pe, --- . ---I GH TYPE

96 CHALCIOLEPIS HOLM,H.T.( AMER. J. SCI. SER.4, 16:21,28. 1903. USA: COLORADO: MINERAL COO : PAGOS A PEAK BAKER, C OF 9 226. NUMBER 12 45

-- AUG 1899) GH SYNTYPE YO SYNTYPE NY SYNTYPE us 368814 SYNTYPE 97. CHAPMAN1 SARTWELL9H.P. EX DEWEYqC.9 AMER. J. SCI. ARTS SER.29 19:254. 1855. USA: FLORIDA: *- (CHAPMAN,A.W., 113. ---I CAS 553918 ISOTYPE us 28433 ISOTYPE 98. CHIAPENSIS HERMANNTF.J.T BRITTONIA 19:68. 1967. MEXICO: CHIAPAS: CHAYULA (BREEDLOVE,D.E. 9 6714. 30 JUL 1964) F 1620435 ISOTYPE NY ISOTYPE us 2460272 HOLOTYPE

99. CHIHUAHUAENSIS MACKENZIE,K.K., BULL. TURREY BOT. CLUB 35:265. 1908. MEXICO: CHIHUAHUA: PUERTA DE ST. DIEGO; ALTO 6500 FT. (HARTMANTC-V., 620. 12 APR 1891) F 49642 ISOTYPE NY HOLOTYPE us 306281 ISOTYPE 100. CHIKUNGANA BAILEY,L.H., GENTES HERB. 1:13. 1920. CHINA: HUPEH AND YONAN: CHIKUNGSHAN ( BAILEYTL.H.T ---. 13 JUN 1917) NY TYPE

101. CILIARIS FERNALD9M.L. 9 PROC. AMER. ACAD. ARTS 43:61. 1907. MEXICO: HIDALGO: LENA STATION (PRINGLETC~G~T10039. 26 AUG 1905) CAS 232050 ISOTYPE CAS 445943 ISOTYPE F 202021 ISOTYPE GH HOLOTYPE YO ISOTYP E hsY ISOTYPE us 462090 ISOTYPE 102- CINNAMOMEA 0LNEYqS.T. IN GRAYTA~TPROC. AMER. ACAD. ARTS 7:396. 1868. USA: CALIFORNIA: HUMBOLDT COO: RED MOUNTAIN (BOLANDERTH.N.~ 6477. -- --- 1866 I CAS 553874 TYPE FRAGMENT GH TYPE COLLECTION us 28457 TYPE COLLECTION us 319228 TYPE COLLECTION

103. CIRCINNATA MEYERTC-A.~ MEMO ACAD. IMP SCI ST.-PETERSBOURG DIVERS SAVANS 1 :209~PL.6. 46 SMITHSONIAN CONTRIBUTIONS TO BOTANY

1831. USA: ALASKA: ALEUTIAN ISLANDS9 UNALASKA 4 ISLAND) ( CHA Y IS S 0 9 L A 9 --- . ---I GH ISOTYPE

1040 CLADOSTACHYA VAR. MAXIMA KUKENTHAL 9 Go IN ENGLER, H .Gm Ao 9 PFL AN ZENR 49 FAM.20:268. 1909. BOLIVIA: --: -- (BANG9M.t 2210. ---I us 350077 TYPE COLLECTION

1050 CLIVICOLA FERNALD9M.L. AND WEATHERBY,C.A. 9 RHODORA 33:233. 1931 CANADA: QUEBEC: GASPE COO: MOUNT SAINT PIERRE (FERNALD9M.L. t WEATHERBYqCoA. AND STEBBINS9G.L.r 24110 05 JUL 19311 GH HOLO TYP E us 1839933 ISOTYPE

106. COLLECTA DEWEY9C.y AMER. J. SCI. ARTS SER.19 11:314. 1826. USA: MASSACHUSETTS: HAMPSHT RE COO : WORTHINGTON (DEWEYpC.9 ---. --- ) GH HOLOTYPE

107- COLUMBIANA DEWEY9C.p AMER. J. SCI. ARTS SER.1, 30:62. 1836. USA: --:COLUMBIA RIVER (SCOULER9J.9 --- . ---I NY HOLOTYPE

168. COMANS VARo STRICTA CHEESEMANrToF.9 TRANS. & PROC. NEW ZEALAND INST. 242415. 18920 NEW ZEALAWD: CANTERBURY (DISTRICT): SOUTH ISLAND9 LAKE TEKAPOi ALL 2sor FT. (CHEESEMAN~LF.~ --- 0 -- JAN 1883) us 2038822 TYPE COLLECTION

COHMUNIS BAILEY9L.H. 9 MEMO TORREY BOT. CLUB 1:41 1889. USA: NEW YORK: YATES COO: PENN VAN (SARTWELL,H.P.9 108. -- --- 1848) CAS 553913 SYNTYPE

1100 CONClNNOIDES MACKENZIEqKoK.9 BULL. TORREY BOT. CLUB 33:440. 1906. USA: MONTANA: FLATHEAD COO: COLUMBIA FALLS (WILLIAMS9R.S.9 ---. 07 JUN 1893) NY TYPE

1110 CONFERTIFLORA BOOTT9F. IN GRAYpA.9 MEMO AMER. ACAD. ARTS NOS., 6:418. 1859. JAPAN: HOKKAIDO (PREFECTURE) : HAKODATE (WRIGHTqC.9 ---. 0- JUN 1855) us 27235 TYPE MATERIAL

1120 CONJUNCTA BOOTT9F.t ILL. GENUS CAREX 3:1229 PL.392. 1862. USA: OHIO: FRANKLIN COO: COLUMBUS (SULLIVANT9W.S. 9 --- . ---I C AS 383550 SYNTYPE GH SY NTYPE

113. CONSPECTA YACKENZIE9KoK.9 No AMER. FL. 182294. 1935. NUMBER 12 47

MEX ICO: PUEBLA: PUEBLA ( ARSENE9 Go ( FRERE) 9 1359. 01 AUG 1907) us 1032323 HOLOTYPE

1140 CONSTANCEANA STACEYTJ.W.T LEAFL. W. BOT. 2:123. 1938. USA: W4SHINGTON: YAKIMA COO: MOUNT ADAMS ("PADDO") 9 WODEN VALLEY (SUKSDORFtW.N.9 6864. 16 AUG 1909) CAS 242987 HOLOTYPE DS 269649 ISOTYPE NY I S OTY P E

CONVOLUTA YACKENZIE9K.K. 9 BULL. TORREY BOT. CLUB 43:428. 1916. USA: NEW JERSEY: MORRIS COO: BUDD'S LAKE (MACKENZIE9K.K.9 20880, 10 JUN 19C6) NY TYPE

COOLEYI DEWEYtC. IN WOODtA., AMER. J. SCI, ARTS SER.19 48:144. 1845 USA: MICHIGAN: MACOMB COO: WASHINGTON (COOLEYtD.9 ------I GH HOLOTYPE

COSTATA SCHWEINITZ~L.D.T ANN. LYCEUM NATO HIST. NEW YORK 1:67. 1824. USA: PENNSYLVANIA: NORTHAMPTON COO: EASTON (SCHWEINITZTL.D.T ------I NY TYPE COLLECT ION

CRANDALLII GANDOGERtM. 9 BULL. SOC. BOT. FRANCE 66:295. 1920. USA: COLORADO: SUMMIT COO: GRAYS PEAK (JONEStM.E., 834. 28 AUG 1878) NY TYPE COLLECTION

CRAWFORDII FERNALDtM.L.9 PROC. AMER. ACAD. ARTS 37:469* PL.1. 19C2. USA: NEW HAMPSHIRE: COOS C0.z MOUNT WASHINGTON, BETWEEN YARSHFIELD AND CRAWFORDS (FAXONvE. AND FAXON9C.E. 9 ---. 06 JUL 1878) GH SYNTYPE

12Qo CRAWFORDII VARo VIGENS FERNALD9M.L. t PROC. AMER. ACAD. ARTS 37:470~PL.1. 1902. CANADA: QUEBEC: GASPE COO: EAST GASPE (MACOUNTJOHN, 6. 01 AUG 1882) GH SY NTYP E

121, CREBRIFLORA WIEGAND9K.M.9 RHODORA 24:197. 1922. USA: FLORIDA: GADSDEN COO : APPALACHICOLA RIVER9 CHATTAHOOCHEE (CURTISStA.H., 3267. -- SEP 1882) F 26304 TYPE MATERIAL GH TYPE NY TYPE

1220 CRINITA VARo BREVICRINIS FERNALDTM.L.T RHODORA 48:54* 1946. USA: VIRGINIA: DINWIDDIE COO: ROWANTA (FERNALD9M.L. AND 48 SMITHSONIAN CONTRIBUTIONS TO BOTANY

LONGTB.T 8143. 08 JUN 1938) GH HOLOTY PE us 278555 ISOTYPE

CRINITA VARo M!NOR BOOTTTF.T ILL. GENUS CAREX 1:18. 1858. USA: NEW YORK: YATES COO: PENN VAN (SARTWELL,H.P*, 78. ---I CAS 553883 TYPE COLLECTION

CRINITA VAR. SIMUCANS FERNALD9Y.L. T PROC. PORTLAND SOC. NATO HIST. 2:135. 1897, USA: MAINE: PISCATAQUIS COO: GREENVILLE (FERNALD,M.L.p 264. 04 JUL 1894) GH SYNTYP E us 278555 SYNTYPE

1250 CRISTATA SCHWEINITZTL.~. T ANN. LYCEUM NAT. HIST. NEW YORK 1:66. 1824. USA: NEW JERSEY: -- (---T --- . ---I GH ISOTY P E

126. CRUS-CORVI SHUTTLEWORTHTR-J. EX KUNZETG.~ SUPPL. SCHKUHR'S RIEDGR. 1281 PL.32. 1844. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMONDTT., 432. -- --_ 1832 1 GH ISOTYPE

CRUS-CORVI VAR. VIRGINIANA FERNALDTY.L.T RHODORA 39:3939 PL.476. 1937. USA: VIRGINIA: SOUTHAMPTON COO: DREWRYVILLE (FERNALDvM. Lo LONGTB. AND SMART,R.F., 5677. 22-23 JUN 1936) GH HOLOTYPE YO 1108572 ISOTYPE NY ISOTY P E us 1682487 ISOTYPE

CRYPTOLEPIS YACKENZIEtKoK., TORREYA 14: 157. 1914. USA: NEW JERSEY: SUSSEX COO: WHITE POND (MACKENZIE,K.K., 4645. 26 JUN 1910) NY TYPE

CUBENSIS KUKENTHAL, Go9 REPERT SP. NOY. REGNI VEG. 23:220. 1926. CUBA: ORIENTE: PIC0 TURGUINO (EKMANqE*L., 14506. 21 JUL 1922) NY ISOTYPE us 1302602 TYPE COLLECTION

CUBENSIS VARo FLACCIDA KUKENTHAL9G.9 REPERT. SP. NOV. REGNI VEG- 23: 221. 1926. HAITI: --:-- (EKMANvE.L., --- 08 AUG 1925) NY TYPE COLLECT ION

CUCHUMATANENSIS STANDLEY,P.C. AND STEYERMARKIJ.A.T CEIBA 4:62. 1953. GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUYATANEST TUNIMA; NUMBER 12 49

ALTO 3400-3500 Me ( STEYERMARK9J.A. 9 48347. 07 JUL 1942) F 1128952 HOLOTYPE

132- CULHENICOLA STEYERYARKTJ.A., FIELDIANA? BOT. 28:65, FIG.7. 1951. VENEZUELA: SUCRE: CERRO fURUMIQUIRE9 EASTERN PEAK; ALTO 2500 Me (STEYERYARKIJ~A.~ 62605. 06 YAY 1945) F 1266170 HOLOTY PE

133. CUHULATA FOR. SOLUTA FERNALDtMoLmr RHODORA 44:285. 1942. CANADA: NOVA SCOTIA: QUEENS COO: BROAD RIVER (FERNALD9M.L. AND BISSELLtC.H.? 20311. 16 AUG 1920) GH HOLOTY PE

134, CUNEATA OHWIIJ., MEMO COLL- SCI. KYOTO IMP. UNIV.9 SERoB9 BIOL. 6:256. 1931. JAPAN: AOYORI (PREFECTURE) : HONSHU ( ISLAND) 9 AOMORI (KINASHI9N.t --- -- JUL 1909) F 1406416 TYPE MATERIAL

135- CURATORIUM STACEYtJoW., LEAFL. We BOT. 2:13. 1937. USA: ARIZONA: COCONINO COO: GRAND CANYON NATIONAL PARK? KAIBAB TRAIL TO ROARING SPRINGS (EASTWOODvA. AND HOWELL9J.T. 9 1101. 23 JUN 1933) CAS 204973 SYNTYPE CAS 204974 SYNTYPE

136. CUSICKIh MACKENZIE9K.K. IN PIPER9C.V. AND BEATTIE,R.K.r FL. NW. COAST 72.. 1915. USA: OREGON: BAKER COO: HEAD OF BURNT RIVER (CUSICKqW.C.9 1331. -- JUL 1886) NY SYNTY P E

-D-

137. OANAENSIS STACEY?J.W., LEAFL. W. BOT. 2:166. 1939. USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (HOWELL9J.T., 14546. 11 AUG 1938) CAS 259874 ISOTYPE CAS 2 59875 HOLOTY PE GH ISOTYPE us 1765700 ISOTY PE

138- DAVYI YACKENZIE 9K.K. 9 BULL. TORREY BOT. CLUB 43:606. 1916. USA: CALIFORNIA: PLACER COO: TRUCKEE RIVER (BURT-DAVY?J.r 3266. 25-30 JUN 1897) GH ISOTYPE JEPS 2511 ISOTYPE NY ISOT YP E uc 50814 HOLOTYPE

139. X DEAHII HERMANNtFmJ. p RHODORA 40:81. 1938. 50 SMITHSONIAN CONTRIBUTIONS TO BOTANY

USA: INDIANA: PIKE CO.: OTWELL (HERMANN9F.J.9 6147. 05 JUL 1934) F 751055 ISOTYPE

DEBILIFORMIS MACKENZIETK.K.T BULL. TORREY BOT. CLUB 37:244. 1910. USA: CALIFORNIA: MENDOCINO COO: -- (BOLANDERTH.N-T 6477. -- -_- 1866) YO TYPE MATERIAL

OEBILIS VARo INTERCURSA FERNALDTM.L*T RHODORA 44:3@79 PL.713. 1942. USA: VIRGINIA: GREENSVILLE CO.: ORION (FERNALDpM-L. AND LONG9B.9 12016. 13 JUN 1940) GH HOLOTY PE us 2003164 ISOTYPE

OEBfLIS VAR. PUBERA GRAYfA.9 MAN. BOT. ED059 593. 1867. USA: PENNSYLVANIA: CENTRE CO.: BEAR MEADOWS (PORTERfT.C.9 ---* --- ) GH HOLOTY PE

DEVIA CHEESEYANTT*F*T TRANS. C PROC. NEW ZEALAND INST. 15:301. 1883. NEW ZEALAND: NELSON (DISTRICT) : SOUTH ISLAND? NELSON (CHEESEMAN9T.F.9 83. -- JAN 1882) GH ISOTYPE

144. DEWEYANA VARo COLLECTANEA FERNALD9M.L. 9 RHODORA 15 :93* 1913. CANADA: QUEBEC: BONAVENTURE CO. : CASCAPEDIA RIVER9 GRAND CASCAPEOIA (W1LLIAMStE.F.; COLLINS9J.F. AND FERNALD9M.L.r --- 12-15 JUL 1905) GH HO LO TY P E

OEHEVANA VARo SPARSIFCORA OLNEYpS-T. EX BAILEY,L.H.r BOT. GAZ. 13:87* 1888. USA: OREGON: MARION CO.: SALEM (HALL9E.9 580. -- --7 1871) F 455703 TYPE COLLECT ION F 1429766 TYPE COLLECTION GH TYPE COLLECT ION NY TYPE COLLECTION

1460 DIGITALIS VARo ASYNMETRICA FERNALDfM.L.9 RHODORA 43:544. 1941. USA: VIRGINIA: SOUTHAMPTON CO. : APPLEWHITE CHURCH (FERNALD9M.L. AND LONGpB.9 11791. 08 MAY 1940) CA S 336835 I SOTYPE GH HOLOTYP E YO 1306423 ISOTYPE us 2003133 I SOTYPE

DIGITALIS VAR. GLAUCA CHAPMANTA.W.T FL. S. U.S. ED.19 541. 1860. USA: FLORIDA: MIDDLE FLORIDA (CHAPMANTA*W.T ---a -- --- 1842 1 NY TYPE COLLECTION us 969118 TYPE COLLECTION NUMBER 12 51

DIGITALIS VARo MACROPODA FERNALDtM*L.-t RHOOORA 40:400t PL-511- 1938- USA: VIRGINIA: GREENSVILLE COm: -- (FERNAL0tM.L- AND LONGtB-e 7767. 08 APR 19381 GH HOLOTYPE YO 1129747 ISOTYPE NY I SOTY PE us 1761151 ISOTYPE

149- DIVERSISTYLIS ROACHtAa We 9 MADRONO 112277. 1952. USA: OREGON: LIYN CO.: CLEAR LAKE JUNCTION (R0ACHtA.W-9 202- 10 JUN 1949) CAS 372834 I SOTYPE 150- DONNELL-SMITH11 BAILEYvL~H.~MEY. TORREY BOT. CLUB 1:56. 1889. GUATEMALA: ALTA VERAPAZ: PANSAMALA; ALT- 3800 FT. (SMITH, J.0. AND TURCKHEIMtHa, 659- -- JUN 1885) us 817314 TYPE COLLECTION

151. DOUGLAS11 VAR. DENSISPICATA DEWEYtCat AMERm J. SCI. ARTS SER-27 32:41e 1861- USA: NEBRASKA: -- (HAYDENPF-V-T 580. ---I GH TYPE MATER IAL

1520 DUDLEY1 MACKENZIE 9K-K. v ERYTHEA 8230. 1922. USA: CALIFORNIA: MONTEREY CO.: TASSAJARA HOT SPRINGS (ELMERtA-DeE-r 3132- -0 JUN 1901) 0s 145619 HOLOTYPE DS 629609 I SOTYPE YO I SOTYP E YY I SOTYPE

153- X OUMANXI LEPAGEtE., NATURALISTE CANAD. 83:143, FIG-4- 1956. CANADA: QUEBEC: VIEUX-COMPTOIR (LEPAGEvE. 9 32078. 30 JUL 1954) GH ISOTYPE us 2176489 I SOTYPE

154. DURANDXI BOECKELER, J.0-t ALLGm BOT- Z. SYST. 2 :189- 1896. COSTA RICA: --: CERRO OE BUENA VISTA (PITTIERpH- AND TONDUZTA-, 3376. 19 JAN 18911 CAS 351155 ISOTYPE us 579795 TYPE MATERIAL

1550 DURIFOLIA BA1LEYtLeH.t BULL. TORREY BOT. CLUB 20:428. 1893- CANADA: SASKATCHEHAN: CARLTON HOUSE 4 52 51"- t 106 13'W. 1

(RICHARDSON, J 9 -0- -0-1 NY SY NTY P E

156. DUTILLY! 0'NEILLtH.T- AND OUMANtM-t RHODORA 43:413t PL.669- 1941. CANADA: MANITOBA: CHURCHILL RIVER, CHURCHILL (DUMAN,Y-, 1506. 08 AUG 1938) GH ISOTYPE 52 SMITHSONIAN CONTRIBUTIONS TO BOTANY

- E-

151. EASTWOOOIANA STACEYTJ.W.T LEAFL. W. BOT. 2:121. 1938. USA: OREGON: GRANT CO.: DIXIE MOUNTAIN (HENDERSONTL.F.T 5583. 25 JUL 1925) CAS 1363 86 HOLOTY PE DS 144009 ISOTYPE GH ISOTYP E

158, EBENEA RYDBERGTP.A. 9 BULL. TORREY BOT. CLUB 28: 266. 1901. USA: COLORADO: EL PAS0 COO: PIKES PEAK (CLEMENTStF.9 ---. _- -_- 1906 I NY TYPE

159. ECHINATA VAR. ORMANTHA FERNALDt M.L. 9 PROC. AMER. ACAD. ARTS 37:483* PL.4. 1902. USA: CALIFORNIA: EL DORADO COO: SIERRA NEVADA RANGE, STRAWBERRY CREEK (BRAINERDTE., 160. 18 JUC 18971 GH HD LO TYP E

160, EGGERTII BAILEYTLOH~TBOT. GAZ. 21:6. 1896. USA: MISSOURI: BUTLER COO: -- (EGGERT9H.q --- 08 AUG 1893) NY TYPE COLLECTION

161. EGGLESTONlI YACKENZIETK~K-TBULL. TORREY BOT. CLUB 422614. 1915. USA: COLORADO: GUNNISON C0.2 MOUNT CARBON, KEBLER PASS (EGGLESTON,W.W.q 6181. 22 AUG 191C) NY ISOTYP E us 857864 TYPE

162 w EGGLESTONII VARw FESTIVELLIFORMIS HERMANN,F.J.r BRITTONIA 12:78. 1960. MEXICO: NUEVO LEON: GALEANA 4 SCHNEIDER,R.A. 9 954. 25 JUL 1938) us 2466328 HOLOTYPE

163 w EGREGIA MACKENZIETK~K~TBULL. TORREY BOT. CLUB 42:414. 1915. USA: WASHINGTON: KLICKITAT COO: FALCON VALLEY (SUKSDORFTW.N.T 5181. 15 JUL 19051 DS 284598 ISOTYPE NY TYPE

164. EKHANII KUKENTHALTG., REPERT. SP. NOV. REGNI VEG. 23:221. 1926. HAIT I: OUEST: PET IONVILLE ( EKMANT E.L. 7 H1453. 12 AUG 1924) GH ISOTYPE NY TYPE YATERIAL us 1411790 COTYPE

165. EKMANIl VAR. HOTTENSIS KUKENTHALvG. AND EKMANTE.L.T ARK. BOT. 22A( 17) :9. 1929. HAITI: .--: MORNE CALUMETTE; ALTO 1200-1300 Me (EKMAN9E.L. T NUMBER 12

H10662. 14 SEP 19281 GH I SOTYPE us 1414090 TYPE COLLECTION

166- ELBERTANA KELSOvL.9 BIOL. LEAFL. 31~3. 1945. USA: COLORADO: LAKE COO: MOUNT ELBERT (KELSOIL~T 4967. C1 AUG 1945) GH TYPE MATES1 AL

1670 ELEOCHARLS BAILEY,L.H.t MEM. TORREY 601. CLUB 1:6. 1889. CANADA: SASKATCHEWAN: SASKATCHEWAN PLAINS ( YACOUN~JOHNI 1665. 12 AUG 1872) GH I SOTYPE

168. ELMER1 KUKENTHAL,G., REPERT. SP. NOV. REGNI VEG. 8:326. 1910. PHILIPPINES: BENGUET: LUZON (ISLANDIT BAGUIO (ELMERqA.D.E.1 8444. -- MAR 1907) YO TYPE MATERIAL us 854950 TYPE MATERIAL

1690 ELROOI JONES9M.E.q BULL. MONTANA STATE UN1V.r BIOL. SER. 15:70. 1910. USA: MONTANA: BEAVERHEAD COO: YONIDA (J0NESvM.E. 7 ---a 08 JUL 1909) DS 149706 I SOTYPE NY I SOTYP E us 1531248 TYPE MATERIAL 170. ELYNOIDES HOLM,H.T.T AMER. J. SCI. SER.4, 9:356. 1900. USA: COLORADO: MIN'ERAL COO: PAGOSA PEAK; ALT. 12000 FT. (BAKER,C.F., 230. -- AUG 1899) GH I SOTYPE YO TYPE COLLECTION us 368818 TYPE COLLECTION

171. ENGELMANNL BAILEY,L.H.9 PROC. AMER. ACAD. ARTS 22:132. 1886 ("1887") USA: COLORADO: EL PAS0 COO: MOUNT FLORA9 "PROBABLY NEAR COLORADO SPRINGS" ( ENGELYANNqG. --- . -- --- 1874) GH HOLOTYPE

1720 EPAPILLOSA M4CKENZIE,K.K. IN RYDBERGIP~A~TFL. ROCKY MOUNT. 138, 1060. 1917. USA: UTAH: PIUTE COO: MARYSVALE (JONES,M*E., 53450 01 JUN 1874) YO I S OTY PE NY HOLOTY PE hsY I SOTYP E us 270933 ISOTYPE

173. EREMOSTACHYA BLAKEIS-T~T J. ARNOLD ARBOR. 28:99. 1947. INDONESIA: WEST NEW GUINEA: LAKE HABBEYA; (COUNTRY AS ItDUTCH NEW GUINEA") (BRASS9L.J.r 10255. -- OCT 1938) 54 SMITHSONIAN CONTRIBUTIONS TO BOTANY

A ISOTYP E

174. ERXLEBENIANA KELSOtL., BIOL. LEAFL. 5121. 19500 USA: COLORADO: GILPIN COO: ROLLINSVILLE (KELS0tL.v 6362. 24 JUL 1948) GH TYPE MATERIAL

175. EURYCARPA H0LYtH.T.t AMER. J. SCI. SER.49 20:303. 1905. USA: WASHINGTON: KLICKITAT Coo: FALCON VALLEY (SUKSD0RFyW.N.t 1284. 26 JUN 1886) C AS 242957 SYNTYPE

176. EURYCARPA VARm ATTENUATA KUKENTHALtGot REPERT. SP. NOV. REGNI VEG. 26: 254. 1929. USA: WASHINGTON: KLICKITAT COO: FALCON VALLEY (SUKSD0RFtW.N. I 11551. 21 AUG 1924) CAS 246772 TYPE C OLLEC TION

117. EURYSTACHYA HERMANNTF~J~TLEAFL. We BOT. 8:109. 1957. CANADA: ALBERTA: JASPER NATIONAL PARK, MOUNT EDITH CAVELLt CAVELL LAKE (HERMANNtF.J.9 13529. 28 AUG 1956) C AS 401490 ISOTYPE GH ISOTYPE us 2265959 HOLOTYPE

1780 EXPLORATORUH NELMES,E.t BULL. MISC. INFORM. 108. 1938. MALAYSIA: SABAH (TERRITORY 1: MOUNT KINABALU; ALT. 4000 FT.; (COUNTRY AS "BORNEO") (CLEMENStJ. AND CLEMENStMoSot 34297. 28 JUL 1933) GH HOLOTY PE NY I SOTYPE

179. X EXSALINA LEPAGEtEot NATURALISTE CANAD. 83:133. 1956. CANADA: QUEBEC: PIAGOCHIWI RIVER (DUTILLYtA.; LEPAGEtE. AND DUMANtMot 32793. 29 AUG 1954) us 2176495 ISOTYPE

-F-

180. FARGESII FRANCHETtA., BULL. SOC. PHILOM. PARIS SER.8, 7:34. 1895. CHIQA: SZECHWAN: TCHEN-KEOU-TIN (FARGES9R.P. 9 --- . ---I NY TYPE MATERIAL us 1123660 ISOTYPE

181. FELIPENSIS CLARKEtCoBot BULL. MISC. INFORM. ADD.SER.8:84. 1908. MEXICO: OAXACA: SIERRA DE SAN FELIPE; ALTO 10000 FT. (PR1NGLEtC.G.t 4838. 25 AUG 1894) GH TYPE COLLECTION NY TYPE COLL ECT ION

182. FENOLERIANA BOECKELERtJ.O.t LINYAEA 39:135. 1875. NUMBER 12 55

USA: NEW MEXICO: -- (FENDLERqA.9 878. -0 -0- 1847 YO 1816497 TYPE COLLECTION NY TYPE

183, FESTIVA DEWEY,C.T AMER. J. SCI. ARTS SER.1, 29:246. 1836. USA: NORTHWEST TERRITORIES: MACKENZIE DISTRICT: GREAT BEAR LAKE ( "BEAR LAKE") (RICHARDSON, J 9 ------J NY SYNTYPE

184. FESTIVA VAR. DECUHBENS HOLMTH~T.9 AMER. J. SCI. SER.4, 16:20,26. 1903 USA: COLORADO: MINERAL COO: PAGOSA PEAK (BAKERfC.F.9 232. -- AUG 1899) F 122779 TYPE MATERIAL NY TYPE COLLECTION

185. FESTIVA VAR. STRICTA BAILEY,L.H.T MEMO TORREY BOT. CLUB 1:51. 1889 USA: CALIFORNIA: -- (KELLOGGTA. AND HARFORD9W .G.W.T 1073. -- --- 1868- 1869 1 NY TYPE MATERIAL

186, FESTIVELLA MACKENZIE,K.K., BULL. TORREY BOT- CLUB 42:609. 1915. USA: WYOMING: ALBANY CO.: -- (NELSONTA., 3275. 02 JUL 1897) GH ISOTYPE h(Y TYPE

187. FETA BAILEYTL.H.T BULL. TORREY BOT. CLUB 20:417. 1893. USA: CALIFORNIA: SONOMA COO: CLOVERDALE SPRING (BOLANDER,H.N., 50. ---I GH HOLOTY PE

188- FILIFOLIA VARm EROSTRATA KUKENTHALqG. IN ENGLERTH.G.A.v PFLANZENR. 41 FAM.20:86. 1909. USA: CALIFORNIA: EL DORA00 COO: ECHO LAKE (BRAINERDtE.9 111. 11 JUL 1897) GH I SOTYP E

189, FISSA MACKENZ?E,K.K.v No AYER. FL. 18:64. 1931. USA: OKLAHOMA: CREEK COO: SAPULPA (BUSHI 6.F. 9 1043. 18 MAY 1895) YO ISOTY P E NY HOLOTY PE

190. FISSA VAR. ARISTATA HERMANNvF.J.9 RHODORA 67:198. 1965. USA: FLORIDA: SEMINOLE COO: OVIEDO (RAY9J.D.; WOOD9C.E.; SYITH,A.C. AND EAT0NtR.J.t 10750. 26 APR 1961) GH HOLOTYPE 4Y I SOTYP E us 2449506 ISOTYPE 191. FISSURICOLA MACKENZIETK.K.T MUHLENBERGIA 5:530 1909. USA: NEVADA: ELK0 COO: RUBY MOUNTAINST HUMBOLDT RIVER 56 SMITHSONIAN CONTRIBUTIONS TO BOTANY

(HELLERTA.A.T 9429. 11 AUG 1908) CAS 234898 ISOTYPE

1920 FLACCIOULA STEUDEL,E.G*, SYN. PL. GLUM. 2:199. 1855. USA: OHIO: MIAMI RIVER VALLEY (FRANK,J.C., 55. -- --- 1835) NY TYPE

193. FLACCIFOLIA MACKENZ1EtK.K.r ERYTHEA 8:92. 1922. USA: CALIFORNIA: SOUTHWEST (PARTI (GRANT,G.B.p ---a 01 MAY 1902) us 468192 TYPE FLACCOSPERHA DEWEYfC.9 AMER. J. SCI. ARTS SER.2r 2:245. 1846. USA: FLORIDA: CAMP SABINE (LEAVENWORTH,M.C. T --- . -- --- 18461 GH HOLOTY PE NY I SOTYPE

195. FLAVA VARo GASPENSIS FERNALDTM.L.T RHODORA 8:200. 1906. CANADA: QUEBEC: BONAVENTURE COO: BONAVENTURE RIVER, BETWEEN BALDE AND BAIE DES CHALEURS (COLLINS,J.F.: FERNALDTM.~.. AND PEASE,A.S., --- 05-08 AUG 19041 GH HO L 0 TY P E

196, FLAVA VARo RECTIROSTRA GAUDINTJ.F.G.P.I FL. HEtV. 6:97. 1830. SWITZERLAND: VALAIS (CANTON) : ZERYATT (---, --- -- AUG 1827) GH ISOTY PE

1970 FOENEA VARo PERPLEXA BAILEYTL.H.T MEMO TORREY BOT. CLUB 1:27. 1889. CANADA: NEW BRUNSWICK: KENT COO: -- (FOWLERTJ., ---. -- --- 18711 GH ISOTYPE

1980 FORMOSA DEWEY9C.v AMER. J. SCI. ARTS SER.1, 8:98. 1824. USA: NEW YORK: YATES COO: PENN YAN (SARTWELLTH.P., ---.---I CAS 102307 ISOTYPE CAS 383156 ISOTYPE GH ISOTY P E

199. FRACTA MACKENZIETK-K. 9 ERYTHEA 8:38. 1922.

USA: CALIFORNIA: SISKIYOU Coo: MOUNT SHASTA (PRINGLE,C.G.t -0-0 23 AUG 1881) us 81781 0 HOLOTY PE

200. FRANKLIN11 BOOTTTF. IN HOOKERTW. J. 9 FL. BOR.-AMER. 2:2179 PL.218. 1839 ("1840") USA: --: ROCKY MOUNTAINS (DRUMMONDTTOT--- . ---I GH ISOTYPE VY TYPE

2010 FULVESCENS YACKENZIETK.K.T BULL. TORREY BOT. CLUB 37:239. 1910. ST. PIERRE AND YIQUELON! --: MIQUELON (ISLAND), LANGLADE (ARSENE,L.(FRERE), ---a 28 JUL 1902) NUMBER 12 57

NY TYPE

202. FUSCOLUTEA BOECKELERtJmO.9 BOT. JAHRB. SYST. 7:278* 1886. MEXICO: SAN LUIS POTOSI: -- (SCHAFFNERTJ~G~T221. -- --- 1877 1 YY TYPE MATERIAL us 397187 TYPE COLLECTION 203- FUSCOTINCTA MACKENZIETK~K~TBULL. TORREY BOT. CLUB 36:478. 1909. MEXICO: 04XACA: SIERRA DE SAN FELIPE (PRINGLETC.G.T 4839. 19 AUG 1894) NY TYPE us 251773 TYPE COLLECTION us 817237 TYPE COLLECTION

- G-

2 RBERI FERN LDTM.L.T RHODORA 37:253* 19 5. USA: PENNSYLVANIA: ERIE CO.: ERIE9 PRESQUE ISLE (PENINSULA) ( GAR B ER T A. P T --- 09 JUN 18691 GH ISOTY P E us 63525 TYPE MATERIAL

2050 GARBERI VAR. BIFARIA FERNALD9M.L. T RHODORA 372253. 1935. CANADA: QUEBEC: GASPE C0-z WEST GASPET SAINTE ANNE DES MONTS ( COLL INS 9 JmF AND FERNALDq M-L 9 --- 03-17 AUG 1905) GH HOLOTYPE

206. GAYANA VARo HYALINA BAILEYTL~H~TPROC. AMER. ACAD. ARTS 22:135. 1886 ( "1887") MEXICO: SONORA: SONORA (THURBERTG-T 652. ---I NY TYPE

207. GEOPHSLA MACKENZIE9KmK.t BULL. TORREY BOT. CLUB 40:546. 1913. USA: NEW MEXICO: RIO ARRIBA CO.: TIERRA AMARILLA (EGGLESTONTW.W.~ 6584. 18 APR-25 MAY 19111 us 66@800 TYPE

208. GEYERS BOOTTTF.~ TRANS. CINN. SOC. LONDON 20:118. 1846. USA: --: ROCKY MOUNTAINS (GEYERIC-A. T 332. ---I NY TYPE COLLECTION

209. GCAREOSA VARm AMPHIGENA FERNALD9M.L.9 RHODORA 8:47. 1906. CANADA: QUEBEC: BONAVENTURE COO: ESCUMINAC BAY1 ESCUMINAC ( FERNALDT M.L 9 --- 28 JUN 1904) GH HOLOTY PE NY I SOTYP E

2100 GLAUCODEA TUCKERMANqE. EX OLNEYTS-T. IN GRAYTA.~ PROC. AMER. ACAD. ARTS 72395. 1868. USA: MASSACHUSETTS: HAMPSHIRE CO. : MOUNT HOLYOKE (TUCKERMAN9 E.9 --- -- JUN 18641 58 SMITHSONIAN CONTRIBUTIONS TO BOTANY

GH ISOTYPE

211. GRACILfOR MACKENZIETKOKOT BULL. TORREY BOTo CLUB 43:6140 1916. USA: CALIFORNIA: SONOYA Coo: CLOVERDALE (BOLANDER,HoNo 9 38220 -- APR 1864) CAS 10 30 33 ISOTYPE 0s 145620 HO LO TY P E GH ISOTYPE MO I SOTYPE us 319177 ISOTYPE

2120 GR CILLIHA SCHWEINITZTLO OT ANN. LYCEUM N To HISTo NEW YORK 1:66. 18240 USA: PENNSYLVANIA: -- (SCHUEINITZ,L.D.T --- 0 ---I GH ISOT Y P E

2130 GRIFFITHII BOOTTIFOT TRANS. LINNo SOCo LONDON 2O:1380 18460

AFGHANISTAN: --:0- (GR1FFITHtW.r 78(KEW 607410 ---I NY TYPE COLLECTION

2140 GRISEA VARo RIGIDA 6AILEYyLoH.y MEMO TORREY B0To CLUB 1:56. 18890

USA: PENNSYLVANIA: BUCKS COO: SELLERSVILLE (FRETZ,C.D.T ---0 -- --_ 1884) GH HOLOTY PE

2150 GUATEMALENSIS HERMANN,FoJo, BRITTONIA Z3:14So 19710 GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, BETWEEN TOJIAH AND CHEMALt ALTO 3380 Yo (BEAMANyJoHoy 38800 31 JUL 1960) GH HOLOTYPE

216. GYHNOCLADA HOLMTHOTOT AMERo Jo SCIo SER.4, 14:424. 19020 USA: OREGON: HURRICANE CREEK: BOGS AT 6000 FTo (cuSIcKyW.co~ 24870 28 AUG 1900) YO TYPE MATERIAL NY ISOTYPE

GYNOOYNAMA OLNEYySoTo IN GRAYyAoq PROCo AMERo ACADo ARTS 7:394o 1868. USA: CALIFORNIA: MENDOCINO Coo: MENDOCINO CITY (BOLANDERqHoNoy 47@@. -- --- 1866) CAS 383986 ISOTYPE 0s 495@0 ISOTYPE DS 490408 ISOTYPE NY I SOTYP E

- H-

218, HAGIANA KELS0,L.p BIOLo LEAFLo 30:20 19450 USA: COLORADO: HAGUES PEAKS (KELSOTLO AND KELSO,E~H.T 5250 C8 AUG 1936) NUMBER 12 59

CAS 328017 ISOTYPE

219. HALE1 DEWEYvC., AMER. J. SCI. ARTS SER.29 2:248. 1846. USA: LOUISIANA: MISSISSIPPI RIVER (LEAVENWORTH9M.C. AND HALEqD.9 683. ---I NY SYNTYPE

220. HALLIANA BAILEY,CoH.y BOT. GAZ. 9:117. 1884. USA: OREGON: -- (HALL9E.q 606. -- --- 18711 GH TYPE COLLECTION

221. HALL11 OLNEY9S.T. IN PORTER9T.C. IN HAYDEN9F.V.9 ANN. REP. U.S. GEOL. SURV. TERR. 5:496. 1872. USA: COLORADO: ROCKY MOUNTAINS: LAT. 39-41 No (HALLvE. AND HARB0URqJ.P.r 617. -- --- 18621 F 314892 SYNTYPE F 456958 SYNTYPE GH SY NTYP E YO SYNTYPE NY SYNTYPE us 29651 SYNTYPE USA: IDAHO: PLEASANT VALLEY (PORTERTT.C. 9 --- 26-29 JUN 1871) NY SYNTYPE

222. HALSEYANA DEWEYTC., AMER. J. SCI. ARTS SER.19 11:313. 1826. USA: MASSACHUSETTS: HAMPDEN COO: UESTFIELD (DAVISTE.~---. ---) GH HOLOTY PE NY I SOTYPE

223. HARFORDII YACKENZIE,Y.K.9 BULL. TORREY BOTo CLUB 43:615. 1916. USA: CALIFORNIA: -- (KELLOGG, A. AND HARFORD,W.G.U.9 1073. _- --- 1868-1 869) NY HOLOTYPE us 28685 ISOTYPE

224. HARPER1 FERNALDTM.L.T RHODORA 8:181. 1906. USA: GEORGIA: JEFFERSON COO: LOUISVILLE, ROCKY COMFORT CREEK (HARPER+R.Y.? 2109. 09 APR 1904) F 176870 ISOTYPE GH HOLOTYPE NY ISOTYP E

225. HASSEI BAILEY9L.H.9 BOT. GAZ. 21:5. 1896. USA: CALIFORNIA: SAN BERNARDINO COO: SAN BERNARDINO MOUNTAINS, SAN ANTONIO CANYON; ALTO 4500 FT. (HASSE9H.E.f ---. -- JUL 1894) NY TYPE COLLECTION

226. HATUSIMANA OHWIqJ., JAP. J. BOf. 7:196. 1934. TAIWAN: FUKIEN: KAOHSIUNG ("TAKA0")t DAIJURIN (OHWIqJ.9 329. -- MAR 1933) F 1411493 TYPE YATERIAL 60 SMITHSONIAN CONTRIBUTIONS TO BOTANY

221. HAVOENIANA OLNEY tS.T. IN WATSON, So 9 BOT. U.S. GEOL. EXPLOR. 40TH PAR. 366. 1871. USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (BOtANDERtHoN., 5074. ---) GH SYNTYPE

228. HAYOENII DEWEYtC.9 AMER. J. SCI. ARTS SER.29 18:103. 1854. USA: SOUTH DAKOTA: STANLEY COO: FORT PIERRE (HAYDENtF*V.t 21. -4 --- 1853-1 854) YO TYPE MATERIAL

229. HEBETATA BO0TTtF.t ILL. GENUS CAREX 4:172t PL.583. 1867. PERU: LIMA: OBRAGILIA? NEAR LIMA (WILKES EXPLOR. EXPED.9 ---. -- --- 1838-1 842 1 GH HOLOTYP E

230. HELLERI MACKENZIEpK.K.9 ERYTHEA 8:80. 1922. USA: MEVADA: WASHOE COO: MOUNT ROSE (HELLERt A.A.9 99750 23 JUL 1910) F 283119 TYPE MATERIAL NY TYPE us 509004 ISOTYPE 231 HEPBURNII BOOTTtF. IN HOOKER9W.J.t FL. BORo-AMER. 222099 PL.207. 1839 ( "1 840" 1 USA: COLORADO: ROCKY MOUNTAINSv SOUTH PARK (DRUMM0ND.T. 9 256. --- ) GH SYNTY PE

232. HETERONEURA BOOTT,W. IN WATS0NtS.t GEOL. SURV. CALIFORNIA9 BOT. 2:239. 1880. USA: CALIFORNIA: LAKE TAHOE TO BEAR VALLEY (KELL0GGtA.t ---. 03 AUG ---- 1 GH TYPE MATERIAL us 28206 TYPE COLLECTION

233. HETEROSTACHYA TORREYtJ. EX 0EWEYvC.t AMER. Jm SCI. ARTS SERo2t 2:248. 1846. USA: MICHIGAN: CHIPPEWA COO: DRUMMOND ISLAND (TORREYtJ. 9 &--. 03 AUG 18391 NY TYPE

234. HINDS11 VARm BREVIGCUHA KUKENTHALtG. IN ENGLER,H.G.Aet PFLANZENR. 49 FAM.20:307. 1909. USA: IDAHO: BONNER CO.: HOPE (SANDBERGvJoHot 9330 20 AUG 1892) NY ISOTYPE

235. HIRSUTA VARm CUSPIDATA DEWEYtC. IN WOODtA., CLASS-BOOK BOT. 758. 1861. USA: XLLINOIS: -- (VASEY9G.t --- . ---I YY TYPE COLLECTION NUMBER 12 61

2360 HITCHCOCKIANA DEWEYtC.9 AMERm J. SCIm ARTS SER.1, 10:274. 1826. USA: MASSACHUSETTS: BERKSHIRE Corn : WILLI AMSTOWN, SADDLE MOUNTAIN (DAVIS,Emt ------1823) GH HOLOTY PE

237- HOLMIANA MACKENZIE+KmKmt BULL. TORREY BOTm CLUB 36:481. 1909. USA: MONTANA: JOHN'S LAKE (VREELANDtFmK., 1121. 19 AUG 1901) NY TYPE

238- HOOD!! VARo NERVOSA BA1LEYtL.H.t MEMm TORREY BOT. CLUB 1:14m 1889. USA: CALIFORNIA: -- (KELLOGGtA. AND HARFORD+W.G.W.t 1069. -- --- 1868-1 869) CAS 103098 SYNTYPE NY SYNTYPE

239. HOOD11 VAR. NEUROCARPA PIPERtCmVm, CONTRm Urns. NATL. HERB. 11:167m 1906. USA: CALIFORNIA: -- (KELLOGGtA. AND HARFORDtW.GmW., 1069. -- _-- 1868-1869) C AS 103098 SYNTYPE NY SYNTYPE

240 0 HOOKERANA DEWEY9C.t AMER. Jm SCIm ARTS SER.1, 29:248m 1836. CANADA: SASKATCHEWAN: CARLTON HOUSE (52 51'N.t 106 13'W.l (R1CHARDSONtJ.t --- -0-1 GH TYPE COLLECTION NY TYPE COLLECTION

2410 HORMATHODES FERNALDTY~L.~RHODORA 8: 165. 1906. USA: RHODE ISLAND: PROVIDENCE COm: PROVIDENCE (OLNEYt SmTet ---m 01 JUL 1867) GH SYNTYPE

242 HORNSCHUCHIANA VARe LAURENTIANA FERNAL0,M.C. AND WIEGAND,KmMmt RHODORA 13: 130. 1911. CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLE MOUNTAIN (FERNALDtMmLm AND WIEGAND,K*Mmt 2897. 16 AUG 19101 GH HOLOTYP E NY I SOTY P E

243 HOSTIANA VARo LAURENTIANA FERNALD, MmLm AND WIEGAND9K.M. 9 RHODORA 26: 122. 1924. CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLP MOUNTAIN (FERNALDtMmL. AND W1EGANDtKmM.t 2897. 16 AUG 1910) GH HOLOTYPE

241. HOUGHTONIANA TORREYpJm EX DEWEY,Cmt AMER. J. SCIm ARTS SER.19 30263. 1836. USA: MINNESOTA: CLEARWATER CO.: LAKE ITASCA ("LAKE LA BICHEt NEAR SOURCES OF MISSISSIPPI RIVER") (HOUGHTON,D.t ---m 13 JUL 18321 NY HOLOTYPE 62 SMITHSONIAN CONTRIBUTIONS TO BOTANY

2450 HUEHUETECA STANDLEYTP~C. AND STEYERMARK9J.A.v PUBL. FIELD YUS. NATO HIST., BOT. SER. 23~195, 1947. GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES 9 CANANA; ALTO 2500 M. (STEYERMARKTJOAOT49055. 18 JUL 1942) F 1128957 HOLOTYPE

2460 HYHENODON OHWI TJ. 9 ACTA PHYTOTAX. GEOBOT. 1:298. 1932. JAPAN: --• HONSHU ( ISLANDIT OSAWAYURA IN SHIMOTSUKE ( SEKIMOTOTH. 9 --- 15 JUL 1932) F 1463659 TYPE MATERIAL

-I-

2470 ICHANGENSIS CLARKETC.B.T J. LINN. SOC.9 BOT. 36:290. 1903. CHINA: HUPEH: -- (HENRYvA.9 7860. -- --- 1885-1888 1 us 802160 TYPE MATERIAL

248. XOAHOA BAILEY9L.H.9 BOT. GAZ. 21:5. 1896. USA: IDAHO: BEAVER CANYON (RYDBERGvP.A.9 2339, 07 AUG 1895) us 235568 TYPE COLLECTION us 235569 TYPE COLLECTION

249. IGNOTA DEWEYTC-T AMER. J. SCI. ARTS SER.29 8:348. 1849, USA: LOUISIANA: RAPIDES PARISH: ALEXANDRIA (HALEpD.9 97. ---I C AS 553902 TYPE COLLECTION YY TYPE COLLECTION

2500 ILLINOENSTS DEWEYTC-T AMER. J. SCI. ARTS SER.29 3:245. 1847. USA: ILLINOIS: HANCOCK COO: AUGUSTA (MEAD~S*B.T --- . ---I NY TYPE COLLECT ION

2510 ILLOTA BAILEY9L.H.t MEM. TORREY BOT. CLUB 1:15o 1889. USA: COLORADO: ROCKY MOUNTAINS: LAT. 39-41 No (HALLTE. AND HARBOURTJ.P.T 591. -- --- I862 I F 314869 I SOTYPE F 456934 ISOTYPE GH HOLOTYPE YO I SOTYPE

2520 INCISOoDENTATA STEUDELTE.G.T SYN. PL. GLUM. 2:189. 1855. CHILE: --: -0 ILECHLERTW.~ 1136. -- OCT 1852) GH I SOTYPE

253. INCOHPERTA BICKNELL9E.P. 9 BULL. TORREY BOT. CLUB 35:494. 1908. USA: MASSACHUSkTTS: NANTUCKET COO: NANTUCKET ISLAND (BICKNELL,E.P.9 --- 20 JUN 1908) NY TYPE

2540 INCONDITA HERMANNTF. J. T LEAFL. W. BOT. 8:112. 1957. CANADA: ALBERTA: RAM RIVER9 NORDEGG (HERMANNTF. J. 9 13347- 15 AUG 19561 NUMBER 12 63

CAS 404489 ISOTYPE us 2265956 HOLOTYPE

255. INCURVIFORMIS MACKENZIE9K.K. IN RYDBERGvP .A. 9 FL. ROCKY MOUNT. 1209 1060. 1917. CANADA: ALBERTA: BANFF NATIONAL PARK9 BANFf; ALTO 8000 FT. ( MACOUNTJOHN 9 --- 0 31 JUL 1891) GH ISOTYP E NY HOLOTYPE

2560 INFLATA VAR. ANTICOSTENSIS FERNALDpM.L.9 RHODORA 44:329~ PL.715. 1942. CANADA: QUEBEC: ANTICOSTI ISLAND, PETITES-RIVIERES (MARIE-V ICTORIN, (FRER El AND ROLLAND-GERMAI NT(FRERE 1 T 25767. 20 JUL 1926) GH HOLOTYPE

2570 INOPS BAILEY,L.H., PROC. AMER. ACAD. ARTS 22:126. 1886 ("1887"). USA: OREGON: CLACKAMAS COO: MOUNT HOOD (HENDERSONTL.F.T ---. -- JUL 1884) CAS 203910 ISOTYPE GH HOLOTY PE YY I SOTYPE

258. INTEGRA MACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 43:608. 1916. USA: CALIFORNIA: PLACER COO: SUMMIT; ALTO 7000 FT. (HELLERTA.A.T 9841. 16 JUL 1909) NY HOLOTY P E

2590 INTERIWUS YAGUIRETB., BRITTONIA 5: 200. 1944. USA: UTAH: CACHE COO: TONY GROVE LAKE (MAGUIRE96.9 16098. 05 AUG 19381 CAS 348506 ISOTYPE GH ISOTYPE NY TYPE us 1872574 ISOTYPE 260. INTERIOR BAILEY9C.H., BULL. TORREY BOT. CLUB 20:426. 1893. USA: NEW YORK: YATES COO: PENN YAN (SARTWELL9H.P.9 36. ---I CAS 553999 ISOTYPE YO 1816496 TYPE COLLECTION NY ISOTYPE

261. INTERIOR VAR. CHARLESTONENSIS CLOKEY9I.W.9 BULL. So CALIF. ACAD. SCI. 38:l. 1939. USA: NEVADA: CLARK COO: CHARLESTON PARK (CLOKEY, I.W.9 7468. 19 JUN 19371 CAS 272528 ISOTYPE CAS 272529 I SOTYPE DS 278190 ISOTYPE F 10 76930 ISOTYP E GH ISOTYPE JEPS 4013 ISOTYPE 64 SMITHSONIAN CONTRIBUTIONS TO BOTANY

YO 1148381 ISOTYPE MO 1190731 I SOTYPE YO 1201697 ISOTYPE NY ISOTYPE uc 910020 HOLOTYPE us 1733722 ISOTYPE 2620 INTERIOR VARo JOSSELYNII FERNALDfY.L.9 RHODORA 8:115. 1906. USA: MAINE: AROOSTOOK CO,: SAINT JOHN RIVER9 FORT KENT ( FERNALD9M.L.9 --- 06 JUL 1904) GH HOLOTY PE NY ISOTYPE us 60 5797 ISOTYPE

263. INTERIOR VAR, KEWEENAWENSIS HERMANN9F.J.9 AMER. MIDL. NATURALIST 25:19. 1941. USA: MICHIGAN: KEWEENAW CO.: EAGLE HARBOR (HERMANN9FoJ.t 7985. 13 JUL 19361 GH HOLOTY PE NY ISOTYPE us 1697057 ISOTYPE 264. INTERRUPTA VAR, DISTENTA KUKENTHAL9G.t REPERT. SP. NOV. REGNI VEG. 26: 254. 1929. USA: WASHINGTON: KLICKITAT COO: BINGEN (SUKSDORFqW.N.9 12333. 22 AUG-05 SEP 1927) CAS 242959 TYPE COLLECTION 265. INTUHESCENS FOR0 VENTRIOSA FERNALD9M.L.v RHODORA 44:321r PL.713. 1942 USA: VERMONT: ADDISON COO Z RIPTON (BRAINERD9E.9 ---a 19 JUL 1898) GH HOLOTY PE

266, fNVOLUCRATELLA MACKENZIEtK.K.9 No AMER. FL. 18:50. 1931. MEXICO: SAN LUIS POTOSI : LAS CANOAS ( PRI NGLE TC. Go 9 3126. 08 JUL 1890) F 263394 TYPE COLLECTION GH ISOTYPE MO TYPE MATERIAL VY TYPE COLLECT ION us 30661 TYPE COLLECTION

-J-

2670 JACINTOENSIS PARISH9S.B.9 BULL. So CALIF. ACAD. SCI. 4:lOOq PL.16. 1905. USA: CALIFORNIA: RIVERSIDE COO: SAN JACINTO MOUNTAINS9 TAMARACK VALLEY; ALT, 9000 FT. (HALLTH.M. 9 2483. -- JUL-AUG 1901) DS 78003 HOLOTYPE NUMBER 12 65

268. JACOBI-PETER1 HULTENTO.E.G.T ACTA UNIV. LUND. N.S.c 38:30C~FIG.4. 1942 USA: ALASKA: TIN CITY (ANDERSON,J.P., 4871. 19 AUG 1938) CAS 477664 ISOTYPE

269. JAMES11 TORREY,J.r ANN. LYCEUM NATO HIST. NEW YORK 3:398. 1836. USA: --: ROCKY MOUNTAINS (JAMESTEDWIN, --- . ---I NY HOLOTY PE

270. JAMESONI VARo SUBFULVA KUKENTHALTG.~ REPERT. SP. NOV. REGNI VEG. 8:7. 1910. BOLIVIA: --• -- (BANG1M.p 2376. ---I us 825890 TYPE MATERIAL

271. JEPSONII HOWELLTJ.T.~ LEAFLO W. BOT. 8:223. 1958. USA: CALIFORNIA: TUOLUYNE COO: YOSEMITE NATIONAL PARK, TUOLUMNE MEADOWS; ALTO 8800 FT. (JEPSON,W.L.r 4477. 20 JUL 1911) J EPS 20008 ISOTYPE NY HOCOTY PE

272- JONES11 BAILEY,L.H., MEMO TORREY BOT. CLUB 1:16. 1889. USA: CALIFORNIA: NEVADA COO: SODA SPRINGS; ALTO 7000 FT. (JONESTY.E., ---. 22 JUL 1881) NY SYNTYPE

-K -

273. KALOIDES PETRIE,D.p TRANS. & PROC. NEW ZEALAND INST. 13:332. 1881. NEW ZEALAND: OTAGO (DISTRICT): SOUTH ISLAND, CARRICK RANGE; ALTO 4000 FT. (PETRIETD., --- 0 ---I GH ISOTYPE

274. KATAHDINENSIS FERNACD,M.L.r RHODORA 3:171~PL.32. 1901. USA: MAINE: PISCATAQUIS COO: MOUNT KATAHDIN, DEPOT POND ( W I L L IA MS T E F ; C H URC H IL L 7 J R AND F E R NAL D T M L T --- 16 JUL 19C.O) GH HOLOTY PE NY ISOTYPE us 1325047 ISOTYPE

275- KAUAIENSIS KRAUSSTR., PACIFIC SCI. 4:279. 1950. USA: HAWAII: KAUAI COO: KAULUWEHI tROCK,J.F.q 9017. -- OCT 1909) us 2074700 TYPE MATERIAL

276. KELLOGGII BOOTTTW. IN WATSONTS., GEOL. SURV. CALIFORNIA, BOT. 2:240. 1880. USA: CALIFORNIA: SIERRA NEVADA RANGE, "LAKE TAHOE TO BEAR VALLEY" (KELLOGG, A. 9 --- . ---1 66 SMITHSONIAN CONTRIBUTIONS TO BOTANY

GH SYNTYPE

277- KOKRINENSIS PORSILD,A.E., RHODORA 41:206, PL.551. 1939. USA: ALASKA: KOKRINES MOUNTAINS (PORSILD, A.E. AND PORSILDIROT. 9 711. 23 JUN-05 JUL 1926) GH ISOTYPE

278 0 KULfNGANA BAILEYTL~H~TGENTES HERB. 1:13. 1920. CHINA: KIANGSI: KULING; ALTO 2500-3500 FT. (BAILEYTL.H.T ---a 18 JUL 1917) NY TYPE

2796 KURILENSIS OHW1,J.y ACTA PHYTOTAX. GEOBOT. 2:270 19330 USSR: RUSSIAN SFSR: SAKHALI N OBLAST: KURIL ISLANDS, SHIKOTAN (ISLAND), NOTORO; (COUNTRY AS "JAPAN") (UHWI 9 J. 9 813. 11 AUG 1931) F 1406403 TYPE M4TERIAL

-L-

280. LACINIATA BOOTT cF. T ILL. GENUS CAREX 4:175, PL.594. 1867. USA: CALIFORNIA: SACRAMENTO RIVER (RICHTWILLIAMT WILKES EXPED. 1241. -- --- 1838-1842) NY TYPE

281. LACUNARUM HOLYTH.T. T AHER. J. SCI. SER.4, 17:316. 1904. USA: CALIFORNIA: SONOMA COO: SEBASTOPOL (HELLER,A.Ao, 5797. @lJUL 1902) F 129242 SYNTYPE F 1566419 SYNTYPE YO SYNTYPE NY SYNTYPE US 430229 SYNTYPE

282- LAEVI-CONICA DEWEYTC.~ AMER. J. SCI. ARTS SER.2, 24:470 18570 USA: NEBRASKA: BIG SIOUX RIVER (HAYDEN9F.V.t --- . ---I GH HOLOTYPE

283. LAHPROCHLAHYS BLAKETSOT., J. ARNOLD ARBOR. 28:1G4. 1947. PAPUA AND NEW GUINEA: PAPUA (TERRITORY): MAFULU; (COUNTRY AS "BRIT1 SH NEW GUINEA") (BRASS TL. J. T 5323. -- SEP-NOV 19331 A I SOTYPE

204. CANCIFOLIA CLARKETCOB.~J. LINN. S0C.r BOT. 36:292. 1903. CHINA: HUPEH: -- (HENRY~A~T5467. ---I us 801132 SYNTYPE

285. LANCIFRUCTUS HACKENZIETK.K.T BULL. TORREY BOT. CLUB 43:607. 1916. USA: CALIFORNIA: TULARE COO: UPPER KERN RIVER, VOLCANO CREEK (HALL9H.M. AND BABCOCKtH.D., 5472. -- JUL 19041 NY I SOTYPE NUMBER 12 67

uc 127723 HOLOTYPE

2860 LANGEANA FERNALDTM~L.~RHODORA 35:217o 1933. CANADA: NEWFOUNDLAND: GARGAMELLE COVE (FERNAL0,M.L.; LONGTB. AND FOGG-JR~TJ~M~T1374. 20 JUL 19291 F 1481645 ISOTYPE GH HOLO TY P E

287. LARENSIS STEYERMARKyJoAov FIELDIAVA, BOTo 28:66~FIG.8. 19510 VENEZUELA: LARA: BETWEEN BUENOS AIRES AND PARAMO DE LAS ROSAS (STEYERMARKTJ.A.T 554700 11 FEB 1944) F 55470 HOLOTYPE us 1932015 ISOTYPE

288. LARICINA MACKENZIETK~KOEX BRIGHTTJoT TRILLIA 9:4,190 19300 USA: INDIANA: KOSCIUSKO Coo: LEESBURG (DEAMyCoCor 10927. 05 JUN 19121 NY TYPE

289- LASIOCARPA VARo AMERICANA FERNALDTM~L.~RHODORA 44:3O40 1942. CANADA: NOVA SCOTIA: YARMOUTH COO: ARGYLE (PEASEvAoSo AND LONGTB.T 205190 09 JUL 19201 GH HOLOTYPE

290 0 LATEBRACTEATA WATERFALL,U.To, RHODORA 56: 23. 19540 USA: OKLAHOMA: YCCURTAIN COO: BROKEN BOW (WATERFALLTU~T~T 11380. 19 APR 19531 C AS 384438 ISOTYPE GH ISOTYPE YO 1692174 ISOTYPE

291 LAXIFLORA VAR. LEPTONERVIA FERNALDtM.L.9 RHODORA 8:184. 19060 USA: MAINE: AROOSTOOK CO.: FORT FAIRFIELD (FERNALD,M.L., 146. 06 JUL 18931 F 267758 ISOTYPE GH HOLOTY PE MO ISOTYPE NY ISOTYPE 292 - LAXIFLORA VARo SERRULATA HERMAVN?F.Jo, RHODORA 40:80. 19380 USA: INDIANA: CLARK Coo: -- (DEAM~C~C~T6458. 25 MAY 1910) GH HOLO TY P E NY ISOTY P E

293 LEAVENUORTHII DEWEYTC~TAMERo Jo SCI. ARTS SER.2, 2:2460 1846. USA: LOUISIANA: -- ( LEAVENWORTHTMOC. T ------4 1845 J NY TYPE COLLECTION

294. LEIOCARPA MEYERTC~A.~ MEMO ACADo IMP. SCIo ST.-PETERSBOURG DIVERS SAVANS 1:208t PL.5. 18310 USA: ALASKA: SITKA (MERTENS?CoHo, --- . ---I GH ISOTYPE 68 SMITHSONIAN CONTRIBUTIONS TO BOTANY

2950 LEIOPHYLLA MACKENZIEqKmKm t Nm AMERm FLm 18:365m 1935- CANADA: YUKON TERRITORY: CARCROSS (EASTWOODTAm, 725Am 16 JUL 1914) C AS 102481 ISOTYPE GH ISOTYPE us 538796 HOLOTYPE

296- LEMANNIANA VARo SIMPLEX KUKENTHALvGm IN ENGLERqHmGmAmv PFLANZENRm 49 FAM.20:405* 1909- COSTA RICA: --: CERRO DE BUENA VISTA (PITTIERTHm AND TONDUZTA.T 3381. 19 JAN 18911 CAS 264341 SYNTYPE

297. LEMMONI BOOTT,Wm, BOTm GAZm 9:93m 1884- USA: CALIFORNIA: SIERRA NEVADA RANGE (LEMMON,JmGmv ---a -- --- 1875) GH TYPE COLLECT I ON us 29211 TYPE COLLECTION

2980 LENTICULARIS VARo PAULLIFRUCTUS KUKENTHAL cGm IN ENGLER, HmGm Am 9 PFLANZENRm 49 FAMm20:308m 1909. USA: WASHINGTON: WHI TMAN COm : PALOUSE CREEK ( ELMER9 AmDm Em T 88 1. -- JUN 1897) NY ISOTYPE

2990 LEPORINA VARo AMERICANA OLNEYvSmTm EX BAILEY~L~H~T PROCm AMERm ACAD. ARTS 22:152. 1886 ("1887")m USA: OREGON: CLACKAMAS COm: MOUNT HOOD (HALLpEm, 583- 01 AUG 18711 F 455706 TYPE COLLECTION F 1425899 TYPE COLLECT ION GH TYPE C OLL ECT ION YO TYPE COLLECTION NY TYPE COLLECTION

3000 LEPORINELLA MACKENZIEtKmKm, BULL- TORREY BOTm CLUB 43:605m 1916- USA: C4LIFORNIA: EL DORADO COm: PYRAMID PEAK (HALL9H.M. AND CHANDLERvHmAm T 4716- 01-02 AUG 19031 DS 490443 ISOTYPE GH I SOTYPE uc 55234 HOLOTYPE

301 LEPTOPODA YACKENZIE,K.Km IN RYDBERGTP~A., FL. *ROCKY MOUNT. 124, 1060. 1917. USA: OREGON: CLACKAMAS COm: OSWEGO, ELK ROCK (HELLER9A.A. 9 10052m 20 YAY 1910) CAS 186427 ISOTYPE DS 13923 ISOTYPE NY TYPE

302 LIWNOPHILA HERMANNTF- Jo T LEAFLm Wo BOTm 8:28. 19560 USA: WYOMING: SUBLETTE COm: PINEDALE (HERMANNfFmJm, 12252- SUMBER 12 69

21 AUG 19551 us 2231577 HOLOTYPE

303. LIVIOA VAR. RUFINAEFORMIS FERNALDtMoLo t RHODORA 28: 80 1926. CANADA: NEWFOUNDLAND: STRAIT OF BELLE ISLE9 FOUR-MILE COVE (FERNAL0tM.L. t WIEGANDvKo Mo AND LONGtBo t 276730 20 JUL 1925) GH HOLOTYP E

3040 LONGICRURIS VARo HENRY1 CLARKEpCoB.9 Jo LINN. SOC.9 BOTo 36:2950 19030 CHINA: HUPEH: -- (HENRYIA~T 42660 ---I us 800846 SYNTYPE

3050 LONGICULMIS PETRIEtDot TRANS. & PROCo NEW ZEALAND INSTo 14:3630 1882. NEW ZEALAND: OTAGO (DISTRICT): SOUTHLAND SUBOIVI SION: STEWART ISLANO PATTERSONS INLET (PETRI ETDo t --- 0 -- JAN 1880) GH ISOTYPE

3060 LONGIROSTRIS VARo MICROCYSTIS BOECKELER,JoOot LINNAEA 41:2410 18770

CANADA: MANITOBA: WINNIPEG (BOURGEAU,Eo, ---0 _- _-- 18 57-1 859 1 NY TYPE COLLECTION

3070 LUNELLIANA MACKENZIETK~K.t BULL. TORREY BOTo CLUB 42:6150 19150 USA: MISSOURI : JACKSON Coo: OAK GROVE (BUSHtBoFo t 70200 02 JUN 1913) NY TYPE

3080 LUZULAEFOLIA VARo STROBILANTHA HOLM,HoTor AMERo Jo SCIo SER.49 20:305. 19050 USA: CALIFORNIA: NEVADA Coo: OONNER PASS; ALTO 7500 FTo (HECLERtAoA.9 7187. 17 AUG 1903) c4s 136 ISOTYPE CAS 231121 ISOTYPE

3690 LUZULINA OLNEYtSoTo IN GRAYtA.9 PROCo AMER. ACADo ARTS 7:3950

1868 0 USA: CALIFORNIA: MENDOCIN0 Coo: YENDOCINO CITY (BOLANDER9HoNop 4740. -- --- 18661 CAS 384084 ISOTYPE 0s 76794 ISOTYPE GH TYPE C OLL ECT ION YO TYPE MATERIAL NY TYPE COLLECT ION us 964880 TYPE COLLECTION

-M-

3100 MACKENZIANA YEATHERBY 9C.A. 9 CONTRo GRAY HERB. 114:360 19360 70 SMITHSONIAN CONTRIBUTIONS TO BOTANY

MEXICO: NUEVO LEON: GALEANA (MULLER9C.H. AND MULLERTM.T.T 892. 28 JUN 1934) CAS 264346 ISOTYPE GH HOLO TYP E us 1746479 ISOTYPE

MACROGLOSSA FRANCHET9A. AND SAVATIERqL.9 ENUM. PL. JAP. 2:148~576. 1879. JAPAN: KANAGAWA (PREFECTURE): HONSHU ( ISLAND) 9 YOKOSUKA (SAVATIER9L.r 1414. -- --- 1866-1874) us 27238 TYPE MATERIAL

MACROKOLEA STEUDELTE-G.~ SYN. PL. GLUM. 2:223. 1855. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMONDtT. 9 420. _- --- 1832 1 NY COTYPE

WACROSPERUA YACKENZIE9K.K.p BULL. TORREY BOT. CLUB 36:477. 1909. YEXICO: OAXACA: SIERRA DE SAN FELIPE; ALTO 6000-7000 FT. (PRINGLE~C.G.T 4840. 27 AUG 1894) NY TYPE

MADRENSIS BAILEY~L*H.T ROT. GAZ. 25:270. 1898. MEXICO: DURANGO: SIERRA MADRE OCCIDENTAL (ROSE9 J.N.9 2357. 16 AUG 1897) NY ISOTYPE lJS 301267 TYPE

315* MAGNIFOLIA Y~CKENZIETK~K.IN SMALLTJ~K~TFL. SE. U.S. ED029 1325. 1913. USA: FLORIDA: -- (CHAPMANvA.W.9 --- . ---I us 969118 TYPE MATERIAL

MANDONIANA BOECKELER9J.O.9 ALLG. BOT. 2. SYST. 2:174. 1896. BOLIVIA: --: -- (MANDONTG., 1429. ---I NY ISOTYPE

HARCIOA VARo OEBILIS BAILEYTL.H.T PROC. AMER. ACAD. ARTS 22:136. 1886 ("1887" USA: OREGON: HARNEY COO: HARNEY VALLEY (HOWELL9T.J.r 937. 27 MAY 1885) F 206587 TYPE COLLECTION NY TYPE

318. MARIPOSANA BAILEY9L.H.p BULL. TORREY BOT. CLUB 43:619. 1916. USA: CALIFORNIA: TUOLUMNE COO: YOSEMITE NATIONAL PARKT TUOLUMNE MEADOWS (JEPSON? W .L 9 4476. 20 JUL 1911 1 JEPS 19722 ISOTYPE NY HOLOTY PE

MEAD11 DEWEYqC.9 AMER. J. SCI. ARTS SER.1, 43:9@. 1842. USA: ILLINOIS: HANCOCK COO Z AUGUSTA (MEAD9S.B.p --- . ---I CAS 553885 ISOTYPE NUMBER 12 71

GH HOLOTY PE YO I SOTYPE NY ISOTYPE

320- MEDITERRANIA YACKENZIETK~K-TBULL. TORREY BOT. CLUB 33:441. 1906. USA: DISTRICT OF COLUMBIA: WASHINGTON (STEELE,E.S., ---. 23 MAY 1898) NY TYPE

321. MEEKII DEWEYTC., AMER. J. SCI. ARTS SER.29 24:48. 1857. USA: NEBRASKA: WHIT€ RIVER (HAYDEN,F.V., --- . ---I GH TYPE COLLECTION

322. MELANOPHORA BLAKE,S.T., J- ARNOLD ARBOR- 28: 106. 1947. INDONESIA: WEST NEW GUINEA: ORANGE RANGE, MOUNT WILHELMINA; (COUNTRY AS "DUTCH NEW GUINEA") ( BRASS, L. J AND MEYER-DREES~E~T9828. -- SEP 1938) A I SOTYPE

323. MELOZITNENSIS PORSILD,A.E., RHODORA 41:209. 1939. USA: ALASKA: KOKRINES MOUNTAINS YELOZ ITNA RIVER (PORSI LD, A. E AND PORSILDTR.T.T 713. 23 JUN-05 JUL 1926) GH I SOTYPE us 1789621 ISOTYPE

324. MENDOCINENSIS OLNEY 9S.T. EX BOOTTTW. IN WATSONvS.9 GEOL. SURV. CALIFORNIA, BOT. 2:249. 1880. USA: CALIFORNIA: YENDOCINO COO: MENDOCINO CITY (BOLANDERTH.N. T 4701. -- --- 1866 1 CAS 553875 TYPE FRAGMENT DS 54832 ISOTYPE GH HOLOTY PE YO I SOTYPE NY I SOTYP E uc 1098 ISOTYPE us 29453 I SOTYPE

325. MERCARENSIS HOCHSTETTER ct OF EX STEUDEL TE.Go T SYN. PL. GLUM. 2 :194. 1855. INDIA: --: NILAGIRI (HOHENACKERTR.F.T 943. -- --- 18511 A I SOT YP E 32b- YERRILLII KUKENTHALTG.~ REPERT. SP. NOV. REGNI VEG. 8:7. 1910. PHILIPPINES: BENGUET: LUZON (ISLAND), PAUAI (MERRILL,E.D.,

6623. -0 MAY 1909) NY TYPE MATERIAL us 711171 TYPE MATERIAL

327. MERRITT-FERNALDII MACKENZIE?K.K., BULL. TORREY BOT. CLUB 49:370. 1923 USA: MAIVE: PENOBSCOT COO: ORONO (FERNALD,M.L., ---- 03 JUL 1897) GH HOLO TYP E 72 SMITHSOXIAN CONTRIBUTIONS TO BOTANY

328. MESOCHOREA MACKENZIE,K.K. y BULL. TORREY BOT. CLUB 37:246. 1910. USA: DISTRICT OF COLUMBIA: WASHINGTON (STEELEvEoSo, ---a -- -_- 1900 1 GH ISOTYPE

329. MICANS BOOTTTF. IN GRAYvA.9 MEMO AMER. ACAD. ARTS NOS., 6:419. 1859. JAPAN: --: SINODA (WRIGHTTC.T --- . ---I us 27281 TYPE MATERIAL

330 MfCRANTHA KUKENTHALTG., BULL. HERB. BOISSIER SER.2, 2:1018. 1902. KOREA: --: KAM-OUEN (FAURIE9U.T 919. 28 JUN 19011 us 2501314 ISOTYPE

331, MICROCHAETA HOLM,H.T., AMER. J. SCI. SER.49 17:305. 1904. CANADA: YUKON TERRITORY: KLONDIKE INDIAN DIVIDE (MACOUN, JOHN9 53877. 14 AUG 1902) GH I SOTYPE

332. HICROGLOCHIN SSPm FUEGINA KUKENTHALqG., BOT. JAHRB. SYST. 27:546. 1899. CHILE: MAGALLANES: TIERRA DEL FUEGO ("FUEGIA"1t ORANGE HARBOR IWI LKE S E XPLOR. EXPED. T --- . -- --- 1838-1 842 1 us 30695 ISOTYPE

333, MfCROPTERA YACKENZI E,K OK T YUHL ENBERGIA 5 :56 1909. USA: NEVADA: ELK0 COO: DEETH (HELLERTA~A.~9067. 21 JUL 1908) CAS 234896 I SOTYPE NY TYPE

336, MICROPTERA VARm CRASSINERVIA HERMANN,FoJ., RHODORA 70:420. 1968. USA: COLORADO: OURAY COO: ENGINEER PASS (JOHNSONTW.M.T 594. 14 AUG 19671 us 2 543 807 HOLOTY PE

335. MILIARIS VARm AUREA BAILEYTL.H.T MEMO TORREY BOT. CLUB 1:37. 1889. CANADA: NEW BRUNSWICK: KING'S Coo: KENNEBECASIS RIVER (FOWLERIJ~T --- . ---I GH HOLOTY PE

336, MIRABILfS DEWEYTC.~ AMER. J. SCI. ARTS SER.1, 30:63. 1836. USA: MASSACHUSETTS: FRANKLIN COO: DEERFI EbD (DEWEYTC~T---. --- ) GH HOLOTY PE

3370 MIRABILIS VAR. PERLONGA FERNALD~M.L.~ PROC. AMER. ACAD. ARTS 37:473, PL.2. 1902. USA: NEW HAMPSHIRE: HILLSBORO COO: NEW IPSWICH (FERNALDvM.L.9 ---. 05 JUN 1896) GH SYNTY PE

3380 MIRABILIS VARm TINCTA FERNALDTY.L*T PROC. AMER. ACAD. ARTS 37:473. NUMBER 12 73

1902. CANADA: NEU BRUNSWICK: SAINT JOHN RIVER (MACOUNtJOHN, 22. 04 JUL 1899) GH SYNTYPE

339, MISANDROIDES FERNALI3,M.L.r RHODORA 17: 158. 1915. CANADA: NEWFOUNDLAND: PORT AU PORT BAY, TABLE MOUNTAIN (ST.JOHN,H. AND FERNALD9M.L. T 10801. 16-17 JUL 1914) GH HOLOTY PE NY ISOTY PE

340 MISERA BUCKLEY,SoB., AMER. J. SCI. ARTS SER.19 45:173. 1843. USA: NORTH CAROLINA: MITCHELL COO: ROAN MOUNTAIN (BUCKLEY cS.B. 9 --- . ---I NY I SOTYPE

341 MISERABILIS YACKENZ1ErK.K.r Ne AMER. FL. 18:385. 1935. USA: WASHINGTON: CHELAN COOZ CHIWAUKUM LAKE (EGGLESTON, Wow. 9 13567. 19-20 AUG 1916) us 886422 HOLOTYPE

3420 MOHRIANA YACKENZIE,K.K.9 No AMER. FL. 18:106. 1931. USA: FLORIDA: HARDEE COO: WAUCHULA (CURTISSrA.H.1 6761. 15 APR 1901) NY HOLOTY PE us 2133195 ISOTYPE

343, MOLESTA MACKENZIE9K.K. 9 N. AMEQ. FL. 18:151. 1931. USA: KANSAS: WYANDOTTE COO: QUINDARO (YACKENZIEqK*K., ---. 30 MAY 18971 NY HOLOTYP E

344, HONTANENSIS BA1LEYpL.H.r BOT. GAZ. 17~152. 1892. USA: MONTANA: FLATHEAD COO: UPPER MARAIS PASS (CANBY,W.M., 350. 03 AUG 18831 NY SYNTYPE us 23257 SYNTYPE

34s. MONTEREVENSIS YACKENZIE~K.K.T ERYTHEA 8:92. 1922. USA: CALIFORNIA: MONTEREY COO: PACIFIC GROVE (SMITH,C.P.t 1055. 24 JUL 1905) GH HOLOTY PE

346, MORRISSEYI PORSILD,A.E.~ SARGENTIA 4:21. 1943. CANADA: NEWFOUWDLAND: LABRADOR9 CAPE MUGFORD (PORSILD,AoE. 173. 26 AUG 1937) us 2095886 ISOTYPE

347. MULTICOSTATA MACKENZIE9K.K. T BULL. TORREY 801. CLUB 43:604. 1916. USA: CALIFORNIA: SAN BERNARDINO COO: SAN BERNARDINO MOUNTAINS BEAR VALLEY DAY (PARISHTS.B.T 3609. -- JUN 1895) 0s 4894C9 HOLOTYPE 74 SMITHSONIAN CONTRIBUTIONS TO BOTANY

348. MURICULATA HERMANN9F.J. IN MCVAUGHTR., FIELD t LAB. 17:132. 1949. USA: TEXAS: CULBERSON COO : GUADALUPE MOUNTAINS? MCKITTRICK CANYON (M0OREpJ.A. AND STEYERMARK?J.A., 3625. 25 JUL 1931) CAS 194659 ISOTYPE

- N-

349. NANA BOOTT,F. IN GRAYtAot MEMO AMER. ACAD. ARTS NOS., 6:418. 1859. JAPAN: HOKKAIDO (PREFECTURE): HAKODATE (WRIGHTTC. 9 ---. -- _-- 1853-1 856 1 NY TYPE MATERIAL us 27280 TYPE MATERIAL

350. NEBRASKENSIS DEWEYTC.~ AYER. J. SCI. ARTS SER.2, 18:102. 1854. USA: NEBR9SKA: -- ( HAY0ENtF.V. T --- . ---I NY I SOTY P E

351. NEBRASKENSIS VARI ERUCAEFORMIS SUKSD0RFjW.N.r WERDENDA 1:5. 1923. USA: WASHINGTON: KLICKITAT COO: FALCON VALLEY (SUKSDORF,W.N., 10249. 22 JUN 1919) 0s 171453 ISOTYPE MO 952735 TYPE COLLECTION NY ISOTYPE us 1438017 TYPE COLLECTION

3520 NEBRASKENSIS VARo ULTRIFORMIS BAILEYTL.H.T BOT. GAZ. 21:8. 1896. USA: WASHINGTON: ADAMS COO: RITZVILLE (SANDBERG?J.H. AND LEIBERGTJ*B.T 194. 09 JUN 1893) NY I SOTYPE

3530 NELSON11 MACKENZ1EtK.K. IN RYDBERG,P.A.v FL. ROCKY MOUNT. 137t 1060. 1917. USA: WYOMING: LA PLATA MINES (NELSONtA. AND NELSONtE., 5264. 30 AUG 1898) GH ISOTYPE NY HOLOTY PE

354. X NEOBIGELOWII LEPAGETE., NATURALISTE CANAD. 91:166. 1964. CANADA: QUEBEC: SAGUENAY COO: ROMAINE RIVER (DUTILLYqA. AND LEPAGE,E.? 41,305A. 12 AUG 1963) GH I SOTYPE YY ISOTYPE us 243371 9 ISOTYPE

3550 X NEOFILIPENDULA LEPAGEtE.9 NATURALISTE CANAD. 83:123. 1956. CANADA: NEWFOUNDLAND: BONNE BAY? MAIN RIVER, MAIN ARM (FERNAL0tM.L.; LONGTB. AND FOGG-JR. 9JoMm~1449. 19 AUG 1929) GH HOLOTYPE

356- NEOMEXICANA YACKENZIE*K.K., BULL. TORREY BOT. CLUB 34:153. 1907. USA: NEW MEXICO: SANTA RITA DEL COBRA (BIGELOWTJ.M.T 1547. NUMBER 12 75

TYPE COLLECTION

357, X NEOPALEACEA LEPAGETE~TNATURALISTE CANAD. 83: 137. 1956. CANADA: QUEBEC: PAINT HILLS (DUTILLY, A.; LEPAGETE. AND DUMANvM.9 32975. 03 SEP 19541 GH ISOTY P E us 2176496 ISOTYPE

358. NERVINA BAILEYTL.H. T BOT. GAZ. 10:203. 1885. USA: CALIFORNIA: SIERRA NEVADA RANGE9 SUMMIT CAMP (KELLOGGTA. 9 --- 10 JUL 1870) GH HOLOTY PE us 286861 ISOTYPE

359, NEUROPHORA MACKENZIE9K.K. IN ABRAYS9L.9 ILL. FL. PACIFIC STATES 122989 FIG.706. 1923. USA: WASHINGTON: CHELAN COO : CASCADE MOUNTAINS, STEVENS PASS (SANDBERG,J.H. AND LEIBERG,J.Bo, 773. 18 AUG 1893) GH ISOTYPE NY ISOTYP E 360 - NIGRICANS MEYER9C.A.9 MEY. ACAD. IMP. SCI. ST.-PETERSBOURG DIVERS SAVANS 122109 PL.7. 1831. USA: ALASKA: ALEUTIAN ISLANDS? UNALASKA ( ISLAND) (CHAYISSO9L.A.9 --- . -0-1 GH ISOTYPE

361 NIGRO-HARGINATA SCHWEINITZTL.D.T ANN. LYCEUM NATO HIST. NEW YORK 1:68. 1824. USA: NORTH CAROLINA: FORSYTH COO z WINSTON-SALEM ("SALEM") (D0StL.t -a- . ---I NY TYPE COLLECTION

362 0 NOVAE-ANGLIAE SCHWEINITZtL. Do 7 ANN. LYCEUM NATO HIST. NEW YORK 1t67. 1824. USA: MASSACHUSETTS: BERKSHIRE COO: WILLIAMSTOWN, SADDLE MOUNTAIN (DEWEYTC.~ *-- . -- --- 1822) GH ISOTYPE

363. X NUBENS LEPAGETE~TNATURALISTE CANAD. 84:40. 1957. CANADA: QUEBEC: JAMES BAY, EASTMAIN (LEPAGETE. T 33131. 27 JUL 1955) GH ISOTYPE

3640 NUBICOLA MACKENZIE9K.K.9 BULL. TORREY BOT. CLUB 36:480. 1909. USA: COLORADO: MINERAL COO t PAGOSA PEAK ( BAKER,C.F., 232. -- AUG 1899) GH ISOTY P E

365 0 NUDATA BOOTTTW. IN WATSON9S.q GEOL. SURV. CALIFORNIA9 BOT. 2z241. 1880. 76 SMITHSONIAN CONTRIBUTIONS TO BOTANY

USA: CALIFORNIA: MARISS (BOLANDERTHON., 2299. -- -_- 1860-1 867) YO TYPE MATERIAL

366. NUDATA VAR. FXRHIOR KUKENfHALqG. IN ENGLER9H.G. A. 9 PFLANZENR. 49 FAM*20:337. 1909.

USA: ARIZONA: WILLOW SPRINGS (PALMERvE.9 546. 0- JUN 1890) DS ISOTYPE

367. NUTANS VAR. JAPONICA FRANCHETTA. AND SAVATIER ,LO9 ENUM. PL. JAP. 2:154. 1879. JAPAN: KANAGAWA (PREFECTURE) : HONSHU ISLAND) 9 YOKOSUKA (SAVATIERvL.9 1404. -- --- 1866- 1874) us 31277 TYPE MATERIAL

368. NUTTALLII DEWEYTC., AMER. J. SCI. ARTS SEft.1~ 43:92. 1842. USA: --: ROCKY MOUNTAINS (NUTTALLTT~T17. ---I GH HOLOTYPE NY ISOTYPE

-0-

369. OAXACANA BAILEY vL.H.9 BOT. GAZ. 25:271. 1898. MEXICO: OAXACA: SIERRA DE SAN FELIPE (PRINGLE,C.Gov 4842. 29 AUG 1894) GH ISOTY PE YO TYPE COLLECTION us 251772 TYPE COLLECTION us 817656 TYPE COLLECTION

370. OBISPOENSIS STACEY1J.W. 9 LEAFL. W. BOT, 1 :240. 1936. USA: CALIFORNIA: SAN LUIS OBISPO COO: SAN LUIS OBISPO, STEINER CREEK (EASTWOODTA. AND H0WELLvJ.T.r 2271. 07 MAY 1936) CAS 235733 HOLOTYPE C AS 237824 ISOTYPE CAS 237908 ISOTYPE DS 270930 I SOTYPE F 866418 ISOTYPE GH ISOTYPE NY ISOTYPE us 1678188 ISOTYPE

371. OBLANCEOLATA KOYAHAtT., WILLDENOWIA 5(3):489. 1969. CHINA: KWANGTUNG: CHUNG TUNGv TAI TSANG, YING TAK (TAK9T.W. AND CHOW,W*K.( 3202. 20 NOV 1926) uc 31 9673 HOLOTYPE

372. OBOVOXDEA CRONQUISTvA. 9 MADRONO 7 :78. 1943. USA: IDAHO: CUSTER COO: STANLEY (CRONQUISTTA. T 2872. 03 JUL 1941) GH ISOTYPE NUMBER 12 77

YO 1220830 TYPE YATERIAL

373, OEDERI VARI ROUSSEAUIANA MARIE-VICTORIN,(FRERE), PROC. & TRANS. ROY. SOC. CANADA SER.3, 23(2), SECT.5:262. 1929 CANADA: QUEBEC: MONTMAGNY COO: L'ESTUAIRE DU ST. LAURENT, BERTHIER-EN-BAS (ROUSSEAU9J. T 24989. 27 JUL 19261 NY TYPE

374. OKLAHOMENSIS YACKENZIE9K.K. T TORREYA 14: 126. 1914. USA: OKLAHOMA: CATALE (BUSH,B.Fm, 993. 22 MAY 1895) YO TYPE MATERIAL NY TYPE

375, OLIGANTHA BOOTT,F., ILL. GENUS CAREX 4:174, PL.589. 1867. CHILE: YAGALLANES: TIERRA DEL FUEGO ("FUEG1A")t ORANGE HARBOR (WILKES EXPLOR. EXPEDoq --- . -- --- 1838-1 842 1 us 30695 ISOTYPE

376, OLlGOCARPA VARo LATIFOLIA GRAYvA. EX TORREY,J.t ANN. LYCEUM NATO HIST. NEW YORK 32415. 1836. USA: NEW YORK: JEFFERSON COO: WATERTOWN (CRAWE,J.B.t --- . ---I GH H OL 0 TY P E

377. OLYYPICA MACKENZIE,K.K., BULL. TORREY BOT. CLUB 43:610. 1916. USA: WASHINGTON: CLALLAY COO : OLYMPIC MOUNTAINS (ELMER, A.D. E. 9 2700. -- JUN 1900) NY SYNTYPE

378. ONUSTA MACKENZIE,K.K.r BULL. TORREY BOT. CLUB 42:618. 1915. USA: TEXAS: TARRANT COO: -- (RUTHfA.9 458. 24 APR 1914) CAS 351152 ISOTYPE NY TYPE us 504456 I SOTYPE

379, OREGONENSIS OLNEYTS~T. EX BAILEY,L.H.T PROC. AMER. ACAD. ARTS 22:73. 1886 ("1887") USA: OREGON: -- (HALLvE.9 605. -- --- 18711 F 455736 SYNTYPE NY SYNTYPE

380. ORHOSTACHYA WIEGANDTK.M.~ RHODORA 24:196. 1922, USA: NEW YAMPSHIRE: GRAFTON COO 2 FRANCONIA, LITTLETON HILL (FAXONpE. AND FAXON,C.E*t --- 27 MAY 1896) GH HOLOTYP E

381. ORONENSIS FERNALD9Y.L 09 PROC. AMER. ACAD. ARTS 37:471, PL.1, FIG.15,16. 1902. USA: MAINE: PENOBSCOT COO: ORONO (FERNALD,M*L.T ---a 30 JUN 1891) GH HOLOTY PE NY ISOTYPE 78 SMITHSONIAN CONTRIBUTIONS TO BOTANY

3820 OXYCARPA HOLYtH.T., AMER. J. SCI. SER.49 20:303. 1905. USA: WASHINGTON: KLICKITAT COO: COLUMBIA (SUKSD0RFtW.N.t 816. 02 JUN 1885) F 96129 TYPE MATERIAL F 211365 TYPE MATERIAL us 27292 TYPE MATERIAL

3830 OXYLEPIS VAR. PUBESCENS UNDERW0ODtJ.K.t AMER. MIDL. NATURALIST 33 :635. 1945. USA: TENNESSEE: CHEATHAM COO: PEGRAM (SVENSON,H.K.t 10469. 12 JUL 1939) NY TY PE COLLECT ION

- P-

384- PACHVCARPA YACKENZIETK.K.V BULL. TORREY BOT. CLUB 43:616. 1916. USA: CALIFORNIA: ALPINE COO: SILVER VALLEY9 BIG TREE ROAD; ALTO 8000 FT. (BREWER9W.H.t 1977. 31 JUL 1863) uc 1060 HOLOTYPE

385. PACHYSTACHYA CHAM1SSOtL.A. EX STEUDEL9E.G.v SYN. PL. GLUM. 2:197. 1855. USA: ALASKA: ALEUTIAN ISLANDS9 UNALASKA (ISLAND) (CHAYI SSOt L *A* T --- . ---I GH ISOTYPE

3860 PACHVSTOMA H0LMtH.T.t AMER. J. SCI. SER.49 20:302. 1905. USA: OREGON: KLAMATH COO: CRATER LAKE NATIONAL PARK? ANNA CREEK CANYON (C0VILLEtF.V.t 1362. 03 SEP 1902) us 415172 TYPE

3870 PADDOENSIS SUKSD0RFtW.N.t ALLG. BOT. 2. SYST. 12:43. 1906. USA: WASHINGTON: YAKIYA COO: MOUNT ADAMS ("PADDO") (SUKSDORF tW.N. 9 1296. 13 AUG 1897) F 223512 TYPE MATERIAL F 1471489 TYPE MATERIAL GH ISOTY PE NY ISOTYPE us 529528 TYPE MATERIAL

3880 PALAWANENSIS KUKENTHALtG. IN ELMERtA.D.E.9 LEAFL. PHILIPP. BOT. 4:1169. 1911 PHILIPPINES: PALAWAN! PUERTO PRINCESA t MOUNT PULGAR (ELMERtA.D.E.9 13146. -- MAY 1911) GH ISOTYP E NY TYPE MATERIAL us 872800 TYPE MATERIAL

3890 PALLESCENS VARo NEOGAEA FERNALDtt4.L.t RHODORA 44:306t PL.712. 1942 CANADA: NEWFOUNDLAND: GANDER RIVER VALLEY 9 GLENWOOD NUMBER 12 79

(FERNALDtMoL. AND WIEGANDtKoM.? 4918. 12-13 JUL 1911) GH HOLO TY P E

390. PANSA BAILEYTL~H~TBOT. GAZO 13:82. 1888. USA: OREGON: CLATSOP COO: CLATSOP (HENDERSONTL.~0) 1482. 31 JUL 1886-20 AUG 1887) DS 490462 SYNTYPE NY SYNTYP E

391 PAPULOSA BOOTTpF. IN GRAYTA~TMEMO AMERo ACAD. ARTS NOSOT6:418. 1859 JAPAN: HOKKAIDO (PREFECTURE): HAKODATE (WRIGHT9C.p ---. -- -4- 1853- 1856 1 GH HOLOTYPE NY ISOTYPE us 31344 ISOTYPE 392 PARCIFLORA BOOTTtF., MEMO AMER. ACAD. ARTS NOS., 6:418. 18590 JAPAN: HOKKAIDO (PREFECTURE): HAKOOATE (WRIGHTTC.? --- 0 ---I us 27275 TYPE MATERIAL

393. PARRYANA DEWEYtC.9 AMER. J. SCI. ARTS SER.11 27:239. 1835. CINADA: --: HUDSON BAY (RICHARDSON9J.t ------I NY TYPE COLLECTION

396. X PATUENSfS LEPAGEtE.9 NATURALISTE CANAD. 89:113~ FIG.1. 1962. CANADA: QUEBEC: UNGAVA BAY? LAKE PATU (DUTILLYtA. AND LEPAGEtEo? 39329: 19 AUG 1961) GH I: SOTYPE

395. PAUCICOSTATA MACKENZIEtK.K., ERYTHEA 8:74. 1922. USA: CALIFORNIA: MARIPOSA COO: YOSEYITE NATIONAL PARK9 YOSEMITE VALLEY (BOLANDER?H.N., 6198. -- JUL 1866) 0s 49738 TYPE COLLECTION F 309086 TYPE MATERIAL YO TYPE YATERI AL NY TYPE COLLECTION

396, PAUPERCULA VAR, BREVISQUAMA FERNALD9M.L.q RHODORA 20:152. 19180 CANADA: QUEBEC: ILE AUX COUDRES (MARIE-VICTORIN,(FRERE), 4021. -- JUN 19171 GH HOLOTYPE

397, PAUPERCULA VAR. PALCENS FERNALD?M.C. T RHODORA 8:77. 19060 USA: MAINE: OXFORD COO: BUCKFIELD (ALLEN?J0A.c 21A. 01 JUL 1878) GH HOLOTYPE

398, PAYSONIS CLOKEYTIOW.~AMERo J. SCI. SER.59 3:90~ PL.2. 1922. USA: WYOMING: TETON COO: GRAND TETON NATIONAL PARK, JACKSON HOLE VALLEY (PAYS0NtE.B. AND PAYSONTL.B., 2224. 06 AUG 1920) GH I SOTY PE uc 905434 HOLOTYPE 80 ShfITHSONIAN COSTRIBUTIONS TO BOTANY

399. PELOCARPA HERMANNTF~J.~RHODORA 39:492. 1937.

USA: UTAH: SUYMIT COO: LAMOTTE PEAK (HERMANNT F.J 09 5983. 15 AUG 1933) CAS 239452 ISOTYPE NY HOLOTYPE

400 a PERCOSTATA HERMANNTF~J~TJ. WASH. ACAD. SCI. 40:282. 1950. MEXICO: CHIHUAHUA: MADERA (MULLERTCml4.r 3520. 27 SEP 1939) CAS 369422 ISOTYPE us 2133207 TYPE

401 a PERGLOBOSA MACKENZ IEI KO K. 9 BULL. TORREY BOT. CLUB 34: 606 1907 USA: COLORADO: SUMMIT COO: MOUNT BALDY, BRECKENRIDGE (MACKENZIETK.K.T 167. -- AUG 1901) YO TYPE MATERIAL NY HOLOTYP E

402 a PERILEIA BLAKEIS~T-T J. ARNOLD ARBOR. 28: 102. 1947. INDONESIA: WEST NEW GUINEA: LAKE HABBEMA; (COUNTRY AS "DUTCH

NEW GUINEA") (BRASS,L.J.p 9583. 1- AUG 19381 A IS OTY P E

403 I. PERLONGA FERNALDTM.L.T PROC. AYER. ACAD. ARTS 43:61. 1907. MEXICO: HIDALGO: TRINXDAD IRON WORKS; ALTO 1585 Me (PRINGLETC~G., 8863. 02 JUN 1904) CAS 155657 ISOTYPE CAS 1930d5 I SOTYPE F 178542 ISOTYPE GH HOLOTY PE YO I SOTYPE NY ISOTYPE us 461358 ISOTYPE

404. PERSTRICTA YACKENZIETK.K.T BULL. TORREY BOT. CLUB 36:479. 19C9. MEXICO: NUEVO LEON: MONTERREY (PRINGLE,C.G., 2630. 05 JUN 1889) GH I SOTYPE NY TYPE

605. PETASATA DEWEYIC., AMER. J. SCI. ARTS SER.1, 29:246. 1836. USA: --: ROCKY MOUNTAINS (DRUMMONDTT.T --- . ---I NY TYPE

406 0 PETRICOSA DEWEYTC., AMER. J. SCI. ARTS SER.19 29:246. 1836. USA: --• ROCKY MOUNTAINS ( DRUMYONDTTo T ------I NY TYPE

407 PETRIEI CHEESEMANTT.F.T TRANS. & PROC. NEW ZEALAND INST. 16:413. 1884. NEW ZEALAND: CANTERBURY (DISTRICT): SOUTH ISLAND, BROKEN RIVER (CMEESEMANTT.F. T --- -- JAN 1883) GH I SOTYPE NUMBER 12 81

408. PHAEOCEPHALA PIPER9C.V.9 CONTR. U.S. NATL. HERB. 11:172. 1906. USA: OREGON: CLACKAMAS Coo: MOUNT HOOD (HALLtE.9 583. 01 AUG 1871) F 455706 TYPE COLLECTION F 1425899 TYPE COLLECTION GH TYPE COLLECTION YO TY PE COLL ECT I ON NY TYPE COLLECTION

409. PHAEOLEPfS HOLM,H.T., AMER. J. SCI. SER.49 17:302. 1904. USA: OREGON: CROOK COO: BEAR BUTTES (LEIBERGqJoB.9 335. 26 JUN 1894) NY ISOTYPE

410. PHALAROIDES VARo PARVULA GROSS9 R. 9 REPERT, SP. NOVo REGNI VEG. 50 :211. 1941 ASGENTINA: JUJUY: TILCARA DEPT.? TILCARA (VENTUR1,S.v 6491. 15 FEB 1927) us 1545831 TYPE

411. PHILOCRENA KRECZETOWICZTVOI09 TRUDY SREDNE-AZIATSK. GOSUD. UNIV.9 SER. 8B9 BOT. 17:75. 1934. USSR: TADZHIKISTAN: PAYIRS; ALTO 82CC FT. (LIPSKY,V.I.T 2732. 19 JUL 1899) NY TYPE MATERIAL

412 PHYLLOMANICA BOOTTTW. IN WATSON, S.9 GEOL. SURV. CALIFORNIA, BOT. 2:233. 1880. USA: CALIFORNIA: MENDOCINO COO: MENDOCINO CITY (BULANDER9H.N.9 4746. -- --- 1866 1 GH HOLOTY P E MO I SOTYPE NY ISOTYPE

413. PHYSOCHLAENA HOLM9H.T.9 AMER. J. SCI. SER.4, 17:317. 1904. USA: ALASKA: YUKON VALLEY9 COAL CREEK HILL (FUNSTON9F.9 139. 30 JUL 1893) F 755322 TYPE MATERIAL MO 920815 TYPE COLLECTION NY TYPE COLLECTION

414. PICTA BOOTT9F. IN GRAY9A.9 MEMO AMER. ACAD. ARTS N.S.9 6:418. 1859. JAPAN: HOKKAIDO 4 PREFECTURE): HAKOOATE (WRIGHTTC. 9 --- . ---I us 31374 TYPE MATERIAL

415. PICTA STEUDEL,E.G., SYN. PL. GLUM. 2m4. ia55. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMOND,T.q ---. ---) NY TYPE COLLECTION

416. PINETORUM VAR. ELAf!OR KUKENTHALvG. IN ENGLER,H.G.A.T PFLANZENR. a2 SMITHSONIAN CONTRIBUTIONS TO BOTANY

49 FAM.20: 195. 1909. MEXICO: OAXACA: SIERRA DE SAN FELIPEt ALTO 10~000FT. (PRINGLETC~G~T4685. 08 JUN 1894) GH ISOTYP E NY ISOTYPE

417. PIPER1 MACKENZIETK.K* IN P1PERqC.V. AND BEATTIETRoK.~ FLo NW. COAST 750 1915. CANADA? BRITISH COLUMBIA: VANCOUVER ISLAND, CEDAR HILL (MACOUNTJOHNT --- 31 MAY 1887) GH ISOTYP E

418. PIRCHINCHENSIS VARm SIMPLEX GROSSvR.9 REPERT. SPo NOV. REGNI VEG. 50:211* 1941. COLOMBIA: --: -- (MUTIS,JoCot KILLIP NO.5715. _- --_ 1760-1888 I us 1563811 TYPE

419. PITYOPHILA MACKENZIETK.K.~ BULL. TORREY BOT. CLUB 40:545. 1913. USA: NEW MEXICO: RIO ARRIBA C0.Z TIERRA AMARILLA

(EGGLESTON, We We I 6605 0 18 APR-25 MAY 1911 1 CAS 383889 I SOTYPE NY TYPE MATERIAL us 660821 TYPE

420. PLANATA FRANCHETvA. AND SAVATIER*Lov ENUM. PL. JAP. 2:126r 555. 1879 0 JAPAN: KANAGAWA (PREFECTURE): HONSHU ( ISLAND) T YOKOSUKA (SAVATIERTL., 20590 -- --- 1866-1874) us 27269 TYPE MATERIAL 421 . PLATVLEPIS MACKENZIEqK.Ko, No AMERo FLo 18:1420 1931. USA: WYOMING: BIG HORN CO.: BIG HORN MOUNTAINS, TEN SLEEP LAKES (WILLIAMS,T.A., 2951. 19 AUG 1897) h(Y HOLOTY PE

422 PLATYPHYLLA CAREYTJ., AMERo Jo SCI. ARTS SER.29 4:23, 1847. USA: NEW YORK: ("NOV. EBOR.") (CAREYTJ~T--- 0 ---I GH TYPE MATERIAL

423 PLECTOCARPA HERMANNTFoJoT LEAFL. Wo BOT. 10:66* 1964. USA: MONTANA: GLACIER CO.: GLACIER NATIONAL PARK, LOGAN PASS, HIDDEN LAKE; ALTO 7300 FTo (HERMANNTF.JOT 18120. 21 AUG 1962) CAS 41 6360 ISOTYPE us 2420276 HOLOTYPE

424. PLUVlCA VARI KOOLAUENSIS KRAUSSvRo T PACIFIC SCI 4: 274. 1950. USA: HAWAII: HONOLULU CO.: OAHU (ISLAND), KOOLAU MOUNTAINS (HOSAKATE~YOT594. 04 JUL 1932) us 2074725 TYPE MATERIAL

42s. PODOCARPA BROWN,R. IN RICHARDSON,Jo IN FRANKLINVJ.9 NUMBER 12

NARR. JOURNEY POLAR SEA 751. 1819. CANADA: --: -- (R1CHARDSONtJ.t ------I GH TYPE MATERIAL

426. POOOGYNA FRANCHETtA. AND SAVAT1ERtL.t ENUM. PL. JAP. 2:131t 557. 1879 JAPAN: TOKYO (PREFECTURE): TOKYO (SAVAT1ERqL.t 1413. -- --- 1866- 1874I us 27270 TYPE MATERIAL

427. PORTER1 0LNEYtS.T.t CARIC. BORo-AMER. 12. 1871. USA: MAINE: PISCATAQUIS COO: MOUNT KINEOt MOOSEHEAD LAKE ( P 0 R T ER 9 T C t --- 28 AUG 1871) GH HOLOTY P E

428. POTOSINA HEMSLEY9W.B.t BIOL. CENTRO-AMER. 3t474. 1885. MEXICO: SAN LUIS POTOSI: SAN LUIS POTOSI (SCHAFFNERtJeGot 546. -- --- 1877) GH ISOTYPE

429. PRAECEPTORIUW MACKENZ1EtK.K.t No AMER. FL. 18:95. 1931. USA: WASHINGTON: KLICKITAT COO: SIMCOE MOUNTAINS, GOLDENDALE (PECKpM.E.7 13. 13 AUG 1917) NY HOLOTY PE

430 PRAEGRACILIS B0OTTtW.t BOT. GAZ. 9:87. 1884. USA: CALIFORNIA: SAN DIEGO COO: SAN DIEGO (SCOTTt(MISSI, ---a -- --- 1880) GH HOLOTYPE

431. PRAINII CLARKE,C.B.t J. LINN. SOC~C801. 36:305. 1904. CHINA: YUNNAN: RED RIVER (HENRYtAot 10839. ---I MO TYPE MATERIAL us 458108 TYPE MATERIAL

432. PRAIREA DEWEYtC. IN W0ODtA.t CLASS-BOOK 601. 414. 1845. USA: MICHIGAN: -- (---t -9-0 ---I GH HOLOTYPE 433 . PRATENSIS DREJERtSoToN., NATURHIST. TIDSSKR. 3:442. 1841. GREENLAND: --: -- (VAHLtJot ------I CAS 105004 TYPE COLLECTION

434 PRATENSIS VARe FURVA BA1LEYtL.H. IN MACOUNTJOHN, CAT. CANADIAN PL. 5:377. 1890. CANADA: BRITISH COLUMBIA: VANCOUVER ISLAND, CEDAR HILL (MACOUNtJOHNt ---.31 MAY 1887) GH ISOTYP E

435. PRATICOLA VAR. SUBCORIACEA HERMANNtF.J.t LEAFC. W. BOT. 8:113. 1957. CANADA: ALBERTA: MOUNTAIN PARK, MCCLEOD RIVER (HERMANNtFoJ. 13453. 26 AUG 1956) 84 SMITHSONIAN CONTRIBUTIONS TO BOTANY

CAS 404488 I SOTYPE us 2265957 HOLOTYPE

4360 PREISSII ESSENSACH,N.VON IN LEHYANNTJ.T PL. PREISS. 2:94. 1846. AUSTRALIA: --: -- (PREISSTL.T 1825. ---I GH SYNTYPE AUSTRALIA: WESTERN AUSTRAL IA: PERTH (PRE ISSTL. T 1861. -- JUL 18391 YO 2002968 SYNTYPE 437. PRESLII STEUDELTE.G.T SYN. PL. GLUY. 2:242. 1855. USA: ALASKA: NUTK,A SOUND ("SINUS NUTKA") (HAENKETT., --- . ---I us 865056 TYPE MATERIAL 438- PRINGLEI BAILEYTL.H.T BOT. FAZ. 17:151. 1892. MEXICO: SAN LUIS POTOSI: HACIENDA DE ANGUSTURA, 100 MILES EAST OF SAN LUIS POTOSI (PRINGLETC.G.~ 3801. 04 AUG 18913 F 105551 TYPE COLLECTION F 1607711 TYPE COLLECTION GH TYPE COLLECTION YO TYPE COLLECTION NY TYPE COLLECTION us 817724 TYPE COLLECTION

439, PRIONPHYLLA HOLM,H.T. T AMER J. SCI SER. 49 14:423. 1902. USA: IDAHO: DIVIDE BETWEEN SAINT JOE AND CLEARWATER RIVERS (LEIBERGTJ~B~T125. 10 JUL 1895) r\lY TYPE COLLECTION

440 PROJECTA MACKENZIE~K~K~TBULL. TORREY BOT. CLUB 35:264. 1908. CANADA: NEW BRUNSWICK: KENT COO: KOUCHIBOUGUAC (FOWLERTJ.T -- --- 18721 GH ISOTYPE NY TYPE COLLECTION

441 PROPOSITA YACKENZIE TKO K. I No AMER. FL. 18: 126 1931. USA: IDAHO: BLAINE COO: SMOKY MOUNTAINS; ALTO 2700 Ma (MACBRIDE9J.F. AND PAYSON*E.B., 3778. 13 AUG 1916) CAS 102638 ISOTYPE GH ISOTYP E NY HOLOTY PE

442. X PSEUDO-FULVA FERNALDpM.L., RHODORA 35:231. 1933. CANADA: NEWFOUNDLAND: PORT AU PORT BAY9 TABLE MOUNTAIN (FERNALD9M.L. AND WIEGAND~K~M~T4258. 16 AUG 1910) GH HOLOTYPE

443. PSEUDOJAPONICA CLARKE9C.B. T BULL. MISC. INFORM. ADD.SERo8:81. 1908. USA: CALIFORNIA: NEVADA COO: SIERRA NEVADA RANGE, DONNER LAKE; ALTO 2750 Mq (HELLER,A.A.T 7187. 17 AUG 1903) F 215918 TYPE COLLECTION NY TYPE COLLECTION SUMBER 12 85

444 PTEROLEPTA FRANCHETTA.T NOUV. ARCH. MUS. HIST. NATO SER.39 8?215. 1896 CHINA: YUNNAN: -- (DELAVAYTR.P.T 4829. 15 JUL 1889) US 1123683 ISOTYPE

445 .I PTYCHOCARPA STEUDEL,E.G., SYN. PL. GLUM. 2:234. 1855. USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (DRUMMONDqT., 424. -- --_ 1832) NY TYPE COLLECTION

446 QURPUREOVAGINATA VARo ITATIAIAE GROSSTR., REPERT. SP. NOV. REGNI VEG. 50:212. 1941. BRAZIL: RIO DE JANEIRO: ITATIAIA (CHASETA., 8283. 17 JAN 1925) !IS 1282178 TYPE 467 PURPURIFERA YACKENZIETK.K.T No AMER. FL. 182253. 1935. USA: TENNESSEE: CAMPBELL COO: CHASKA (BRIGHTTJ., ---a 18 MAY 1923) NY TYPE COLLECT ION

4480 PYCNOTHYSOS KUKENTHALTG~TPHILIPPo J. SCI. 6:60. 1911. PHILIPPINES: NEGROS OCCIDENTAL: MOUNT CANLAON (VOLCANO) (MERRILLTE.D., 543. r.- APR 1910) us 1398830 TYPE MATERIAL

-Q-

449. QUADRIFIDA BAILEY,L.H.T PROC. CALIF. ACAD. SCI. SER.2, 3: 104. 1891 USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA, TUOLUMNE RIVER (ROLANDER,H.Y. T 5046. -- --- 1866) DS 55002 SYNTYPE NY SYNTYP E

450 QUADRIFIDA VARo LENIS BAILEYTL-HoTPROC. CALIF. ACAD. SCI. SER.2, 3:105. 1891. USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA, TUOLUMNE RIVER (BOLANDERTH~N~T5046. -- --- 1866) NY SYNTYPE

451 X QUEBECENSIS LEPAGEtE.9 NATURALISTE CANAD. 91:168. 1964. CANADA: QUEBEC: SAGUENAY COO: ROMAINE RIVER (DUTILLYtA. AND LEPAGE9E.r 41,305. 12 AUG 1963) GH I SUTYP E NY I SOTYP E us 243371 8 ISOTYPE

4520 QUICHENSIS HERMANNTF~J~TJ. WASH. ACADo SCI. 40:284. 1950. GUATEMALA: QUICHE: NEBAJ (SHARP, A. J. T 45144. 07 FEB 1945) us 2133193 TYPE 86 SMITHSONIAN CONTRIBUTIONS TO BOTANY

-R-

453. RACHILLIS YAGUIREt B.9 BRITTONIA 5: 199. 1944. USA: UTAH: GILBERT PEAK (MAGUIREpB. AND MAGUIRE,R.R., 14668. 16 AUG 1936) CAS 325253 ISOTYPE NY TYPE us 1872576 TYPE MATERIAL

4540 RAMOSII KUKENTHAL~G.T REPERT. SP. NOV. REGNI VEG. 8:8. 1910. PHILIPPINES: RIZAL: LUZON ( ISLANDIT MORONG (RAMOSpM.9 BUR. SCI. 1434. -- AUG 1906) us 626608 TYPE MATERIAL

4551 RAYNOLOSII DEWEYTC., AMER. J. SCI. ARTS SER.29 32:39. 1861. USA: IDAHO: FREMONT COO: PIERRE'S HOLE AND HENRY'S FORK

t HAY D EN 9 F V 9 -0- 20 JUN 1860) GH SYNTY P E

456. RETROCURVA DEWEYTC. IN WOODyA., CLASS-BOOK BOT. 423. 1845. USA: NEW YORK: JEFFERSON COO: -- (WOODqW.A.9 --- . ---I GH HOLOTYPE

451. RETROCURVA VAR. COPULATA BAILEYTL.H.T HERB. DISTR. N0.161. 1886. USA: MICHIGAN: INGHAM COO: LANSING (BAILEYTL.H.T 161. 01 JUN 1886) GH IS OTY P E

458. RETROFLEXA MUHLENBERGTH. EX WILLDENOWTC.L.T SP. PC. ED041 4:235. 1805. USA: TEXAS: -- (DRUMYONDTT*T ------I NY TYPE

459. RHYNCHACHAENIUM CLARKEvC. Be IN MERRILL p E. 0.9 PUBL. BUR. SCI. GOV. LAB. 35:5. 1906 ("1905"). PHILIPPINES: BATAAN: MOUNT MARIVELES; ALTO 1100 Me (ELMERTA.D.E.T 6983. -- NOV 1904) MY COTYPE

460. RICHAROSONII BROWNTR. IN RICHARDSONTJ. IN FRANKLINyJ.9 NARR. JOURNEY POLAR SEA 751. 1819. CANADA: --: -- ( R IC HARD SON T J T --- . ---I GH TYPE MATERIAL

461. RfCHAROSONII FOR. EXSERTA FERNALD,M.L., RHODORA 44:290. 1942. USA: ILLINOIS: HANCOCK COO: AUGUSTA (MEADTS~B~T . ---I GH HOLOTYP E

462. ROANENSIS HERYANN~F~J~TCASTANEA 12:113. 1948 ("1947"). USA: TENNESSEE: CARTER COO: ROAN MOUNTAIN: ALT.4700 FT. NUMBER 12 87

(BR0WNtD.M.t 255. 02 AUG 1936) us 213319C HOLOTYPE

463. RORAIWENSIS STEYERMARK, J.A. t FIELDIANAt BOT. 28:67t FIG.7. 19510 VENEZUELA: BOLIVAR: MOUNT SORAIMA; ALTO 2700-2740 M. (STEY ERMARK t J A. 7 5887C 28 SEP 1944) F 1263854 HOLOTYPE NY I SOTYPE

464. ROSAEOIOES HOW€( EoC. IN GORDINIERtHoCo AND H0WEtE.C. 9 FL. RENSSELAER COO 33. 1894. USA: NEW YORK: RENSSELAER COO: LANSINGBURGH (HOWE9E.C.t ---a 30 MAY 1887) NY ISOTY PE

465. ROSEA VARo ARKANSANA BA1LEYtLoH.t BOT. GAZ. 13:87. 1888. USA: ARKANSAS: PULASKI COO: LITTLE ROCK, LA FOURCHE CREEK (HASSEtHoEot --- 0 01 MAY 1886) GH HOLOTY P E NY I SOTYP E

466. ROSEA VARo PUSILLA PECKtCsHmt ANNUAL REP. NEW YORK STATE MUS. 48 132. 1895. USA: NEW YORK: LIVINGSTONE COO: PORTAGE (PECKtCoHot ---. -- JUN 18941 NY TYPE

467- ROSEA VAR. STAMINATA PECK,C.H.t ANNUAL REP. NEW YORK STATE MUS. 47:164. 1894. USA: NEW YORK: OTSEGO Coo: COOPERSTOWN JUNCTION (PECK9C.H.t 5. -- JUN 1893) NY TYPE

468. RUBRO-BRUNNEA VAR. ELINEOLATA MERR1LLtE.D.t LINGNAN SCI. J. 13818. 1934. CHINA: KHANGTUNG: LOH-FAU-SHAN (MOUNTAIN) 9 POK-LO (TSU1tT.M.t 74. -- MAR-APR 1932) A ISOTYPE GH I SOTYPE MO 1260436 ISOTYPE NY TYPE us 1754487 ISOTYPE

469. RUGATA FERNALDt MoL t RHODORA 43: 5459 PLo 671. 1941. USA: VIRGINIA: SUSSEX COO: HOMEVILLE (FERNALD9M.L. AND LONGtBo t 117870 07 MAY 1940) CAS 336836 I SOTYP E F 1489429 ISOTYPE GH HOLOTYP E YO 1306478 ISOTYPE NY ISOTYPE us 2003132 ISOTYPE 88 SMITHSONIAN CONTRIBUTIONS TO BOTANY

470. RUGATA OHWI TJOT ACTA PHYTOTAX. GEOBOT. 1:76. 1932. JAPAN: --• HONSHU (ISLAND), MOUNT DAIMONJI IN YAMASHIRO (OHWI TJOT 29. 04 MAY 1931) F 1463953 TYPE MATERIAL

4710 RUGOSPERMA MACKENZIETK~K., BULL. TORREY BOT. CLUB 42:621, 1915. USA: NEW JERSEY: OCEAN Coo: TUCYERTON (MACKENZIETK~K~T98710 -- MAY 1911) NY TYPE

4720 RUSBYI MACKEYZIETKOK*T SMITHSONIAN MISC. COLLECT. 65(7):20 1915. USA: ARIZONA: YAVAPAI COO: -- (RUSBY,H.H., 859. -- --- 1883) NY TYPE us 30267 TYPE COLLECT1ON

4730 RUTH11 MACKENZIETK.K.T No AMER. FL. 18:112. 1931. USA: NORTH CAROL1 NA: BUNCOMBE Corn: CRAGGY MOUNTAIN (RUTH, A. 9 --_0 -- JUL 1900) YY HOLOTYP E

-S-

474, SAL!NAEFORHIS MACKENZIETK.K.T BULL. TORREY BOT. CLUB 36:477. 19090 USA: CALIFORNIA: MENDOCINO Coo: MENDOCINO CITY (BOLANDERTHON. T 4702. -- --- 18661 CAS 383801 TYPE COLLECTION DS 293480 TYPE COLLECT ION F 30885 TYPE MATERIAL YO TYPE MATERIAL NY TYPE COLLECTION us 29888 TYPE COLLECTION us 319226 TYPE COLLECTION

6750 SALTAENSIS GROSSTROT REPERT. SPo NOV. REGNI VEGo 50:211. 19410 ARGENTINA: SALTA: ROSARIO DEPTo: CAMP0 QUIJANO (VENTURITS.T 8650. -- --- 1929) us 1545752 TYPE

476. SANGUINEA BOOTTTF.T TRANS. LINN. SOC. LONDON 20:137. 1846.

AFGHANISTAN: --: T- (GRIFFITHTW.T 96(KEW 6094). ---I NY TYPE COLLECTION

477. SARAWAKETENSIS KUKENTHALTG~TBOT. JAHRBo SYST. 69:2620 19380 PAPUA AND NEW GUINEA: NORTH-EAST NEW GUINEA (TERRITORY): MOROBE DISTRICT: FIN1STERRE RANGE, MOUNT SARAWAKET (CLEMENSTJ. AND CLEMENSTYOS~T5546. -- MAR 1937) A ISOTYPE

4780 SARTHELLIANA OLNEY9S.T. IN GRAYPA., PROC. AMER. ACAD. ARTS 7:396. 1868. USA: CALIFORNIA: YARIPOSA COO: YOSEYITE NATIONAL PARK, NUMBER 12 89

YOSEMITE VALLEY (BREWER,W.Hoq 1636. -- --- 1863) GH HO LOTY P E

4790 SARTWELLII DEWEYTC.~ AMER. J. SCI. ARTS SER.1, 43:90. 1842. USA: NEW YORK: SENECA COO: JUNIUS (SARTWELLIH~P~T 12. ---I CAS 383407 TYPE COLLECTION CAS 553975 TYPE COLLECT ION NY TYPE COLLECTION

480. SAVAIIENSIS KUKENTHAL9G.q BERNICE P. BISHOP MUS. BULL. 128:24. 1935. WESTERN SAMOA: --: SAVAII ISLAND (CHRISTOPHERSEN, E.9 800. 24 SEP 1929) NY ISOTY PE

481 X SAXENII NMo FERRUGINEA LEPAGE9E.9 NATURALISTE CANAD. 83:142. 1956. CANADA: QUEBEC: JAMES BAY9 FORT GEORGE (DUTILLY9A.G LEPAGE9E. AND DUYAN9M.9 32357. 14 AUG 1954) GH I SOTYPE us 2176493 ISOTYPE

482 SAXIMONTANA YACKENZIETKOK-T BULL. TORREY BOT. CLUB 33:439. 1906. USA: COLORADO: LARIMER COO: FORT COLLINS (BAKER,C.F., ---. _--) MO TYPE MATERIAL NY TYPE MATERIAL

4830 SCABRIOR SARTWELL9H.P. EX DEWEY9C.9 AMER. J* SCI. ARTS SER.2, 8:349* 1849. USA: NEW YORK: YATES COO: PENN YAN (SARTWELLfH.P.9 72. ---) CA S 553877 I SOTYPE

4840 SCABRIUSCULA MACKENZIEIK~K~TBULL. TOQREY BOT. CLUB 35:268. 1908. USA: WASHINGTON: CASCADE MOUNTAINS (CUSICK9W.C. 9 2849. 30 JUN 1902) DS 490735 I SOTYPE YO ISOTYPE NY TYPE us 52 8631 I SOTYPE

485. SCAPOSA HOOKERTJ.D.T BOT. MAG. PL.6940. 1887. CHINA: KWANGTUVG: LOH-FAU-SHAN (MOUNTAIN) ('FORDTC~T---. -- --- 18831) YO 2002967 TYPE COLLECTION

4860 SCHAFFNERI BOOTTrW. IN WATSON,S., PROC. AMER. ACAD. ARTS 18:172. 1883. MEXICO: SAN LUIS POTOSI : SAN LUIS POTOSI (SCHAFFNER. JOG. 9 546. -- --- 1877 1 GH ISOTYPE 481. SCHNEIDERI NELMES,E.y BULL. MISC. INFORM. 201. 1939. 90 SMITHSONIAN CONTRIBUTIONS TO BOTANY

CHINA: YUNNAN: LIKIANG (SCHNEIDERIC~T 2738. 06 AUG 1914) GH HOLOTY PE

488. SCHUEINITZII OEWEYTC-T AMER. J. SCI. ARTS SER.19 9268. 1825. USA: MASSACHUSETTS: BERKSHIRE COO : WILLIAMSTOWN (DEWEYvC. 9 ---. ---) GH HOLOTYP E

489. SCIRPIFORHIS YACKENZIE?K.K., BULL. TORREY BOT. CLUB 35:270. 1908. CANADA: ALBERTA: BANFF NATIONAL PARK? BANFF (MCCALLA9W.C. 9 2348. 28 JUL 1899) rYY TYPE

490. SCIRPOIDEA VAR. GIGAS HOLMTH.T.T AMER. J. SCI. SER.4, 18Z20. 1904. USA: CALIFORNIA: SISKIYOU COO: MOUNT EDDY (PRINGLE?C.G., ---. 19 AUG 1881) F 210109 TYPE COLLECTION NY TYPE COLLECTION

491 SCIRPOIDEA VAR. STENOCHLAENA HOLMTH.T.T AMER. J. SCI. SER.49 18:20. 1904. CANADA: BRITISH COLUMBIA: CHILLIWACK VALLEY (MACOUNT J.M. T 33728. 12 JUL 1901) NY TYPE COLLECTION

492 SCOPARIA VARo CONOENSA FERNALD9M.L.t PROC. AMER. ACAD. ARTS 37:468? PL.1. 1902. USA: MASSACHUSETTS: MIDDLESEX C0.2 MEDFORD (BOOTTTW.T ---. 26 JUL 1865) GH HOLOTY PE

493 SCOPARIA VAR. FULVA BOOTTvW. IN WATSONTS., GEOL. SURV. CALIFORNIA? BOT. 2:237. 1880. USA: CALIFORNIA: ALPINE COO: SILVER VALLEY (6REWERtW.H.t 1969. 31 JUL 1863) GH SYNTYPE

494. SCOPARIA VAR. MINOR BOOTT,F., ILL. GENUS CAREX 3:116~ PL.369. 1862 USA: NEW HAMPSHIRE: WHITE MOUNTAINS (TUCKERMANTE.~ ------I NY TYPE COLLECTION

495. SCOPARIA VARo UONILIFORMIS TUCKERMANTE.? ENUM. CARIC. 17. 1843. USA: MASSACHUSETTS: MIDDLESEX CO.: CAMBRIDGE (TUCKERMAN,E.r --_- ---_1 GH HOLOTY P E

496. SCOPARIA FOR. PfRACUlA FERNALD, M.L.9 RHODORA 23:234. 1921. CANADA: NOVA SCOTIA: YARMOUTH COO: SAND 8 EACH ( FERNALD? M. Lo AND LONGVB.T 20296. 14 JUL 1920) GH HOLOTY PE

497. SCOPARIA VARo SUBfURBINAfA FERWALD1M.L. AND WTEGANDTK.M.T RHODORA NUMBER 12 91

14:116. 1912. CANADA: NEWFOUNDLAND: EXPLOITS RIVER VALLEY, GRAND FALLS AREA, RUSHY POND (FERNALD9M.L. AND WIEGANDTK. Me T 4796. 28 AUG 19111 F 464432 SYNTYPE GH HOLO TY PE NY ISOTYPE

SCOPARIA VARo TESSELLATA FERNALDTM~L. AND WIEGANDTK*M.~ RHODORA 12: 135. 1910. USA: MAINE: WASHINGTON COO: PEYBROKE (FERNALDTM~L.~1464. C8 JUL 19091 GH HOLOTYPE NY ISOTYPE

SCOULERI TORREYTJ., ANN. LYCEUM NAT. HIST. NEW YORK 3:399. 1836. USA: BRITISH COLUMBIA: "WESTERN SHORE OF AMERICA, OBSERVATORY INLET" (ARM OF PORTLAND INLET) (SC0ULERtJ.q 296. ---I NY HOLOTY P E

SEATONIANA BAILEYTL.H., BOT. GAZ. 25327C. 1898. MEXICO: HIDALGO: TULA (PRINGLETC.G.T 7452. 24 JUN 18971 CAS 445940 ISOTYPE GH ISOTYPE us 30 5734 TYPE C OLLEC TION

SHELDON11 MACKENZ1EvKmK.r BULL. TORREY 601. CLUB 42:618. 1915, USA: OREGON: UNION Coo: CLARK'S CREEK (SHELDON,EoP., 8854. 09 SEP 1897) NY I SOTYP E NY TYPE us 528495 TYPE COLLECTION

SICCATA DEWEYTC~TAMER. J. SCI. ARTS SER.19 1@:278o 1826. USA: MASSACHUSETTS: HAMPDEN COO: WESTFIELD 4 DAVIS,Ee ---o ---) NY TYPE COLLECTION

SIMULATA MACKENZIE,K.K., BULL. TORREY BOTo CLUB 34:604. 1907. USA: WYOMING: ALBANY CO.: CHUG CREEK (NELSON,&., 7316. 29 JUN 1900) NY HOLO TYPE

SMALLIANA YACKENZIETK~K.T BULL. TORREY BOT. CLUB 36:484. 1909. USA: GEORGIA: TATTNALL COO: REIDSVILLE (HARPERTR~M.T 2159. 26 APR 1904) us 511177 TYPE COLLECTION

SONOMENSIS STACEYqSeW., LEAFL. We 601. 2:63* 1937. USA: CALIFORNIA: SONOMA CO.: SEBASTOPOL, PITKIN MARSH (HOWELLTJ.~. AND STACEYTJ.W.~ 13042. 06 JUN 19371 CAS 246086 HOLOTYPE CAS 246636 ISOTYPE 92 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DS 258275 TSOTYPE DS 374718 ISOTYPE F 907841 ISOTYPE GH ISOTYP E NY ISOTY PE uc 835699 ISOTYPE us 1736782 ISOTYPE

506- SOPERI RAUPTH.M.T SARGENTIA 6:129, FIG.12. 1947. CANADA: NORTHWEST TERRITORIES: MACKENZIE DISTRICT: BRINTELL LAKE (R4UPtH.M. AND SOPERTJ~H., 9534. 18 JUL 1939) GH HOLOTY P E

507. SPECIFICA BAILEY,L.H.r MEMO TORREY BOT. CLUB 1:21. 1889. USA: CALIFORNIA: ALPINE COO 2 SILVER VALLEY (BREWERtWoHot 1969. 31 JUL 1863) CAS 232289 SYNTYPE us 30329 SYNTYPE

5080 SPECUICOLA HOWELL, J .To t LEAFL. W. BOT. 5: 148. 1949. USA: ARIZONA: COCONINO COO: INSCRIPTION HOUSE (HOWELL~J~T~T 24609. 23 JUN 1948) CAS 342552 ISOTYPE CAS 342553 HOLOTY PE DS 337970 ISOTYPE GH ISOTYPE NY ISOTYPE us 2C06386 ISOTYPE

509. SPRETA BAILEYTL~H., MEMO TORREY BOT. CLUB 1:6. 1889. USA: OREGON: MULTNOMAH COO: SAUVIE ISLAND (COLUMBIA RIVER AT MOUTH OF WILLAMETTE RIVER) (HOWELLTT~J~T--- -- MAY 1880) GH TYPE COLLECTION YO TYPE COLLECTION

STANDLEYANA STEYERMARK, J. A. 9 CEIBA 3:23. 1952. GUATEMALA: JALAPA: AGUACATE (WILL1AMSyL.O.t 13178. 06 JUL 1947) F 1252385 HOLOTYPE

STANTONENSIS JONES,M.E. 9 BULL. MONTANA STATE UN1V.t BIOL. SER. 15:2Cy PL.3. 1910. USA: MONTANA: STANTON LAKE ( WILLIAMS,R.S.', --- 11 AUG 1894) NY TYPE

STELtATA M4CKENZIE,K.K.~ No AMER. FL. 18:226. 1935. MEXICO: HIDALGO: IXMIQUILPAN (ROSE, J. No i PAINTER, JoHm AND ROS€TJ.S. 9 9019. -- --- 1905) NY ISOTYPE us 452499 HOLOTYPE

5130 STELLULATA VAR. CONFERfA CHAPYAN,A.W., FL. So U.S. ED.1, 534. 1860. NUMBER 12 93

USA: FLORIDA: GADSDEN COO: -- (CHAPMANpA.W.9 --- . ---I NY TYPE

514. STENOPHYCLA VARI DESERTORUH LITVINOVTD.I.T ALLGO BOT. Z. SYST. 5( BEIH. 1): 56. 1899. USSR: TUSKESTAN: -- (LITVINOV,b.I., 1530 -- --- 1897) us 616142 TYPE MATERIAL 515. STENOPTERA MACKENZIE9K.K.9 ERYTHEA 8:28. 1922. USA: CALIFORNIA: COS ANGELES COO: SAN ANTONIO MOUNTAINS9 ICE HOUSE CANYON (JOHNSTONTI.M.T 1505. 31 JUL 1917) DS 83850 ISOTYPE NY TYPE

5160 STERILIS VAR. EXCELSIOR BAILEYTLOH. T BULL TORREY BOT. CLUB 20:424. 1893. USA: NEW YORK: SENECA COO: JUNIUS (SARTWELL9H.P.9 35. ---I NY I SOTY P E

517. STEUDELII KUNTHTC~S.9 ENUM. PL. 2:480. 1837. USA: OHIO: MXAMI RIVER VALLEY (FRANK9J.C. 9 --- . -- --- 1835) NY TYPE COLLECTION

518. STEVERHARKII STANDLEYTP.C.T PUBL. FIELD MUS. NAT. HIST.? BOT. SER. 23: 196. 1947. GUATEMALA: HUEHUETENANGO: CERRO HUI TZ (STEYERMARK, J. A. 9 48542 14 JUL 1942) F 1129096 HOLOTYPE 519. STIPATA VAR. LAEVIVAGINATA KUKENTHALvG. IN ENGLERvHeG.A.9 PFLANZENR. 49 FAM.20: 172. 1909. USA: NORTH CAROL1 NA: BUNCOYBE COO: BILTMORE (NEAR ASHEVILLEI (BILTMORE HERBARIUM9 262A. 28 MAY 1897) F 813737 TYPE COLLECTION GH TYPE COLLECT ION MO TYPE COLLECTION NY TYPE COLLECTION

52 8 ST ATA VAR. NAXIHA CHAPMAN9A.W. EX BOOTT9F.9 ILL. GENUS CAREX 3:1219 PL.391. 1862. USA: FLORIDA: APPALACHICOLA (CHAPYAN,A.W., --- . -0-1 us 969091 TYPE COLLECTION

521. STIPATA VARo SUBSECUTA PECKTC.H.T ANNUAL REP. NEW YORK STATE MUS. 48: 128. 1895. USA: NEW YORK: RENSSELAER COO: BERLIN (PECK,C.H.p ---a -- JUN 1894) WY TYPE COLLECTION

522. STIPATA VAR. UBERIOR MOHR9C.p CONTR. U.S. NATL. HERB. 6:417. 1910. USA: ALABAMA: MOBILE RIVER (MOHRvC.9 --- 26 APR 1897) NY ISOTYPE 94 SMITHSONIAN CONTRIBUTIONS TO BOTANY

5230 STRAHINEA VARo CUMULATA BAILEY, L .He T YEY. TORREY BOT. CLUB 1: 23. 1889.

CANADA: NEW BRUNSWICK: KENT COO: -0 (FOWLERTJ., ---a

-0 JUL 1870) GH SY NTY P E YO SY NTYPE NY SYNTYP E

STRANINEA VARo ECHINODES FERNALD,M.L., PROC. AMER. ACAD. ARTS 37:474, PL.20 1902. CANADA: ONTARIO: LAYBTON COO: WYOMING (MACOUNvJOHN, 26624. 24 JUN 1901) GH SYNTYPE

STRAWINEA VARo RENIFORMIS BAILEY,L.H., MEM. TORREY BOT. CLUB 1~73. 1889 USA: MISSISSIPPI: OKTIBBEHA COO: STARKVILLE (TRACY,S.M.t 17. 23 MAY 1888) NY SYNTYPE

STRICTA VARo XEROCARPA BRITTON,N.L., BULL. TORREY BOT. CLUB 22:222. 1895. USA: NEW YORK: STEUBEN COO: PRATTSBURGH (WRIGHT,S.H., .--. ---) NY TYPE COLLECTION

STYLOFLEXA BUCKLEYtS.B., AMER. J. SCI. ARTS SER.1, 45:174. 1843. USA: NORTH CAROLINA: MACON COO: MOUNTAINS (BUCKLEY,S.B., ---a --- ) YO TYPE COLLECTION NY TYPE COLLECT ION

STYLOSA VARe VlRENS BAILEYpLoH., PROC. AMER. ACAD. ARTS 22:79. 1886 ("1887"). USA: OREGON: MULTNOMAH COO: SAUVIE ISLAND (COLUMBIA RIVER AT MOUTH OF WILLAMETTE RIVER) (HOWELLIT~J., --- -- MAY 1880) GH SYNTYPE YO SYNTYPE

SUB-BRACTEATA MACKENZIE,Y.K., BULL. TORREY BOf. CLUB 43:612. 1916. USA: CALIFORNIA: ALAYEDA COO: OAKLAND (BOLANDER,H.N.r ---. _- _-- 1860 CA.1 GH ISOTYPE NY HOLOTY PE us 28683 ISOTYPE SUBFUSCA ROOTTTW. IN WATSONTS., GEOL. SURV. CALIFORNIA, BOT. 2:234. 1880. USA: CALIFORNIA: LAKE TAHOE TO BEAR VALLEY (KELLOGG,A., ---. ---) GH ISOTYPE

SUBIWPRESSA CLOKEY,I.W., RHODORA 21:84. 1919. NUMBER 12 95

USA: ILLINOIS: MACON COO: -- (CLOKEYTI.W.T 2338. 06 AUG 1915) C AS 162423 ISOTYPE uc 9054 33 HOLOTY P E

5320 SUBNfGRICANS STACEYTJ.W.T LEAFL. W. BOT, 22167. 1939. USA: CALIFORNIA: TUOLUMNE COO: MOUNT DANA (HOWELLTJ~T~T14519. 11 AUG 1938) CAS 259816 HOLOTYPE us 1765699 ISOTYPE

5330 SUBORBICULATA MACKENZIE9K.K. IN ABRAMS,L.r ILL, FL. PACIFIC STATES 1:338. 1923. USA: WASHINGTON: KL ICKITAT COO : -- (SUKSDORF rW.N. T 1315. -- JUL 1883) NY TYPE

534. SUBTRANSVERSA CLARKE9C.B. T PHILIPP. J. SC1 2:108. 1907. PHILIPPINES: BENGUET: LUZON I ISLAND), PAUAI (MERRILLqE.D.1

473C. -9 OCT-NOV 1905) YY TYPE YATERIAL us 710428 TYPE MATERIAL

5350 SUKSDORFII KUKENTHALTG., REPERT. SP. NOV. REGNI VEG. 16:434. 1920. USA: WASHINGTON: YAKI MA CO. : MOUNT ADAMS ( loPADDO1g 1 (SUKSDORFTW-N., 7383. 27 AUG 1912) CAS 152864 ISOTYPE CAS 243333 I SOTYPE DS 171455 ISOTYPE 0s 269641 ISOTYPE GH ISOTYPE YO 95212 TYPE COLLECTION NY 1 SOTYPE us 1437926 TYPE COLLECTION

5360 SUKSDORFII VARo OVALIS KUKENTHALqG., REPERT. SP. NOV. REGNI VEG. 16 :434. 1920. USA: WASHINGTON: YAKI MA COO : MOUNT AOAMS ( "PADDO") (SUKSDORF,W.N., 5259. 21 AUG-20 SEP 1905) DS 269625 SYNTYPE NY SYNTYPE

537. SURCULOSA RAYMONDTM.T MEMO JARD. BOT. MONTREAL 52:22. 1959. CHINA: KUANGTUNG: SIN-FUNG DISTRICT: SAI-LIN-SHAN VILLAGET NGOK SYING SHAN (TAAMTY.W.T 502. 1-16 APR 1938) A HOLOTYPE

538. SYCHNOCEPHALA CAREYTJ~TAYER. J. SCI. ARTS SER.2, 4:24. 1847. USA: NEW YORK: JEFFERSON COO: WATERTOWN (CAREYTJ~T--- . ---I GH ISOTYPE 96 SMITHSONIAN CONTRIBUTIONS TO BOTANY

-T-

5390 TACHIRENSIS STEYERMARKtJoAot FIELDIANA, BOT. 28:68t FIG080 1951. VENEZUELA: TACHIRA: COLOMBI AN-VENEZUELAN BOUNDARY t PARAMO DE TAMA; ALTO 3045-3475 M. (STEYERMARK9J.A.t 573670 15 JUL 1944) F 1263858 HOLOTYPE us 1932033 ISOTYPE

5400 TAHITENSIS BROWNTF.B.H.~ OCCAS. PAP. BERNICE PAUAHI BISHOP MUS. 9(4):8. 1930. FRENCH POLYNESIA: --: SOCIETY ISLANDS t TAHITI t MOUNT OROHENA (MACDAYIELStL .Hot 1542. 15 YAY 1927) A ISOTY P E

561. TAHAKXI K0YAMAtT.t BULL. ARTS SCI. DIV. RYUKYU UNIV. 3:75. 1959. RYUKYU ISLANDS: OKINAWA (PREFECTURE): OKINAWA (ISLANDIT YONA OKINAWA (ISLAND), YONA EXPERIMENTAL FOREST OF RYUKYU UNIVERSITY; (COUNTRY AS "JAPAN") (K0YAMAtT.t ---a 23 NOV 1958) NY HOLOTY PE

542, TAHANA STEYERMARKTJ~A~TFIELDIANAt BOT. 28:70. 1951. VENEZUELA: TACHIRA: COLOMBIAN*VENEZUALAN BOUNDARY9 PARAMO DE TAMA; ALTO 3045-3475 Mo (STEYERYARKtJ0A.t 57401. 15 JUL 1944) F 1263857 HOLOTYPE 543. TENERA DEWEYTC.~ AMER. J. SCI. ARTS SER.19 8:97. 1824. USA: MASSACHUSETTS: BERKSHIRE COO: WILLIAMSTOWNt SADDLE MOUNTAIN (DEWEYtC. T --- 20 JUN ---- 1 GH HOLO TY PE

544. TENERA VARm RICH11 FERNALDtMoLo, PROCo AMER. ACAD. ARTS 37:475T pL.2. 1902. USA: MASSACHUSETTS: YIDDLESEX COO : MIDDLESEX FALLS (R1CHtW.P. t --- 05 JUN 1894) GH HOL OTY P E

5450 TENERAEFORWIS MACKENZIE,K.K.T BULL. TORREY BOT. CLUB 4326090 1916. USA: CALIFORNIA: BUTTE Coo: JONESVILLE; ALTO 5100 FT. (HALLpHoMot 97810 25 JUL 1914) NY HOLOTY P E

546. TENTACULATA VARo ACTIOR BOOTTTF.T ILL. GENUS CAREX 2:94r PL.278. 1860. USA: NEW YORK: YATES COO: PENN YAN (SARTWELLTH.P.~ 1380 ---I CAS 553943 TYPE COLLECTION

547- TENUIFLORA VARo SETACEA KUKENTHALtG. IN ENGLERTH~G~A~TPFLANZENR. NUMBER 12 97

49 FAM.20:224. 1909. USA: MICHIGAN: INGHAM COO: LANSING (WHEELERtCoF. 9 28 JUN 1890) GH ISOTYPE

548. TERETIUSCULA VAR. AMPLA BA1LEYtL.H.t MEMO TORREY BOT. CLUB 1:53. 1889 USA: OREGON: BAKER COO: HEAD OF BURNT RIVER (CUS1CKtW.C.t 1331. -- JUL 1886) NY SY NTY P E

549. TERRAE-NOVAE FERNALDtMoL.? RHODORA 44:290, PL.711. 1942. CANADA: NEWFOUNDLAND: SAINT JOHN BAY? SAINT JOHN ISLAND (FERNAL0,M.L.t W1EGANDtK.M.; LONGyB.; GILBERT-JR.,F.A. AND HOTCHKISS,N., 27657. 31 JUL 1925) GH HOLOTYPE

550. TETSUOI 0HWIpJ.q MIX. REP. NATL. SCI. MUS. 5:2t PL.2. 1952. RYUKYU ISLANDS: OKINAWA (PREFECTURE): OKINAWA (ISLAND); (COUNTRY AS "JAPAN") (AMAN0,T.t 6358. -- MAY 19511 us 2092356 TYPE MATERIAL

5511 THURBERI DEWEYpC. IN TORREYtJ. IN EM0RYtW.H.t REP. U.S. MEX. BOUND. SURV.7 BOT. 2(11:232. 1859. MEXICO: SONORA: MABIBI (THURBER9G.t --- 0- JUN 1850) GH HOLOTY P E

552. TOJQUIANENSIS STANDLEYtPoC. AND STEYERMARK,J.A.q CEIBA 4:64. 1953. GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, BETWEEN TOJQUIA AND CAXIN (STEYERMARKtJ*A.t 50150. 06 AUG 1942) F 1129085 TYPE MATERIAL F 1129086 TYPE MATERIAL

553. TOHPKINSI H0WELLtJ.T.t LEAFL. W. BOT. 9:185. 1961. USA: CALIFORNIA: FRESNO COO: KINGS RIVER CANYON? COPPER CREEK TRAIL (H0WECLtJ.T.t 35333. 06 JUN 1960) CA S 428953 HOLOTYPE CAS 429306 ISOTYPE NY I SOTYPE us 2604281 ISOTYPE

554. TOREAOORA STEYERMARKTJ.A.r PHYTOLOGIA 9:338* 1954, ECUADOR: AZUAY: TOREADOR: ALTO 3810-3930 Me (STEYERMARK,J.A., 53095. 15 JUN 1943) F 1266183 TYPE MATERIAL NY I SOTYPE 555 . TORTA VAR. STAMINATA PECKpCeH., ANNUAL REP. NEW YORK STATE MUS. 46: 131. 1893. USA: NEW YORK: ONEIDA COO: TABERG (PECKtC.H.9 ---. -- JUN 1892) NY TYPE COLLECTION 98 SMITHSONIAN CONTRIBUTIONS TO BOTANY

556. TOWNSENDII MACKENZIE,K.K.t No AMER. FL. 18:111. 1931. MEXICO: CHIHUAHUA: COLONIA GARCIA; ALTO 2250 M. (TOWNSEND,C.H.T. AND BARBER~COM~T157. 21 JUL 1899) CAS 351161 ISOTYPE NY ISOTYPE us 568126 HOLOTYPE

557. TRACY1 YACKENZIE,K.K.T ERYTHEA 8:41. 1922. USA: CALIFORNIA: HUMBOLDT COO: BALD MOUNTAIN (TRACYTJ~P~T4547. 04 JUL 1914) NY TYPE

558. TRIANGULARIS BOECKELERTJ.~. 7 FLORA 39:226. 1856. USA: TEXAS: -- (0RUYMONDtT.T --- . ---I NY I SOTY P E

559, TRIBULOIDES VARm SANGAMONENSIS CLOKEYtIoW., RHODORA ?1:84. 1919. USA: ILLTNOI~: MACON rn.: -- (CLnKFY.1-Wet 2364- 37 AUG 1915) uc 5,"3+41 LlnLOTYPE

560o X TRICHINA FERYALD, HoLot 9HOr)OSA 35z2l.9. 19330 USA: MAINE: ARnOSTOOK CO.: FORT FAIRFIELO (WICLIAMS,E.F.; C~~~LINSTJ.F.ANn FERYILD~M*L.T IIC. I9 JUL 1902) GH HOLOTYPE

561. TRICHOPHYLLA NELYESTE.~ MEMO YUS. NATL. HIST. NATO9 SER. Bt BOT. 4:lCb. 1955. VIET-NAY? NORTH: TONKIN: CHAPlJ: (rOUNTRY AS "TNDOCHINA") foCTCLOTqDoA+ c324. -- JUI. :"?I GH HT)LOTYDE

562. TRISPERYA DEWEYtC.9 AMER. J. SCI. 4qT5 SER.1, 9:ha. 1825. USA: MASSACHUSETTS: -- (DEWEYTC., -0- . ---I NY 1 SOTY PE

563 TRISPERWA VAR. BILLINGS11 KNIGH1,O.W. T RHOOORA 8 :185. 19C6. USA: M4INE: SOMERSET COO: PLEASANT RIDGE9 JEWETT BROOK BOG (WARE9R.A.t ROLLINStS. AlVD KVIGHTTO.W., 5066. G5 JUL 19C6) GH I SOTYPE

5640 TSOI MEqRILLT F.D. 4N3 CH?INvY.K. 9 SUYYATSEYIA 2:207. 1935. CHIYA: KW4NGTUNI;: YAIN4N (ISLAY@IT DlJhlG KA TO WEY FA SHI (CHUIVtY-K. 4Nr) TSOTC.L.T 43683. -- --- 1932-1 933 1 GH I SnTYPE NY ry PF us 167'1 2C TVPE "4TESI 4L

565. TUHULICOLA MACKENZ1EtK.K. T BULL. TORREY BOT. CLUB 34:154. 1907. USA: CALIFORNIA: ALAYEDA COO: LAKE TEYESCAC (BIOLETTI TF-T. T 1. 25 JUN 18931 NY HOLO TY P E

566- TUNIHANENSIS STANDLEYvP .C. AND STEYERMARKTJ. A. 9 CEIBA 4:65. 1953. NUMBER 12 99

GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUYATANES, TUNIMA ( STEYERMARK?J.A. T 48334. 07 JUL 1942) F 1128966 TYPE 567. TURGESCENS TORREYTJ.~ ANN. LYCEUM NATO HIST. NEW YORK 3:419. 18360 USA: LOUISIANA: ORLEANS PARISH: NEW ORLEANS (INGALLSTTOT--to --- ) h(Y TYPE

5680 TURUMIQUIRENSIS STEYERMARKtJoAoq FIELDIANA, BOTo 28:70, FIG.7. 1951. VENEZUELA: SUCRE: CERRO TURUMIQUIRE, RIDGE DIVIOING HEADWATERS Of RIC) YANZANARES AND RIO DE AMANA; ALTO 1900-2000 Ma (STEYERMARKTJOA~T62705. 10 YAY 1945) F 1266150 HOLOTYPE GH ISOTYPE NY ISOTYPE us 1933688 I SOTY PE

- U-

569. ULTRA BA1LEYtLoH.r PROC. AYERo ACAD. ARTS 228830 1886 ("1887"). USA: ARIZONA: COCHISE COO: HUACHUCA ( LEMMON, J.G.9 2. 21 JUN 1882) DS 63991 ISOTYPE DS 64032 I SOTY P E

5710 UNCOMPAHGRE KELSO9L.r BIOL. LEAFLO 38:l. 1947. USA: COLORADO: LA PLATA COO: LA PLATA YOUNTAINSt MOUNT HESPERUST GOLD KING MINE (KELSOTL., 6058. 03 JUL 1947) GH I SOTYPE

572, UNOERHOODII BRITTDN*N.L.q TORREYA 5: 100 1905. JAMAICA: --: SALT HILL MARSH (UNDERWOOD,L.M., 158. 29 JAN 19031 YY TYPE

573. UNILATERALIS MACKENZIETK~K~,ERYTHEA 8243. 1922. USA: CALIFORNIA: HUYBOLDT Coo: ALTON (TRACY, JoPot 3783. 21 JUL 1912) NY TYPE 100 SMITHSONIAN CONTRIBUTIONS TO BOTANY

- V-

576. VAGANS H0LMrH.T.~ AMER. J. SCI. SER.4, 17:301. 1904. USA: OREGON: HARNEY COO: STEEN MOUNTAIN, ANDREWS (LEIBERG,J.B.* 2558. 10 JUL 1896) NY I SOTYP E

5750 VAGINATA VAR. ALTO-CAULIS DEWEY,C.* AMER. J. SCI. ARTS SER 0 2 41: 227. 1866. USA: NEW YORK: GENESEE C0.Z BERGEN (PAINE,J.A., --to ---I GH HOLOTY PE

576. VALLICOLA DEWEY,C.r AMER. J. SCI. ARTS SER.2, 32:40. 1861. USA: WYOYING: TETON COO: SNAKE RIVER, JACKSON HOLE VALLEY (DEWEYIC.~ 1Q. 18 JUN 18601 GH HOLOTYPE

5770 VENOSIVAGINATA STANDLEY,PoC. AND STEYERMARK,J.A.T CEIBA 4267. 1953. GUATEMALA: HUEHUETENANGO: SIERRA DE LOS CUCHUMATANES, CERRO HUTTZ, BETWEEN MIMANHUITZ AND YULHUITZ; ALTO 2600 M. (STEYERYARK,J.Ao, 48554. 14 JUL 1942) F 11 2 9094 HOLOTY PE GH I SOTYPE

578. VERNACULA VAR. HOBSONII YAGUIRE,Boq BRITTONIA 5:199. 1944. USA: UTAH: BEAR RiVER RANGE, WHITE PINE LAKE (MAGUIRETB.~ H0BSONtD.A. AND MAGUIRE+R.R. 9 14013. 16 JUL 1936) CAS 348507 ISOTYPE GH ISOTYPE NY HOLOTYPE us 1872573 ISOTYPE

5?9. VESfCARIA VAR. JEJUNA FERNALD,M.L.t RHODORA 3:530 1901. USA: MAINE: AROOSTOOK COO: MADAWASKA LAKE (WILLIAMS,E.F.~ ---. -- AUG 1900) GH HO LOTY P E

5800 VESICARIA VARo LAURENTIANA FERNALDqY.L.9 RHODORA 35:232. 1933. CANADA: NEWFOUNDLAND: SAINT JOHN'S (FERNALD7M.L.; LONGvB. AND FOGG-JR.,J.M., 1474. 31 JUL 1929) GH HOLOTYPE us 205@647 I SOTYPE

5810 VESTITA VAR. KENNEDY1 FERNALDIM~L~T RHODORA 2:170o 1900. USA: MASSACHUSETTS: MIDDLESEX COO: WILMINGTON, SILVER LAKE (KENNEDYIGSG.~ --- 11 JUN 1899) GH HOLOTYPE

582. VEXANS HERYANNIF-J., RHODORA 57:156. 1955. NUMBER 12 101

USA: FLORIDA: HENDRY COO: CLEWISTON (DEAMTC.C.T 61177. 19 M4R 1941) US 2231425 HOLOTYPE

5830 VICARIA BAILEY9L.H. T HEM. TORREY BOT. CLUB 1:49. 1889. USA: OREGON: -- (HALLTE.T ------.II 1871 1 r,H HOLOTYPE

5840 VIOLACEA CLARKE,C.B., BULL. MISC. INFORM. ADD.SER.8:87. 1908. USA: CCILORADO: ROCKY MOUNTAINS; 141. 39-41 No (HALLTE. AND HARBOURTJ.P., 587. -- --- 1862) YO TY PE COLLECT ION

5850 VIRIOlOR MACKENZ1EvK.K. IN ABRAHS,L.T ILL. FL. PACIFIC STATES 1.: 331. 1923. USA: WASYINGTON: OKANOGAN COO: SHEEP MOUNTAINS ( EGGLEST0NvW.W. 9 3329. 30 JUL-01 AUG 1916) IJ S 886234 TYPE 5860 VlRIDULA FOR. PVGMAEA LEPAGE9E.p NATURALISTE CANAD. 89:115. 1962. CANADA: QUEBEC: UNGAVA BAY, SWAMPY BAY (DUTILLYTA. AND LEPAGE9E.r 39274. 16 AUG 1961) GH ISOTYP E

5870 VITIENSIS ST.JOHNTH.~ PACIFIC SCI. 1:116~ FIG.1. 1947. FIJI: --: VITI LEVU (ISLAND) (STIJOHNTH.T 18330. 18 AUG 1937) us 1967819 ISOTYPE 5880 VITREA HOLMTH.T.T AMER. J. SCI. SER.4, 17:302. 1904. USA: CALIFORNIA: RIVERSIDE COO: PALM SPRINGS (PARISHTS.B.~ 4144. 04-13 APR 18961 F 89120 TYPE HATERIAL YO ISOTYPE us 279151 ISOTYPE 589- VULPINOlOEA VARo PYCNOCEPHALA HERMANN9F.J.v RHOOORA 38:363. 1936. USA: MICHIGAN: EMYET COO: BIG STONE BAY (HERHANNvFoJ., 64C8. 14 AUG 1934) GH HOLOTYPE

-W-

5900 UAHUENSIS VARo RUBlGfNOSA KRAUSSTR., PACIFIC SCI. 4~2579F16.2934-Do 1950. USA: HAWAII: KILAUEA IKI (BEANqR.S.t HOSAKATE~Y. AND ST.JOHNTH.T 11228. 21 DEC 1931) us 2074653 TYPE COLLECTION 591. HATSONI OLNEY9S.T. IN WATSONTS., BOT. U.S. GEOL. EXPLOR. 40TH PAR. 370. 1871. USA: NEVADA: ORYSBY COO: CARSON CITY (WATSONTS., 1246. 102 SMITHSONIAN CONTRIBUTIONS TO BOTANY

-- MAY 1868) GH HOLOTY PE

5920 WEROERMANNIT GROSSTR. IN WERDERMAYNtE.1 NOTIZBL. BOT. GART. BERLIN-DAHLEM lot7630 1929. CHILE: LLANQUIHUE: ALTO 700 M. (WERDERMANNTEOT 1687. -- MAR 1925) YY TYPE COLLECTION

5930 WHIfNEYI OLNEY,S.T. IN GRAY,A., PROC. AMERo ACAD. ARTS 7:394. 1868. USA: CALIFORNIA: TUOLUMNE COO: YOUNT DANA; ALTO 1200C FT. (BOLANDERTHON., 5086. -- --- 1866) YO SYNTYPE NY SYNTYPE USA: CALIFORNIA: MARIPOSA COO: YOSEMITE NATIONAL PARK, YOSEYITE VALLEY (BOLANDERIH.N.~ 6198. -- --- 1866 1 F 309085 SYNTYPE F 309086 SYNTYPE YO SYNTYPE NY SYNTYP E USA: CALIFORNIA: NEVADA COO: SODA SPRINGS; ACT. 9000 FT. (BREWERTU.H.T 1778. 04 JUL 1863) YO SYNTYP E NY SY NTY PE

5940 WIEGANDII MACKENZIETK.K*T No AMER. FL. 18:lCS. 1931. CANADA: NEUFOUNDLAND: RAY OF ISLANDST HUMBER ARM, CURLING (FERNAL0,M.L. AND WIEGANDTK.M.T 2776. 21 JUL 1910) GH ISOTYPE

5950 W1LKESII TORREYTJ. IN WILKESTC. 9 U.S. EXPLOR. EXPED. 17:477r PL.17. 1854. USA: CALIFORNIA: SACRAMENTO RIVER (WILKES EXPLOR. EXPEDOT ---a -_ --- 1 8 3 8- 1 842 1 NY TYPE COLLECTION

596. WILLDENOVII VARm PAUCIFLORA 0LNEYvS.T. EX BAILEY,LoH. IN COULTER9J.M.r CONTR. U.S. N4TL. HERB. 2:482. 1894. USA: TEXAS: HARRIS COO: HOUSTON (HALLTE.~--- . -- -0- 1872) GH TYPE COLLECTION \

597. WILLOENOWII VARI HEGARRHYNCHA HERYANNTFoJ.~ AMER. MIDL. NATURALIST 51:277, 1954. USA: GEORGIA: JASPER CU.: OCMULGEE (SMITH,S.J. AND DUNCAN,W.H., 4872. 06 APR 1949) us 2133191 TYPE COLLECTION 598- WILLfAMSIT BRITTONtN*L., BULL. NEW YORK BOT. GARD. 2:159. 1901. CANADA: YUKON TERRITORY: DAUSON (WILtIAMS,R.S., ---a 12 JUN 1899) NY TYPE NUMBER 12 103

5990 UOODII DEWEYTC., AMER. J. SCI. ARTS SER.2, 2:249. 1846. USA: NEW YORK: JEFFERSON COO: PERCH LAKE, PERCH RIVER (CRANE, 1.B. AND WOOD,W.A., ---.I.---I GH HOLOTY PE

600. WRIGHTI! DEWEYTC. IN TORREYTJ. IN EYORY,W.H., REP. U.S. MEX. BOUND. SURV.r BOT. 2(1):232. 1859. USA: TEXAS: -- (WRIGHT,C., 1561. -- --- 1850 1 NY TYPE COLLECTION

- X-

601. X XANTHINA FERNALDTM.~., RHODORA 35:230. 1933. CANADA: NEWFOUNDLAND: MAIN RIVER (FERNALD,M.L. AND LONGTB.~ 1455. 27 AUG 1929) GH HOLOTY PE us 2050636 ISOTYPE

602. XANTHOCARPA VAR. ANNECTANS BICKNELLTE.P.T BULL. TORREY BOT. CLUB 23:23. 1896. USA: NEW YORK: LONG ISLAND, RICHMOND VALLEY (BRITTONTN.L.T ---. 06 JUI. 1895) NY TYPE

6630 XERANTICA BAILEYTL~H~TBOT. 642. 17:151. 1892. CAN4OA: SASKATCHEWAN: FILE HILLS; 50.5N. 9 1041. ( MACOUNTJOHN, --- 04 JUL 1879) GH SYNTYPE NY SYNTYP E

604. XEROCARPA WRIGHTTSOH. IN DEWEYTC.? AMER. J. SCI. ARTS SER.21 42:334. 1866. USA: NEW YORK: STEUBEN COO: PRATTSBURGH (WRIGHT,S.H.r ---. ---) NY TYPE COLLECTION

- Y-

6050 YUKONENSIS BRITTON9N.L.1 BULL. NEW YORK BOT. GARD. 2:159. 1901. CANADA: YUKON TERRITORY: BONANZA RIVER (WILLlAMS,R.S., ---a 18 JUN 1899) NY TYPE

- Z-

606. ZIZANIAEFOLIA RAYMONDvM. T MEMO JARD. BOT. MONTREAL 53:36. 1959. 104 SMITHSONIAN CONTRIBUTIONS TO BOTANY

CHINA: YUNNANI: -- (TSA1,H.T.p 62809. -- 0-w 1934 1 4 HO LO f Y P E NUMBER 12 105

AUTHOR INDEX

AUTHOR DATE SPECIES

BAILEY,L.H. 1935 ABLATA 1889 ACUTI NA 1889 ALBIDA 1889 ALMA 1893 AUSTRO-CAROLINIANA 1886 *BRONGNIARTII 1889 CALI FORNICA 1884 *CANEXENS 1920 CHIKUNGANA 1889 COMMUNI S 1889 DONNE LL -SM ITHI I 1893 DUR IFOL IA 1896 EGGERTI I 1889 ELEOCHARIS 1886 ENGELMANN I 1889 *FEST IVA 1893 FETA 1889 *FOENEA 1886 *GAYANA 1889 *GR ISEA 1884 HALL IANA 1896 HASSEI 1889 *HOOD1 I 1896 IDAHOA 1889 ILLOTA 1886 INOPS 1893 INTERIOR 1889 JONES II 1920 KUL INGANA 1898 MADRENS IS 1886 *MARC1 DA 1916 MARIPOSANA 1889 *MI LIARI S 1892 MONTANENSI S 1896 *NEBRASKENSIS 1885 NERVINA 1898 OA XAC ANA 1888 PANSA 1892 PRINGLE I 1891 QUADRI F IDA 1891 *QUADRIF IDA 1886 *RETROCURVA 1888 *ROSEA 1898 SEATONIANA 1889 SPECIFICA 1889 SPRETA 1893 *STERILI S 1889 *STRAM1 NEA 1889 *STRAMINEA 106 SMITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPECIES

BAILEY9L.H. 1886 *STYLOSA 1889 * TE R ET IU SCUL A 1886 ULTRA 1889 VICARIA 1892 XERANTI CA BAILEY1L.H. IN MACOUNTJOHN 1890 *PRATENSI S BICKNELL,E.P. 1908 ABDITA 1908 INCOMPERTA 1896 *XANTHOCARP A BLAKE, SOT 1947 ACROPHILA 1947 BREVI S 1947 ER EMOSTACHY A 1947 LAMPR OCHL A MY S 1947 MELANOPHORA 1947 PERIL E IA 1877 BURCHELL IANA 1896 OURAN~I I 1875 F ENOL ER IANA 1886 FUSCOLUTEA 1877 *LONGIROSTRI S 1896 MANDONI ANA 1856 TR IANGULAR IS BOOTTqF. 1867 *ACUTA 1846 BANKS1 I 1867 BREWER1 1862 CONJUNCTA 1858 *CR INITA 1846 GEYERI 1846 GRIFF ITHII 1867 HE BET AT A 1867 LAC1NI ATA 1867 OL IGANTHA 1859 PARCIFLORA 1846 SANGUINE A 1862 *SCOPARI A 1860 *TENTACULATA BOOTTqF. IN GRAYTA. 1863 *BONPLANDI I 1859 CONFERTIFLORA 1859 MICANS 1859 NANA 1859 ’ PAPULOSA 1859 PICTA ROOTT,F. IN H0OKERqW.J. 1839 APERTA 1839 BACK II 1839 FRANKLINI I 1839 HEPBURNI I BOOTT I W. 1884 ASS INI BOIN ENS IS 1884 LEMMONI 1884 PRAEGRACILIS NUMBER 12 107

AUTHOR DATE SPECIES

BOOTTTW. IN WATSONTS. 1880 HETERONEURA 1880 KELLOGGI I 1880 NUDATA 1880 PHYLLOMANI CA 1883 SCHAF FNE RI 1880 *SCOPAR I A 1880 SUBFUSCA 1901 BONANZENSI S 1895 *ST RICTA 1905 UNDERWOOD11 19C 1 W ILLI AMSI I 190 1 YUKONENSIS BROWN, F Bo He 1930 TAHITENSIS BROWN,R. IN RICHARDSONTJ. IN FRANKLINVJ. 1819 PODOC A R PA 1819 RICHARDSON I I 1843 CAROL I N IANA 1843 MISERA 1843 STYLOFLEXA CAREY T J 1847 PLATYPHYLLA 1847 SYCHNOC E PHAL A CHAMISS0,L.A. EX STEUDELTE~G~ 1855 PACHY S T ACHY A CHAPMAN T A. W 1860 *DIGITALIS 1860 *ST ELLUL ATA CHAPMAN,AoWo EX BOOTTfFo 1862 *ST IPATA CHAPMAN9A.W. EX DEWEYTCO 1847 BALTZELLII CHEE SEMAN 9 T. F 1892 *COMANS 1883 DEVIA 1884 PETRI E I 1908 AEQUA 1908 FELIPENSIS 1903 ICHANGENSI S 190 3 LANC IFOL I A 1903 *LONGICRURI S 1904 PRAINII 1908 PSEUDOJAPONICA 1907 SU BTR ANSVE R S A 1908 VIOL AC EA CLARKE,C.Bo IN MERRILLTEoD. 1906 RHYNCHACHAENIUM CLAUSEN9R.T. AND WAHL9H.A. 1939 *ANGUSTIOR CLOKEYTI.W* 1922 APODA 1919 ARAPAHOENS IS 1939 *INTERIOR 1922 PAYSONI S 1919 SUBIMPRESSA 1919 *TRIBULOIDES CRONQUI STTA. 1943 OBOVO IDEA CURTIS,M.A. EX GRAYqA. 1841 AE STIVALIS DEWEY 9 C 1849 *ALOPECOIDEA 1835 ARCTI CA 108 SMITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPECIES

DEWEY Co 1854 *ARI STATA 1836 BACKANA 1845 BUCKL EY I 1842 *CE PHALOPHORA 1826 COL L ECT A 1836 COLUMBIANA 1861 *DOUGLAS1 I 1836 FESTIVA 1846 FLACCOSPERMA 1824 FORMOSA 1846 HALE1 1826 HALSEYANA 1854 HAYDENI I 1826 HITCHCOCK IANA 1836 HOOKERANA 1849 IGNOTA 1847 ILLINOENSIS 1857 LAEVI-CON1 CA 1846 LEAVENWORTHII 1842 MEAD1 I 1857 MEEK1 I 1836 MIRAB ILI S 1854 NEBRASKENSIS 1842 NUTTALL I1 1835 PARRYANA 1836 PETASATA 1836 PETRICOSA 1861 RAYNOLDSII 1842 SARTWELLII 1825 SCHWEINITZI I 1826 SICCATA 1824 TENERA 1825 TR ISPERMA 1826 *UMBELLATA 1866 *V AG IN AT A 1861 VALLICOLA 1846 WOOD1 I DEWEYvCo IN TORREYvJo IN EMORY,WoHo 1859 BARBARAE 1859 THURBERI 1859 WRIGHT11 DEWEYvCo IN WOODqAo 1845 COOLEYI 1861 *HI RSUTA 1845 PRAIREA 1845 RETROCURVA DREJER, S .To No 1841 PRATENSI S ESSENBACH,NoVON IN LEHMANNpJo 1846 PREISSI I FERNALD9M.L 1942 *ABSCONDITA 1942 X ABSCONDITIFORMIS 1902 AENEA NUMBER 12 109

AUTHOR DATE SPECIES

FERNALO 9 M L 190 2 *ALATA 1942 *AMPHIBOLA 1942 BAYARDI 1902 *CANESCENS 1907 CILIARI S 1902 CRAWFORDII 1902 *CRAWFORD11 1946 *CR INIT A 1897 *CRI NITA 1937 *CRUS-CORVI 1942 *CUMULATA 1942 *DEBILI S 1913 *DEWEYANA 1941 *DIGITALIS 1938 *DIGITALIS 1902 *ECHINATA 1906 *FLAVA 1935 GARBERI 1935 *GARBERI 190 6 *GLAREOSA 1906 HARPER1 1906 HORMATHODE S 1942 *INFLATA 1906 *I NTER I OR 1942 * I NTUMESCEN S 190 1 KATAHDINENSIS 1933 LANGEANA €942 *LASIOCARPA 1906 *LAXI FLORA 1926 *L IVIDA 1902 *MIRABILIS 1902 *MIRABILIS 1915 MISANDROIOES 1902 ORONENS IS 1942 *PALLESCENS 191 8 *PAUPERCULA 1906 *PAUPERCULA 1907 PER LONGA 1933 X PSEUDO-FULVA 1942 *RICHARDSON11 1941 ~UGATA 1902 *SCOPARI A 1921 *SCOPARI A 1902 *STRAMINEA 1902 *TENERA 1942 TERRAE-NOVAE 1933 X TRICHINA 19C 1 *VESICARIA 1933 *V E S I CAR IA 110 SMITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPECIES

FERNALD 9 M L 1900 *VE STITA 1933 X XANTHINA FERNALDT M. L AND W EATHERBYT C. A. 1931 CL IVI COLA FERNALDpt4.L. AND WIEGANDTK.M. 1911 *HORNSCHUCH IANA 1924 *HOST1 ANA 1912 *SCOPARIA 1910 *SCOPAR IA FRANCHET 9 A 1895 FARGESI I 1896 PTERO L E PTA FRANCHETTA. AND SAVATIERTL. 1879 MACROGLOSS A 1879 *NUTANS 1879 PLANATA 1879 PODOGYNA GANDOGER 9 M 1920 CRANDALLII GAUOIN,J.F.G.P. 1830 *FLAVA GRAY 9 A. 1867 *DEB1 LIS GRAYTA. EX TORREYTJ. 1836 *OL IGOCARPA GROSSTR. 1941 *PHALAROIDES 1941 *PIRCHINCHENSIS 1941 *PURPUREOVAGINATA 1941 SALTAENSIS GROSSpR. IN WERDERMANNyE. 1929 WERDERMANNII HEMSLEY9W.B. 1885 POTOSINA HERMANN9F.J. 1955 AMPL ISQUAMA 1938 *ART ITECTA 1957 ATHABASCENSIS 1950 ATRACTODES 1963 *BI PART1TA 1967 CHIAPENSIS 1938 X DEAMII 1960 *EGGLESTONI I 1957 EURYSTACHYA 1965 *FISSA 1971 GUATEMALENSI S 1957 INCONDITA 1941 *INTERIOR 1938 *LA XIFLORA 1956 LIMNOPHILA 1968 *MICROPTERA 1937 PELOCARPA 1950 PERCO STATA 1964 P L E C T OC A R P A 1957 *PRATI COLA 1950 QU ICHENSIS 1948 ROANENSIS 1955 VE XANS 1936 *VULPINOIDEA 1954 *HI LLDENOWI I HERMANNT F. J. IN MCVAUGHTR. 1949 MURICULATA NUMBER 12 111

AUTHOR DATE SPECIES

HOCHSTETTER,Co Fo EX STEUDEL, EoG. 1855 MERCARENSI S HOLM 9 H .T 19C3 ACCEDENS 1905 BRAC HYPODA 1905 CAMPLYOCARPA 1903 CHALCIOLEPIS 1900 ELYNU IDES 190 5 EURYC AR PA 190 3 *FESTIVA 1902 GYMNOCLADA 1904 LACUNARUM 190 5 *LUZULAEFOLI A 1904 MICROC HAET A 190 5 OXYCARPA 190 5 PACHY STOMA 1904 PHAEOLEPIS 1904 P HY SOC HL AE N A 1902 PRIONPHYLLA 1904 *SC IRPOIDEA 1904 *SCIRPOIDEA 1904 VAGANS 1904 VITREA HOOKERtJoDo 1887 SCAPOSA HOWE9E.C. IN GOROINIER,HoCo AND HOWE,EoCo 1894 ROSAEOIDES HOWELL 9 J To 1958 JEPSONI I 1949 SPECUICOLA 1961 TOMPKINSI HULTENy0.E.G. 1942 JACOB1-PETER I JONES9M.E. 1910 ABORIGINUM 1910 ELROOI 19 10 ST ANTONENS IS KELS0,Lo 1945 ELBERTANA 1950 ER XL E BENI ANA 1945 HAGIANA 1947 UNCOMPAHGRE KNIGHT,OoW. 1906 *TR ISP ERMA KOYAMAIT. 1969 OBLANC EOLATA 1959 TAMAK II KRAUSS, R 1950 KAUA IENS IS 1950 *PLUVI CA 1950 *WAHUENSIS KRECZETOWICZTVOI 1934 PHILOC RENA KUKENTHALtGo 1929 *AP ERT A 1929 *APERTA 1910 ARSENII 1910 *BRUNNEA 1926 CUBENSI S 1926 *CU BENS IS 1926 EKMANI I 1910 ELMER1 112 SMITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPECIES KUKENTHAL ,Go 1929 * EURY C AR P A 1929 *INTERRUPTA 1910 *JAMESONI 1910 MERRILL II 1902 MICRANTHA 1899 *M ICROGLOCH IN 1911 PYC NOTHY SOS 1910 RAMOSI I 1938 S ARAW AKETENS IS 1935 SAVAIIENSIS 1920 SUKSDORFII 1920 *SUKSDORFII KUKE NT H A L 9 G AN D E KM AN 9 E L 1929 *EKMANI I KUKENTHALvGo IN ELMER9 AoDoEo 1911 PALAWANENSI S KUKENTHALTG. IN ENGLER9HoGoAo 1909 *AQUAT IL IS 1909 *CLADOSTACHYA 1909 *F IL IFOLI A 1909 *HINDS1 I 1909 *LEMANNI ANA 1909 *LENTICULARIS 1909 *NUDATA 1909 *PI NETORUM 190 9 *ST IPATA 1909 *TENUIFLORA KUNTH9C.S. 1837 STEUDELI I LEPAGEvEo 1956 X DUMANII 1956 X EXSALINA 1964 X NEOBIGELOWI I 1956 X NEOF ILI PENDULA 1956 X NEOPALEACEA 1957 X NUBENS 1962 X PATUENSIS 1964 X QUEBECENSI S 1956 *X SAXENII 1962 *VI RIDULA L ITVINOV9 Do I 1899 *ST ENOPHYLL A MACKENZIE9K.K. 1909 ABRAMSI I 1916 ABRUPTA 1935 ACUTI NELLA 1910 AESTIVALIFORMIS 1906 AGGLOMERATA 1910 AGGREGATA 1907 AGROSTOIDES 1931 ARCTAEFORMIS 1912 ATROSQUAMA 1931 AUTUMNAL IS 1935 AZTEC ICA 1910 B ILTMORE ANA 1913 BRAINERDII NUMBER 12 113

AUTHOR DATE SPECIES

MACKENZI Ec K.K. 1913 BREVICAULIS 1907 BREVI SQUAMA 1915 BULBOSTYLI S 1910 BUSHI I 1935 CAESAR1 ENS IS 1908 CHIHUAHUAENSI S 1906 CONCINNOIDES 1935 CONSPECTA 1916 CONVOL UTA 1914 CRY PTOL EP IS 1916 DAVYI 1910 DEBIL IFORM IS 1922 DUDLEY I 1915 EGGLESTONI I 1915 EGREG IA 1915 FEST IVELLA 1931 FISSA 1909 FISSURICOLA 1922 FLACC IFOL IA 1922 FRACTA 1910 FULVE SCENS 1909 FUSCOT IN CT A 1913 GEOPH ILA 1916 GR AC IL IOR 1916 HARFORD II 1922 HELLERI 1909 HOLMIANA 1916 INTEGRA 1931 I NVOLUC RAT ELLA 1916 LANCIFRUCTUS 1935 L E IOP HYL L A 1916 LEPORINELLA 1915 LUNELL IANA 190 9 MACROSPERM A 1906 MEDITERRANIA 1923 MERRITT-FERNALDII 1910 ME SOC HORE A 19G9 MICROPTERA 1935 MISERABILIS 1931 MOHR IANA 1931 MOLESTA 1922 MONTEREYENSIS 1916 MULT ICOSTATA 1907 NEOME XICANA 1909 NUBICOLA 1914 OKLAHOMENSIS 1916 OLYMPICA 1915 ONUSTA 1916 PACHYCARPA 114 SMITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPECIES

MACK EN21E 9 K K 1922 PAUC ICOSTATA 19@7 PERGLOBOSA 19C9 PER STRICTA 191 3 PITYOPHILA 1931 PLATYLEPIS 1931 PRAECEPTORIUM 1908 PROJ ECTA 1931 PROPOS ITA 1935 PURPURI FERA 1915 RUGOSPERMA 1915 RUSBY I 1931 RUTH1 I 1909 SALINAEFORMIS 1906 SAX1MONTANA 1908 SCABRIUSCULA 1908 SC IRPIFORM IS 1915 SHELDON1 I 190 7 SIMULATA 1909 SMALLIANA 1935 STELLATA 1922 STENOPTERA 1916 SUB-BRACTE ATA 1916 TENERAEFORMIS 1931 TOWNSENDII 1922 TRACY I 1907 TUMULICOLA 1922 UN ILATERAL IS 1931 WIEGANDI I MACKENZIEtKoKo EX BRIGHT,Jo 1930 LARICINA MACKENZIE,KoKo IN ABRAMSqLo 1923 NEUROPHORA 1923 SUBORBICULATA 1923 VIRIDIOR MACKEN2IEtK.K. IN PIPER9C.V. AND BEATTIE, 191 5 CUSICK II 1915 PIPER1 MACKENZIE,KoKo IN RYOBERG, PaAo 1917 ALBO- N IGR A 1917 ANGUST IOR 1917 EPAPILLOSA 1917 INCURVIFORM1 S 1917 LEPTOPODA 1917 NELSON1 I MACKENZIE,K.Ko IN SMALL,JoKo 1913 MAGN IFOL I A MAGUIRE,B* 1944 INTERIMUS 1944 RACHILL IS 1944 *VERNACULA MAGUIRE 9 B AND HOLMGRE N 9 A He 1946 WAMPYLOCARPA MAR1E-VI CTORI N q t FR ERE 1 1929 *OEDER I MERRILL 9Eo Do 1918 BAMBUS ETOR UM 1934 *RUBRO-BRUNNEA MERRILL1EoDo AND CHUNtNaKo 1935 TSOI NUMBER 12 115

AUTHOR DATE SPECIES

MEYE R TC A 1831 CIRCINNATA 1831 LEIOCARPA 1831 N IGR IC ANS MOHR T C 1910 *ST I PATA MUHLENBERG 9 H. EX W ILLDENOW T C L. 1805 RETROFLEXA NELMES, E. 1938 EXPLORATORUM 1939 SCHNEIDERI 1955 TR ICHOPHYLLA 0' NE ILLTH. To 1940 BARTLETTII 0'NEILLpH.T. AND DUMANTM. 194 1 DUTILLY I OHWI T J. 1934 APODOSTACHYA 1931 CUNE AT A 1934 HATUSIMANA 1932 HY MENODON 1933 KURILENSIS 1932 RUG AT A 1952 TETSUOI OLNEY 9 SoT 1871 PORTER I 0LNEYvS.T. EX 6AILEYgL.H. 1884 *ADUSTA 1888 *DEWEYANA 1886 *LEPORINA 1886 OREGONENSI S 0LNEYvS.T. EX BAILEYTL.H. IN COULTERTJ~M. 1894 *WILLDENOVI I 0LNEYTS.T. EX BOOTTvW. IN WATSONpS. 1880 MENDOCINENSIS 0LNEYvS.T. IN GRAYTA. 1868 ATHROST ACHY A 1868 CINNAMOMEA 1868 GYNODYNAMA 1868 LUZUL INA 1868 SARTWELLIANA 1868 WH ITNEY I 0LNEYgS.T. IN PORTER9T.C. IN HAYDEN9F.V. 1872 HALL1I 0LNEYyS.T. IN WATSONvS. 1871 *AUREA 1871 HAYDENIANA 1871 WATSON I PAR1 SH 9 S B 190 5 AUSTROMONT ANA 190 5 JACINTOENSIS PECK 9C.H. 1895 *ROSEA 1894 *ROSEA 1895 *ST IPATA 1893 *TORTA PETRI E, D 1881 KALOIDES 1882 LONGICULMI S PIPERqC. V. 1906 *HOOD I I 1906 PHAEOCEPHALA PORS ILDTA. E. 1943 *ATROF USCA 1939 KOKRINENSI S 1939 MELOZITNENS I S 1943 MORRISSEYI PRESL TKOB. 1828 ANTHOXANTHERA 116 5AlITHSONIAN CONTRIBUTIONS TO BOTANY

AUTHOR DATE SPEC IE S

RAUP H oM 1947 SOPERI RAYMOND, Mo 1959 SURCULO SA 1959 2 IZ AN IAEFOL IA ROACH,AoWo 1952 DIVERSI STYL IS RYDBERG, Po A. 1901 EBENEA SARTWELL9H.P. EX DEWEY,Co 1855 CHAPMAN I 1849 SCABR IOR SCHWEINITZ,LoDo 1824 COSTATA 1824 CR ISTAT A 1824 GRAC ILL IMA 1824 NIGRO-MARGINATA 1824 NOVAE-ANGLIAE SCHWEINITZyLoDo AND TORREY,Jo 1824 BARRATTI I SHUTTLEWORTH,RoJo EX KUNZEqGo 1844 CRUS-CORVI SToJOHNyH. 1947 VITI ENS IS STAC EY 9 J W 1938 CONSTANCEANA 1937 CURATOR IUM 1939 DANAENS IS 1938 EA STWOOD IANA 1936 OBISPOENSIS 1937 SONOMENSI S 1939 SUBNIGR ICANS STANDLEY,P.Co 1947 STEYERMARKI I STANDLEY,PoCo AND STEYERMARK,JoAo 1953 CUCHUMATANENSIS 1947 HUEHUETECA 1953 TOJQU IANENS IS 1953 TUNIMANENSI S 1953 VENOSIVAGINATA STEUDEL,EoGo 1855 AMPHI BOLA 1855 AUROLENSIS 1855 FL ACC ID ULA 1855 INC ISO-DENT ATA 1855 MAC R OKOL E A 1855 PICTA 1855 PRESL II 1855 PTYCHOCARPA STEYERMARK,JoA. 1964 A2 UAYAE 1951 CULMEN ICOLA 1951 LARENSI S 1951 RORAIMENSI S 1952 ST ANDLEY ANA 1951 TACH1 RENSI S 1951 TAMANA 1954 TOREADORA 1951 TURUMIQUIRENSIS SUKSDORF ,WoNo 1923 *NEBRASKENSIS 1906 PADDOENSIS TORREY J 1836 ALATA 1836 JAMES II NUMBER 12 117

AUTHOR DATE SPECIES

TORREY 9 J 1836 SCOULERI 1836 TURGE SCENS TORREY, J. EX DEWEY ,C. 1836 CAREYANA 1846 HE TEROSTAC HYA 1836 HOUGHTONI ANA TORREYTJ~IN WILKESyC. 1854 WILKESI I TUCKERMAN T E. 1843 ALOPECOIDEA 1843 *CANESCENS 1843 *SCOPARIA TUCKERMAN,Eo EX DEWEY,C. 1860 ARGYRANTHA TUCKERMANvE. EX 0LNEYvS.T. IN GRAYTA. 1868 GL AUCODEA UNDERWOOD, JoK. 1945 *OXYLEPI S WAHLENBERG ,Go 1803 AQUATI L IS WATERFALLTU~T. 1954 LATEBR ACTE AT A WEATHERBY,C.Ao 1936 MAC KE NZ I AN A W1EGANDyK.M. 1922 CR EBR I FLORA 1922 ORMOST ACHY A WRIGHTTS.H. IN DEWEYpC. 1866 XEROCARPA 118 SMITHSONIAN CONTRIBUTIONS TO BOTANY

PUBLICATION-DATE INDEX

DATE TAXUN AUTHOR

1803 AQUAT ILIS WRHLEmERG TG. 1805 R ETPOFLEXA MUHLENBERG TH EX W ILLDENOW T C L 1819 PODOCARPA BROWNTR. IN RICHARDSONqJ. IN FRANKLINfJ. R I CHARDSONT I BROWNTQ~IN RICHARDSONpJ. IN FRANKLINTJ. 1824 BARRATTI I SCHWE INI TZ TL D. AND TORREY T J COSTATA SCHWEINITZ 9L.D. C R I S TATA SCHWE INIT2 TL 0. FORMOSA OE WEY TC GRAC ILLIMA SC HWE INI TZ TL 0. NIGPO-MARGINATA SCHWE INIT2 TL 000 NOVAE-ANGLIAE SCHWEINIT2,L.D. TENERA DEWEY T C 1825 SCHWEINITZI I DEWEY +C. T R IS PEP MA OEWEY TC. 1826 Cot1ECTA DEWEY T C HAL SEYANA DEWEY TC. HITCHCOCK IANA DEWEYqCo S ICCATA DEWEY TCo *UMBELLATA DEWEY 9 c 0 1828 ANTHOX ANTHER A PR ESL TK*R 1830 *FLAVA GAUDINTJ.F.G.P. 1831 CIRCINNATA MEYER TC. A. LE IOC AR Pb MEYERTC.A. NIGRICANS MEYER9C.A. 1835 ARCT ICA DEWEY TCo PARRYANA DEWEY 9 C 1836 AL AT A TORREY T J. BACKANA DEWEY 9 C CAREYANA TORREYtJ. EX DEWEYTC. COLUMB I AN4 DE WEV TC. FEST IVA DEWEY T C HOOKE RANA DEWEY 9 C HOUGYTONI ANA TORREYTJ. EX DEWEYfC. JAMES11 TORREY 9 J MIRABILIS DEWEY TC. *OL I GOCARP A GRAYTA. EX TORQEYqJo P ETA SAT A DEWEY T C. PETR ICOSA DEUEY TC. \ SCOULER I TORREY T J. TURGESCENS TORREY + J 0 1837 STEUDEL II KUNTHTCOS. 1839 AP ERTA BOOTTvF. IN HOOKERTW.J. BACK1 I BOOTTIF. IN HOOKERTW.J. FRANKLIN1 I 6013TT~F. IN HOOKERTW.J. HEPBURNI I ROOTTTF. IN HOOKERTW.J. 1841 AESTIVALI S CURTISTM.A. EX GRAYTA. PR ATENS IS ORE JER T S. To No 1842 *C EPHALOPHORA DEWEY TC. MEAD11 DEWEY TC. NUMBER 12 119

DATE TAXON AUTHOR

1842 NUTTACL II DEWEY ,Co SARTWECL II DEWEY T f 1843 ALOPECOIDEA TUCKERMANTE. *CANESCENS TUCKERMANTE. CAROLINIANA BUCKLEY9 SoRo MISERA BUCKL EY p S Bo * S COP AR IA TUCKERMAN, Eo STY1OFL E XA BUCKLEYTSOB. 1844 CR US -COR V I SHUTTLEWORTHTR.J. EX KUNZEtG. 1845 BUCKLEY I DEWEY 9Co COOLEY I DEWEYTC. IN WOODPA. PRAIREA DEWEYTC. IN WOODTA. R E T R OCU RV A DEWEYTC. IN WOODTA. 1846 BANK SII BOOTT 9 F 0 FLACCOSPERMA DEWEY T C GEYERI BOOTT T F GRIFFITHII B(30Tf 9 F HALE I DEWEY T C HETEROSTACHYA TORREYTJ. EX OEWEYTC. LE4VENWORTHI I DEWEYTC. PREISSI I ESSENRACHTNmVON IN LEMMA", J SANGUI NEA BOOTT F WOOD1 I DEWEY rC. 1847 BACTZELL II CHAPMAN, 4oWo EX DEWEY TC ILL INOENS IS DEWEY cCo P L ATY P HYL L A CAREY, Jo S YCH NOC E P H AL A CAREY 9 J 1849 *ALOPECOIDEA DEWEY 9Co IGNOTA DEWEY ,to SCABRIOR SARTWELLTHOP. EX DEWEYvC. 18 54 *ARI STATA DEWEY tCo HAYDENI I DEWEY ,Co NEBRASKENSIS DEWEY ,Co WILKESI I TORREYTJ. IN WICKEStCo 1855 AMPH IBOLA STEUDEL,EoGo AUROLENSI S STEUDEL T E. Go CHAPMAN I SARTWELLTHOP. EX DEWEYTC. FL ACC IDUL 4 STEUDEL E *Go INCISO-DENTATA STEUDEL T €0 Go MACROKOCEA STEUDEL T €.Go MERCARENS IS HOCHSTETTERTC~F. EX STEUOELTE.G~ PACHYSTAC HYA CHAYISS0,L.A. EX STEUDELTE~G. PIETA STEUDEL T E Go PRESLI I ST EUDEL 9 E oG P TYC HOC AR PA STEUOELTE.G. 1856 TRIANGULARIS BO ECK E LER p J 0 1857 LAEVI-CONICA DEWEY ,Co MEEKII DEWEY T C. 1858 *CRIN ITA BOOTT T F 120 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DATE TAXON AUTHOR

1859 BAR 8 AR A E DEWEY,C. IN TOQREYyJ. IN EMORY,W.H. CONFERT IFLORA BOOTTvF. IN GRAYyA. MICANS BOOTTVF. IN GRAYqA. NANA ROOTTtF. IN GRAYtA. P APUL 0s A BOOTTIF. IN GRAY,A. PARC 1 FLORA ROOTT 9 F P ICTA BOOTT,F. IN GRAYyA. THURBER I DEWEYvC. IN TORREY,J. IN EMORYTW.H. WR IGHTI I DEWEYtC. IN TORREY,J. IN EMORY9W.H. 1860 ARGYR ANTHA TUCKERMAYgEo EX DEWEYqC. *DIGITALIS CHAPMAN, A.W. * S TEL LUL AT A CHAPMAN 9 A. W *T E NT AC UL AT A BOOTT T F 1861 *DOUGLAS II DEWEY 9C. *HI RSUTA DEWEYpC. IN WOODtA. RAYNOLDS I I DEWEY T C VALL ICOLA DEWEY cC. 1862 CONJUNCfA BOOTT T F *SCOPARI A BOOTT 9 F *STIPATA CHAPMAN,A.W. EX BOOTTpF. 1863 *BONPLANDI I BOOTTTF. IN GRAYVA. 1866 *VAGINATA DEWEY TC. XEROCARPA WRIGHT9S.H. IN DEWEYqC. 1867 *ACUTA BOOTT 9 F BREWER1 BOOTT F *DEBILIS GRAY, A, HE BET AT A ROOTT 9 F LACINIATA BOOTT,F. OL IGANTHA BOOTT 9 F. 1868 ATHROSTACHYA 0LNEYtS.T. IN GRAYtA. CINNAMOMEA OLNEY,S.T. IN GRAYvA. GLAUCODEA TUCKERMANtE. EX 0LNEYtS.T. IN GRAYqA. GY NO DY N AM A OLNEY,S.T. IN GRAYqA. CUZULINA OLNEY,S.f. IN GRAYvA. SAQTWELLIANA OLNEY9S.f. IN GRAYqA. WH ITNEY I 0LNEYtS.T. IN GRAYqA. 1871 *AUREA OLNEY,S.T. IN WATSONqS. HAY DEN IAN A OLNEYI SOTO IN WATSON9 So PORTER1 OLNEY 9 5 .T WATSON1 OLNEYTS.T. IN WATSON,S. 1872 HALL II 0LNEYtS.T. IN PORTER,TbC. IN HAYDEN9F.V. 1875 FENDLER IANA BOECKELERtJ.0. 1877 BURCHELL IANA BOECKELERt J.0. *LONG IROSTRI S BOECKELERtJ.0. 1879 MACR OGL 0s S A FRANCHETtA. AND SAVATfERvL. *NUTANS FRANCHETt A. AND SAVAfIERpC. PLANATA FRANCHET, A. AND SAVATIER,L. PO DOGYN A FR ANCHET 9 A AND SAVATIER?L. 1880 HE T E P ONEU R A BOOTTtW. IN WATSON, So NUMBER 12 121

DATE TAXON AUTHOR

1880 KELLOGGI I BOOTTtW. IN WATSONtS. MENDOCINENSIS 0LNEYtS.T. EX BOOTTtW. IN WATSON, So NUDATA BOOTTtW. IN WATSONtS. P HYLL OM AY IC A BOOTTtW. IN WATSONtS. *SCOPARI A BOOTT t W I IV WATSON T So SUBFUSCA BOOTTtW. IN WATSONTS. 1881 KALOIDES PETRI EtD. 1882 LONGICULMIS PETRI EtD. 1883 DEVIA CHEESEMANt1.F. SCHAFFNER I BOOTTtW. IN WATSONqS. 1884 * ADU S T A 0LNEYtS.T. EX BA1LEYtL.H. ASSINIBOINENSIS BOOTT t W *CANE SCENS BA ILEY 9 L H. HALL IANA BAILEYtL OH. LEMMONI BOOTT t U. PETRIEI C HEE S EMAN 9 To F PRAEGRAC IL IS BOOTTtW. 1885 NERVINA BAILEY 9L.H. POT0 S INA HE MSL EY t W B 1886 *BRONGNIARTI I BAILEY TL.He ENGELMbNNI BAILEY ,L.H. FUSCOLUTEA BOECKELERt J.0. *GAYANA BAILEYtLoH. INOP S BAILEYtL.H. *LE POR INA 0LNEYtS.T. EX BIILEY,L*H. *MARC IDA BA I LEY tL OH OREGONENSIS Ol-NEYtS.T. EX BA1LEYtL.H. *R ETROCURV A BAILEY tL .He *STYLOSA BAILEY tL.H* ULTRA BAILEY 9L.H. 1887 SCAPOSA HOOKE R t J 0 1888 WEWEYANA 0LNEYtS.T. EX BA1LEYtL.H. PANSA BAILEY L H. *ROSE A RAILEY,L.H. 1989 ACUTINA BA ILEY ,L .He ALBIDA BAILEYtL *H ALMA BAILEY L .H CALIFORNICA 6AILEYtL.H. COb4MUN I S BA ILEY t L eH DONNELL-SMITH11 BA1CEYtL.H. ELEOCHARIS BAILEY t L. H *FEST IVA RA1LEYvL.H. *FOENEA BAILEY ?LoH *GR IS EA BA1LEYtL.H. *HOOD II 8AILEYqL.H. ILLOTA BA1LEYtL.H. JONES11 BA I LEY 9 L H *MILIARIS BA ILEY t L .H SPECIFICA BA1LEYtL.H. 122 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DATE TAXON AUTHOR

1889 SPSETA BA ILEY 9 L oH *STRAMINEA BAILEYTL~H~ *STRAMlYEA BA I LEY I Lo He *TERETlUSCULA BAILEYTL~H~ VICAR1 A BAILEY ,L Ho 1890 *PRATENSIS BAILEY,LoHo IN MACOUNTJOHN 1891 QUADRIFIOA BAILEYTLoHo *QUAOR1 f IDA BAILEYTLOH 1892 *COMANS CHEESCYANy ToFo YONTANENSIS BAILEY TLOH PRINGLEI RA ILEY TL H X E 9 ANT IC A BA ILEY 9 L oH 1893 A USTRU-CAROL1 NIANA BAILEY TLoHo OURIFOLIA BA ILEY T L Ho FETA BAILEY rLoHo 1NTE RIOR BAILEY TL Ho *STER 1L IS BA ILEYT L H *TORTA PECKTCOH. 1894 ROSAEOI DE S HOWETEOC. IN GORDINIER,HoCo AND YOWEtEoCo *ROSEA PECK, CoHo *WILLDENOVII OLNEY,SoTo EX BAILEYyLoHo IN COULTERTJOM. 1895 FARGESI I FRANCHET, A. *ROSEA PECK, C OH *STIPATA PECK, C eHo *STR ICTA BR ITTON T No Lo 1896 DURANOI I BOECKELERrJoOo EGGERTI I BAILEY ,L .Ho HASSEI BA ILEY (Lo Ho I DAHOA BA ILEY ,L Ho MANDONIANA BOECKELERt JoOo *NERRASK ENS IS BAILEYTLoHo PTE ROLE PT A FRANCHET, A. * X ANT HOC AR P A BICKNELLTEOP. 1897 *CR INITA FERNALDTM ot 1898 MADRENSI S BA ILEY TLoHo 0 AX ACAN A BAILEY TL H SEATON1 ANA RA ILEY TL Ho 1899 *MI CR OGLOC H IN KUKENTHALvG. *STENOPHYLLA L I TV I YOV T D I 1900 ELYNOIDE S HOLM, H oT *VESTITA FE RN4L D T Mo L 1901 BONANZENSIS BR I TTONt No Lo EBENEA RYDBERG? Po A. KATAHDI NENSI S FE SNALD T Mo L *VESTCAR14 FERNALD,MoLo WILL I AMSI I BRITTUN7 No Lo YUKONENSI S RRITTON,N.Lo 1902 AENEA FE RNAL DTM oL *ALATA FERNALD, Mo Lo YUMBER 12 123

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1902 *CANESCENS FERNALDTY-L. CPAWFORDI I FE QN AL DI M L *CRAWFORD11 FERNALD, M. L. *€CHI NATA FERNALDt Me L. GY MNOC L AD A HOLM1 HOT. MICRANTHA KUKENTHAL ,Go *MIRABILIS FE QNALD T H L *MIRABILIS FERNAtD, M*L ORON ENS IS FERNALDt Me L. PR IDNPHY L L A HOLM, H T* *SCOPARIA FERNALDt M.L. *STRAMINEA FERNALD9M.L. *T EN ERA FERNAL0tM.L. 1903 ACCEDENS HOLM9H.T. CHALCIOLEPIS HOLM, HOT. *FESTIVA HOLM, HOT. ICHANGENS IS CLARKE TC R LANCIFOLIA CLARKE tC 6 *LOWGICRUR IS CLARKE TC B 1904 LAC UNA P UM HOLM, H .T MICROCHAETA H0LMtH.T. PHAEOLEPIS HOLM, H.T. PHYSOCHLAENA HOLM 9 HOT. PRAINII CLARKE C*B *SCIRPOIDEA HOLYIH*T* *SC IRPOlOEA HOLM9 H .T* VAGANS HOLM, HOT. VITREA HOLM,H.T. 1005 AUSTROMONTANA PAR1 SH 9s. 6. BRACHYPOOA HOLYTl4.f C A YPLY OC A RP A HOLM t H .T EURY CAR PA HOLM 9 H T J AC I NTOENS IS PAR ISH 9 S B *LUZULAEFOLf A HOLM, H .To OXYC ARP A HOLM, H T PACHYSTOMA HOLM,H.T. UNDERWOOD11 BRITTON, N. L. 1906 AGGLOMERATA MACKENZIE,K.K. CONCINNOIDES MACKENZ IE TK Y *FLAVA FERNAL0tM.L. *GLAR EOSA FESNAL0,M.L. HARPER I FERNALD9M.L. *HOOD1 I PIPER TC. V. HO R M AT HOD E S FERNALDTMOL. *INTE RIOR FERNALDt M L. *LAXI FLORA FERNALDtM.L. MEDITERRANIA MACKENf1EtK.K. PADDOENSI S SUKSD0RFtW.N. *PAUP ERCUL A FERNALDTY-L. 124 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DATE TAXON AUTHOR

1906 P HA E OC E P H AtA PIPERT C oV RHYNCHAC HAE NIUM CLARKETC~B. IN MERRILLTEOD. SAXIMONTANA MACKENZIE KoKo *TR I S PERMA KNIGHTTO~W. 1907 AGROSTOIDES MACKENZI E T Ko Ko BREVISQUAMA MACKENZIErKoKo CILI ARI S FE RN ALD T M L NEOMEXI C AWA MACKENZI E KO Ko P E RGLO BOS A MACKENZIETKOK. P E RLONG A FE RNALD M L S IYULATA MACKENZI E Ko Ko SU RTR AN SV ERS A CL ARK E 9 C B TUMUL I COL A MACKENZI E T K Ko 1908 ABDI TA BICKNELLTEOPO AEQUA CLARKE 7 C B CHIHUAHUAENSI S MACKENZI E 9 Ko Ko FELIPENSIS CLARKE 9 C 6. INCOMPERTA BICKNELL, E *Po PROJECTA MACKENZIETYo K. P S EIJ DOJ APON I C A CL AQKE T C 06 SCAB RT USC UL A MACKENZIETK.K. SC IRPIFORMI S MACKENZI E T K Ko V IOL AC EA CLARKEvC oBo 1909 ABRAMSI I MACKE NZIE K K *AQUATILIS KUKENTHAtrG. IN ENGLERTH*G~A~ *CL4DOSTACHYA KUKEN~HALTG. IN ENGLERTHOG-A. *F I L IFOt I A KUKENTHALTG. IN ENGLERTH~G~A~ FISSUPICOLA MACKENZIEt KI Ko FUSCOT INC T A MACKENZIE,K.K. *HINDS1 I KUKENTHALTG. IN ENGLERTH~G~A~ HOLYIANA MACYEN 21E T K K *L EYANN I AN A KUKENTHALqG. IN ENGLERTH~G~A~ *LENTICULARIS KUKENTHALTG. IN ENGLER,HoGoAo M ACR OSP ERMA MACKENZIE,K*Ko MICROPTER A MACKENZIE, KoKo NUBI COLA MACKENZIETK~K. *NUDATA KUKENTHALTG. IN ENGLESTH.G*A* PERSTRICTA MACKENZI E TK K *P I NETORUM KUKENTHALTG. IN ENGLER,HoGoAo SAL I NAEFORM IS MACKENZIETKOK. SMALL1 ANA MACKEYZIETK-K. *STI P AtA KUKE N THA L G IN ENGL E R T H G A *T ENU I F L OR A KUKENTHALqG. IN ENGLERTHoG-A. 1910 ABOR IGINUM JONESTMOE. AESTIVALIFORMIS MACKENZIETK~K. AGGREGATA MACKENZIE TKOK ARSENII KUKENTHAL T Go B I LTMORE AN A MACKENZI E TKOK. *BRUNNEA KUKENTHAL T Go NUMBER 12 125

DATE T AXON AUTHOR

1910 BUSHI1 MACKENZIE9K.K. DEB IL IFORM IS MACKENZIEtKoK. ELVER1 KUKENTHAL t G. ELRODI JONES 9 MeE. FULV ESC ENS MACKENZIE9K.K. *JAW€ SON1 KUKENTHALtG. MERR ILLI I KUKENTHALtG. M E S OC HO RE A MACKENZI EtK OK RAMOSI I KUKENTHAL tG. *SCOPARIA FERNAtO 9 M. L . AND WIEG AND t K. M STANTONENSIS JONES 9 MoE *STI PATA MOHR, C 1911 *HORN SCHUC H I AN A FERNALOTM.L* AND WIEGANDTK. Ma PALAW ANEN S IS KUKENTHAL 9 G I N ELMER t A D E PYCNOTHYSOS KUKENTHAL (Go 1912 ATROSQUAM A MACKENZ1EtK.K. * SCOPAR I A fEQNALD7M.L. AND WIEGAND9K.M. 1913 BRAINERDII MACKENZI E1K.K. BREVICAUL IS MACK ENZ IE 9 K K *DEWEYANA FE RN AL D. M L GEOPH! LA MACKENZI E t K.K. MAGN IFOL IA MACKENZ1EtK.K. IN SMALL9 J.K. PITYOPHILA MACKEYZ1EtY.K. 1914 CRYPfOLEPIS MACKENZIE t K. K. OKLAHOMENS IS MACK ENZ IE t K OK 1915 RULBOSTYLIS MACKENZIE 9 K.K. CUSICKII MACKEN2IEtK.K. IN P1PERtC.V. AND BEATTIE9R.K. EGGLESTONII MACKENZIE9K.K. EGREGIA MACKENZI E1K.K. FEST IVELL A MACKENZ1EtK.K. LUNELLIANA MACKENZ1EtK.K. MISANDROIDES FERNALOp Ma Lo ONUSTA MACK EN2 I E 9 K K PIPER1 MACKENZIE9K.K. IN PIPER7C.V. AND BEATT1EtR.K. RUGOSPERYA MACKENZIE 9K.K. RUSBYI MACKENZIE9K.K. SHELDON I I MACKENZIE9K.K. 1916 ASRUPTA MACKENZ1EtK.K. C ONV OL UT A MACK ENZ IE t K K DAVYI MACKENZ IE 9 KO K GRAC ?L?OR MACKENZIE TY.K. HARFORD II MACKENZIE9K.K. INTEGRA MACKENZI E t K. K. L ANC I FRUCTUS MACKENZIEt KOKO LEPORINELLA MACKENZIE tK OK MAR1 POSANA BAILEY,L.H. MULT ICOSTATA MACKENZIE9K.K. OLYMPIC A MACKENZIE9K.K. PACHYCARPA MACKENZI E t K. K 126 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DATE TAXON AUTHOR

1916 SUB-BRACTEATA MACKENZ IE9 K OK TENERAEFORMI S MA CKENZI E T K K 1917 ALBO-NIGRA MACKENZIE9K.K. IN RYDBERG,P.A. ANGUST IOR MACKENZIE,KoK. IN RYDt3ERGtP.A. EPAPILLOSA MACKENZIE ,KoKo IN RYDBERGTP A. INCURV IFORMI S MACKENZIE,K.K. IN RYDBERG,P.A. L E PT OPODA MACKENZI E,K. K. IN RYOBERG, P A. NELSON1 I MACKENZIE,K.K. IN PYDBERGTP.A. 1918 BAMB US ETORUM MERRILLqEoDo * P AUPERC IJL A FERNALD, Me L 1919 ARAPAHOENSI S CLOKEY 1. W SUBIMPRESSA CLOKEYTIOW. *TR IB ULO ID E S CLOKEY 9 I ow. 19 20 CHIK UNGAN A BA ILEY T L H CRANDALLJ I GANDOGER9M. KULI NGANA BAILEY 9 L OH SUKSDORFI I KUKENTHAL I Go *SUKSDORfI I KUKENTHALtGo 1921 *SCOPARI A FERNALDTM. Lo 1922 APOOA CLOKEY,I.W. CREBRI FLORA WIEGAND9K.M. Z)UOL EY I YACKENZIE,K.K. FLACCIFOLIA MACKENZIEIK~K. FRACTA MACKENZIETK.K. HELL EP I MACKENZIE9K.K. MONT E RE Y E N S IS MACKENZIETK.K. ORMO ST AC HYA WIEGAN0,K.M. PAUCICOSTATA MACKENZIETK.K. P AY S ON I S CLOKEY 9 I W STENOPTER A M4CKENZI E T K. K. TRACY1 MACKENZI E 9 K. K. UNILATERALIS MACKENZIE,K.K. 1923 MERRITT-FERNALDII MACKENZIE,K.K. *N EBR ASKENS IS SUKSDORF9W.N. NEUROPHOP A YACKENZIE,K.Ko IN ABRAMSvC. SUBORBICULATA MACKENZIETK.KO IN ABRAMSTL~ VIRIDIOR MACKENZIEvK*K. IN ABRAMSTL. 1924 *HOSTIANA FERNALD, MoLo AND W IEGAND, K OM. 1926 CUBENSI S KUKENTHAL ,Go *CUBENS IS KUKENTHALTG. EKMANI I KUKENTHAL *L IVIDA FERNALD9M.L. 1929 *APERTA KUKENTHAL (Go *APEP TA KUKENTHAL ,Go *EKMAN II KUYENTHALqG. AND EKMAN,E.L. *EURY CAR P A KUYENTHAL (Go *INTERRUPTA KUKENTHAL 9 G* *OEDERI MARIE-VICTORIN, (FPERE) WEROERMANNI I GROSSTRO IN WERDERMANNqE. NUMBER 12 127

DATE f AXON AUTHOR

1930 LARICINA MACKENZIE9K.K. EX BRIGHTTJ. TAHITENSIS BROWN 9 F B OH. 1931 ARCTAEFORMI S MACKENZI E y K. K AUTUMNAL I S MACKENZIE9K.K. CL IVICOLA FERNALD9 M. L ANlr WEATHERBY 9 C.A CUNEATA OHWI 9 J. FISSA MACKENZIE 9K.K. INVOLUCRATELLA MACKENLIE9K.K. MOHRI ANA MACKENZI E 9K. K. MOLE STA MACKENZIE9K.K. PLATYLEPI S MA CKE NZ IE 9 K K PRAECEPTORIUM MACKENZI E 9 K.K. PROPOSITA MACK EN2 IE 9 K K RUTH II MACKENZ IE 9 K OK TOWNSENDII MACKENZ IE 9 K. K W I EGANDI I MACKENZIE9K.K. 1932 HYMENODON OHWITJ. RUGATA OHMI T J. 1933 KURILENSIS OHWIT J. L ANGE AN A FERNAL0,M.L. X PSEUDO-FULVA FERNALD9M.L. X TRICHINA FERNALD9M.L. *VESICARIA FERNALD9M.L. X XANTHINA F E RNALDTM. L 1934 APOOOST ACHY A OHWITJ. HATUSIMAlVA OHWI 9 J. PHILOCP ENA KR EC 2 ETOWI CZ T V I *RURRO-BRUNNEA MERRI LL 9 E .Dm 1935 AB'LATA BAICEY9L.H. ACUT INELLA MACKENZIE 9 K.K AZTECICA MACKENZIE9K.K. CAESAR I EN SI S MACKENZIE,K.K. CONS PECTA MACKENZ I E 9 K K GARBERI FERNALD9M.L. *GARBERI FERNALD9M.L. L E I OPHY L L A MACK ENZ I E 9 K .I( MISERABI LIS MACKENZIE 9K.K. PURPURIFERA MACKENZIE9K.K. SAVAIIENSTS KUKENTHAL 9 Go S T ELL AT A MACKENZIE9K.K. TSOI MERRI LL 9 E D. AND CHUN 9N.K 1936 MACKENZIANA W EATHERBY ,C. A* OBISPOENSIS STACEY 9 J. We WULP INOIDEA HEQMAlVN9 F. J. 1937 *CRUS-CORVI FE RNAL D 9 M L CURATOR IUM STACEY 9J. W. P ELOCAR P 4 HERMANNTF. J. SONOMENS1 S ST ACEY T J W 1938 *ART1 TECTA YERMANNTF. J. 128 SMITHSONIAN CONTRIBUTIONS TO BOTANY

DATE TAXON AUTHOR

1938 CONSTANCE ANA STACEYTJmW. X DEAMII HERMANNqFm Jm *DIGITAL IS FERNALD, MmLm EASTWOODI ANA STACEYVJoWm E XP L OR A TO RUM NELMES ,Em *LAXI FLORA HERMANN, FmJm SARAWAKETENSI S KUKENTHALTGm 1939 *A NG U ST I OR CLAUSENTRmTm AND WAHL?HmAm DANA ENS IS STACEYvJmWm *INTERIOR CLOKEY?I.Wm KOKRINENSIS PORSI LDTAm Em MELOZITNENS IS PORSILD, Ao Em SCHNEIDER I NELMES ,E SUBN I GR ICANS STACEY 9 JmW. 1940 BARTLETTII O'NEILL? HoTm 1941 +DIGITAL I S FEQNALDpMmLm DUTILLY I O'NEILL9H.T. ANT) DUMANqMm *INTERIOR HERMANN? Fm J *PHACAROI DE S GROSS *PIRCHINCHENSIS GROSS? Rm * P U RP UR E OV AG I N AT A GROSS 9 Rm R UGATA FE RNALD 9 M L SALTAENS I S GROSS T Ro 1942 +A6 SC OND ITA FERNALO, MoLm X A B SC OND I T I FORM 'I S FERNALDTM-Lo +PMPH IROLA FERN AL 0 T M L BAYARDI FE RNAL 09 MmLm *CUMUL AT A FERNALDT MmLm *DEBILIS FERNALD?MmLm *I NFCATA FERNACDVMm Lm *INTUYEStENS FERNALDTMm Lm JACOBI-PETER1 HULTEN, 0. E mG *LASI OCARPA FE RN ALD 9 Mo L *PALL ESCENS FEQNALD, Ye Lo *RICHARDSON11 FERNALDV MmLm TERR AE-NOVAE FERNALDTM-Lo 1943 *ATRO FUS C A PORSI LD, AmEm MORRI SSEYI PORSI LDTAm Em OBDVOI OEA CRONQUIST? Am 1944 I NTEQ I MUS MA GU I R E 7 B RACH ILL I S MAGUI RE 7 R WERNACULA MAGUI RE 9 6 1945 FLBERTPNA KELSO 9 L HAG1 ANA KELSOTLm *OXYL EP IS UNDERWOOD? JmKo 1946 *CA MPYL OC A RPA MAGUIRETBm AND HOLMGREN,AmHm *CRI N I TA FERNALD?Mm Lm 1947 ACROPHILA BLAKE?SOT. 6REVI S BLAKETSmTm NUMBER 12 129

DATE T AXON AUTHOR

1947 EREMOSTACHYA BLAKE 9 So1. HUEHUETECA STANDLEY 9 P .C AN0 STEYERMAQK 9 J. A. LAMPROCHC AMY S BLAKE9S.T. MELANOPHORA BLAKE 9 SOTO PEP ILEIA BLAKE 9 S of SOPER I RAUP 9 H. M. STEYERMARKT I STANDLEY TP.C UNCOY PAHGRE KELSO 9 L VITIENSIS ST.JOHN9H. 1948 RO AN EN S IS HERMANNTF. J. 1949 M UR ICU L AT A HERMANN9 F J IN MCVAUGHTR SP ECUICOLA HOWELL 9J.T. 1950 ATP ACTOOE S HERMAhSN9 F. J. ERXLEBENI ANA KELSO 9 L. K AUA IENS IS KRAUSS TR PERC OST AT A HESMANN9 F. J. *PLUVI CA KRAUSS 9R. QUICHENSI S HERMANNvF. J. *WAHU ENS IS KRAUSS 9 R 1951 CULM EN ICOLA STEYERMARK9J.A. L ARENS I S STEYERMARK9J.A. RORAIMENSIS STEYEQMARK 9 J A. TACHIRENSIS STEYEP MARK 9 J A. TAMANA SfEYERMARK9J.A. TURUMIQUIRENSIS STEY ERMARK 9 J A 1952 OIVERSI STYLI S ROACH 9 A. W ST ANOL EYANA STEYERNASK9J.A. TETSUOI OHWI 9 J 1953 C UCHUMATANENS I S STANDLEY9P.C. AN0 STEYERMARK9J.A. TOJQUIANENSIS STANDLEYTP~C~AN0 STEYERYARK9J.A. TUN1 MANENSI S STANDLEYTP.C. AND STEYERMARK9J.A. VENOSIVAGINATA STANDLEY9P.C. AND STEYERMARK9J.A. 1954 LATEBRACT EAT A WATERFALL 9 U. To TOREADORA STEYfRMARK9J.A. *WILLDENOW I I HERMANNfF. J. 1955 AMPLISQUAMA HERMANN9 F J TRICHOPHYLLA NELMESTE. VEXANS HERMANN9F.J. 1956 X OUMANII L E P AG E 9 E X EXSALINA LE PAGE 9 E L IhNOPHIL A HERMANN9 F. J. X NEOFILIPENDULA LEPAGE 9 E X NEOPALEACEA LEPAGE 9 E *X SAXENII LEPAGEvE. 1957 ATHABASCENSI S HERMANN9F.J. EURY STACYYA HERMANNTF. J. INCONDI T A HERMA"N9 F. J. X NUBENS LE PAGE 9 E *PRATICOLA HERMANNvF. Jo 130 SMITHSONIAN CONTRIBUTIONS TO BOTANY

D4TE TAXON AUTHOR

1958 JEPSONI I HO WEL L T J T 1959 SURC UL 0 S A RAYMONDTM~ TAMAKII KOYAMA ,To ZIZANIAEFOLIA RAYMONDTM~ 1960 *EGGLESTONII HERMANN, Fo J. 1961 TPMPKINSI HOWELL T J T 1962 X PATUENSIS LEPAGE T E *VIP IDULA LEPAGE TE 1963 *BIPARTITA HERMANNTF~Jo 1964 AZUAYAE STEYERMARK, J A. X NEOBIGELOWI I LEPAGE TEo PLECTOC AP PA HERMANN, F J. X QUEBECENSIS LE PAGE T E 1965 *CISSA HERYANNvFo Jo 1967 CHIAPENSIS HEPMANN, FoJo 1968 * M I C R OPT E R A HERMANN,FoJo 1969 ORLANCEOLATA KOYAMAyT. 1971 GUAT EMALENS IS HERM~NNTF~J~ NUMBER 12 131

COLLECTOR INDEX

NUMBER DATE COLLECTED TAXON --- 2688 25 MAY 1897 BI LTMOREANA ------.-- 1843 *CANE SC E NS --- --.L CR ISTATA --- -- AUG 1827 *FLAVA -"- --c PRAIREA ABRAM S ,La 2816 31 JUL 1902 ABR4MS II ALLEN, JaAa 21A 01 JUL 1878 * PAUP ERCUL A AMANOtT a 6358 -- MAY 1951 TETSUOI ANDERSON, Ja Pa 4871 19 AUG$€938 JACOB I -PETER I ARSENEtG.(FRERE) 30 54 16 JUL 1909 ARSENII 1359 01 AUG 1907 CONSPECTA ARSENEtLa(FRERE1

-9- 28 JUL 1902 FULVESC E NS BAILEY tLaHa _-- 13 JUN 1917 CHIKUNGANA --- 18 JUL 1917 KULI NGANA 161 01 JUN 1886 *RE TROC URVA BAKER 9C. Fa 811 10 MAY 1902 AE QUA 226 -- AUG 1899 CHA LCI OLE PIS 230 -- AUG 1899 ELYNO1 DES 232 -- AUG 1899 *FESTIVA 232 -- AUG 1899 NUB1 COLA ------SAX I MONTANA BANG9 Ma 2210 --- *CL ADOSTACHYA 2376 --- *JAM€ SON I BANKS,Ja AND SOLANDER,DaC* -__ -- --- 1769 BANKS1I BARTLETT I Ha Ha 11718A 24 FEB 1931 BAxRTtETTII BEAMANTJaHa 3880 31 JUL 1960 GUATEMALENSIS BEAN9R-S.; HOSAKAvE-Y. AND STaJOHN9Ha 11228 21 DEC 1931 *W A HUENS I S B ICKNELL 9 E -P -- t 11 YAY 1904 ABDI TA --9 20 JUN 1908 INCOHPERTA B IGELOW 9 J a M a --- *- --- 18 53- 1854 ALBIDA ------1853 -18 54 *BRONGN IARTII 1547 -7- NEOMEX ICANA BI LTMORE HER BAR IUY 132 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUYRER DATE COLLECTED TAXON

B ILTMORE HER BAR IUY 262A 28 YAY 1897 *STI PATA BIOLETTI 9F.T. 1 25 JUN 1893 TUMULICOLA BOL ANDE R 9 Ho N 6213 1863 ATHROST ACHY A 4741 1866 CALIFORNICA 6477 1866 CINNAMOMEA 6477 1866 DEB1 LIFORMIS 50 FETA 3822 1864 GRACIL IOR 4700 1866 GYNODYNAMA 50 74 HAYOENI ANA 4740 1866 L(J ZUL INA 470 1 1866 YE ND OC I NE NS IS 2299 1860-1867 NUDATA 6198 1866 PAUC ICOST AT A 4746 1866 PHYL LOMANI C A 5046 1866 QUADRIFIDA 5046 1866 * QU ADR I F IDA 4702 1866 SALINAEFORMI S --- 1860 CA. SU 6-BRACT€ ATA 5086 1866 WHITNEY I 6198 1866 WHI TNEYI BOOTT p W --- 26 JUL 1865 *SCOPARI A BOURGEAUpE. --- ..- --- 1857-1859 *LONGIROSTRIS BRA INERDTE 121 19 JUL 1897 BRAINERDI I 160 18 JUL 1897 *ECHINATA 111 11 JUL 1897 *FILIFOLIA -- - 19 JUL 1898 *INTUMESCENS BRASSpLo Jo 9515 -- AUG 1938 ACROPHI LA 441 8 -- YAY-JUL 1933 BREVIS 10255 -- OCT 1938 ERE MBST ACHY A 5323 -9 SEP-NOV 1933 LAMPROCHLAMYS 9583 -- AUG 1938 PERILE I A BRASS9L.J. AND MEYER-DSEES,E. 1 98 28 -- SEP 1938 YE L ANOPHOR A BREEOLOVEpO. Eo 67 14 30 JUL 1964 CHIAPENSIS BREWER+ W. He 1650 17 JIJN 1863 ATHROST ACHY A 1422 -- --- 1863 BREWER1 1977 31 JlJL 1863 PACHYCARPA 1636 -- --- 1863 S ART W E LL IANA 1969 31 JUL 1863 * SCOPAR IA 1969 31 JUL 1863 SPECIFICA NUMBER 12 133

NUMBER DATE COLLECTED TAXON

BREWER, W Ho 1778 04 JUL 1863 WHITNEY I BRIGHT, Jo --- 18 YAY 1923 PURPURI FERA BRITTON ,No Lo --- 06 JUL 1895 * XANTHOC AR P A BROWY,O* Yo 25 5 02 AUG 1936 ROANENS IS BUCKLEY 9S.B. --- AUSTRO-CAROL INIANA --- BUCKLEY I --- CAROLINIANA --- MISERA -- - STYLOFLEXA 6URCHEL.L ,W Je 1911 BURCHELL IANA BURT-DAVYTJO 3266 25-30 JUN 1897 DAVY I BUSH9B.F. 1718 25 YAY 1902 AGGLOYERATA 1718 25 YAY 1902 AGGREGATA 2514 36 APR 1905 BUSH1 I 1043 18 MAY 1895 FISSA 7020 02 JUN 1913 LUNE LL IANA 99 3 22 MAY 1095 OKLAHOMENSIS CANBY 9 W Y 350 03 AUG 1883 MONT ANENS IS CAREY 9 J --- -- MAY 1832 CARE YANA ------PLATYPHYLL A --- --9 SYCHNOCEPHALA CARY 9 Y. 613 11 JUL 1910 ACBO-NIGRA CHAM ISSO, L Ao

_-- CIRCINNATA ~ -- - N IGR ICANS _-- PACHYSTACHYA CHAPYAN1Ao Wo --- BA LTZE LL II 113 C \A P MANI --- *DIGITALIS -- - MAGNIFOL IA -- - *STE L LULATA --- *ST IPATA CHASE, A 8283 17 JAN 1925 *PURPUREOVAGINATA CHEESEMANITOF. --- -- JAN 1803 * COMANS 83 -- JAN 1882 DEVIA --- -- JAN 1883 PETRIE I 134 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

CHRISTOPHERSENT€. 800 24 SEP 1929 SAVA II ENS1S CHUN9N.K. AND TS0,C.L. 43680 -_ --- 1932-1933 TSOI CLAUSEN9R.T. AND WAHL9H.A. 2532 06 JUN 1937 *ANGUSTIOR CLEMENSTJ. AND CLEMENSTM.S. 34297 28 JUL 1933 EXPLORATORUM 5546 -- MAR 1937 SARAW AK ETENS IS CLEMENTS, Fe ------c- 1900 EBENEA CLOKEY II We 3227 29 JUL 1918 AR AP AHOENS I S 7468 19 JUN 1937 * INTER IOR 2338 06 AUG 1915 SUBIMPRESSA 23 64 07 AUG 1915 *TRIBULOIDES COLLINS+JeFe AND FERNAL0pM.L. --* 03-17 AUG 1905 *GARBERI COLLINS,J.F*; FERNALDTM~L. AND PEASETA~S. --- 05-08 AUG 1904 *FLAVA COLLINS p Z a ------BARRATTI I COOL E Y T D 74 _-- * AL OPEC01 DEA ------COOL E Y I COVILLE 1Fe Ve 1455 11 SEP 1902 BR ACHY PODA 1457 11 SEP 1902 CAYPLYOCARPA 1362 03 SEP 1902 PACHY STOMA CRAWETI~B. AND WOODTWOA. ------WOOD1 I CRAWEt J. Be t-- q-- *OLIGOCARPA CRONQUI STTA. 2872 03 JUL 1941 OBOVOIDEA CROOM,HeBe c------1834 ALAT A CURTIS 9 M A. --- -- JUL 1841 , AESTIVALIS CURT1 SS 9 A H 3267 -- SEP 1882 CREBRIFLORA 676 1 15 APR 1901 MOHR I ANA CUSICKTW~C. 1331 -- JUL 1886 CUS ICKI I 2487 28 AUG 1900 GY MN OC L ADA 2849 3@ JUN 1902 SCABRI USCULA 1331 -- JUL 1886 *TERETIUSCULA DAVIS,€ ------HALSE YANA NUMBER 12 135

'NUMBER DATE COLLECTED TAXON

DAVIS,E --- -1 --- 1823 HITCHCOCKI ANA ------S ICCATA DEAM, C .C 54764 05 MAY 1934 *ART ITECTA 10927 05 JUN Is12 LARICINA 6458 25 MAY 1910 *LA XIF LORA 61 177 19 MAR 1941 VEXANS DELAVAY 9 ReP 15 JUL 1889 P TEROLE PTA

COLLECTA YIRABI LIS NOVAE-ANGL IAE SCHWEINITZI I TENERA T R IS PERMA *UMBELLATA VALL ICOLA

NIGRO-MARGINATA

AMPHI BOLA AUROLENSI S CRUS-CORVI FRANKLINI I HEPBURNI I MACROKOLEA PET A SATA PETR ICOSA PICTA PT YCH OC ARP A RETROF L E XA TRIANGULAR IS 08 AUG 1938 DUTILLYI ND LEP GEvE. 41, 3054 12 AUG 1963 X NEOBIGELOWI I 39329 19 AUG 1961 X PATUENSIS 41,305 12 4UG 1963 X QUEBECENSIS 39 2 74 16 AUG 1961 *VI RIDULA DUTILLYqA. i LEPAGEvE. AND DUMAN,M. 32793 29 AUG 1954 X EXSALINA 32975 03 SEP 1954 X NEOPALEACEA 32357 14 AUG 1954 *X SAXENII EASTWOOD T A 725A 16 JUL 1914 LE IOPHYLLA EASTWOODTA. AND HOWELL, JOT. 1101 23 JUN 1933 CURATOR1 UM 2271 07 MAY 1936 OB ISPOENSI S 136 SMITHSONIAN COiXTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED T AXON

EGGERT T Ho *-- 08 AUG 1893 EGGERTII EGGLESTONTW~W. 6181 22 AUG 1910 EGGLESTONI I 65 84 18 APR-25 MAY 1911 GEOPHI LA 13567 19-20 PUG 1916 MI SERABILIS 6605 18 APR-25 YAY 1911 PITYOPHILA 3329 3@ JUL-C1 AUG 1916 VIRI DIOR EKMANT EoLo 14506 21 JUL 1922 CUBENSI S --- 08 AUG 1925 *CU BENS IS H1453 12 AUG 1924 EKMANI I H10662 14 SEP 1928 *EKMANI I ELMER T A .DOE 3132 -- JUN 1901 DUOLfYI 8444 -- MAR 1907 ELMER I 88 1 -- JUN 1897 *LENT’ICULARIS 2700 -- JUN 190C OLYHPI CA 13146 -- YAY 1911 PALAWANENSIS 6983 -* NOV 1904 RHYNCHACHAE NIUM ENGELM~NNTG~ ------1874 ENGELMANNI FARGESTROP~ ------F ARGESI I FAURI E T Uo 919 28 JUN 1901 M IC RANT H A FAXONqEo AND FAXON,CoEo --- 23 JUN 1888 AENEA --- 06 JUL 1878 CRAWFORDII --- 27 YAY 1896 OR MOST ACHYA FENDLER TAO 878 -- --- 1847 FENDLE R IANA FERNALDqMoLo 264 04 JUL 1894 *CRINITA -- - 28 JUN 1904 *GLAREOSA --- 06 JUL 1904 *INTERIOR 146 06 JUL 1893 *LAXIFLORA -- - 03 JUL 1897 MERRITT-FERNALDI I I-- 05 JUN 1896 *MI RABIL IS --_ 30 JUN 1891 ORONENSIS 1464 08 JUL 1909 *SCOPARIA FERNALD,M.Lo AND BISSELLTCOH. 203 11 16 AUG 1920 *CUMULATA FERNALD1M.L. AND LONGqBo 12012 08 JUN 1940 *ABSCONDITA 12969 13 JUN 1941 X A6 SCOND I TIFORM IS 8143 08 JUN 1938 *CRINITA 12016 13 JUN 1940 *DEB1 LIS 11791 08 YAY 1940 *DIGITALIS NUMBER 12 137

.NUMBER DATE COLLECTED TAXON

FERNALDTMOL. AND LONGTB. 7767 08 APR 1938 *DIGITAL IS 11787 07 MAY 1940 RUGATA 20 2 96 14 JUL 1920 *SCOPARIA 1455 27 AUG 1929 X XANTHINA FERNALD9 YoL AND WIEGAND 9 K. Mo 2897 16 AUG 1910 *HORNSCHUCHI ANA 2897 16 AUG 1910 *HOST IANA 4918 12-13 JUL 1911 *PALL E SCE NS 42 58 16 AUG 1910 X P SEUDO-FULVA 4796 28 AUG 1911 *SCOPAR I A 2776 21 JUL 1910 WIEGANDI I FERNALD9M.L.; LONGvB. AND FOGG-JR.,JoYo 1374 20 JUL 1929 LANGEANA 1449 19 AUG 1929 X NEOFILIPENDULA 1474 31 JUL 1929 *VE S ICARIA FERNALDtYoLo: LONG980 AND SMARTTROF. 5677 22 JUN 1936 BAYARDI 5677 22-23 JUN 1936 *CR US-CO R V I FERNALD9M.L.; WEATHERBY,C.A- AND STEBBINSTGOLO 241 1 05 JUL 1931 CLIVICOLA FERNAL0,M.L.; WIEGAND7K.M. AND LONG, Be 27673 20 JSJL 1925 *LIVIDA FERNAL0,Y.L. f W IEGANDTK~Mo~LnNGt B. ; GILBERT-JR. TF~A.AND HOTCHKI SSqNo 27657 31 JUL 1925 T ERR AE-NOV AE FORDTC. -_------18833 SCAPOSA FOWLER, Jo _-- _- --- 1872 *ADUSTA ------1871 *FOENEA ------*MILIARIS ------1872 PROJECTA --- -- JUL 1870 *STRAM1 NEA FRANKTJOC. 55 -_ --- 1835 FL ACC IDULA ------_- 1835 STEUDE L II FRET2,C. Do ------1884 *GRI SEA FUNSTONTF. 139 30 JUL 1893 PYY SOCHL AEN A GARBER, AoP --- C9 JUN 1869 *AUREA --- 09 JUN 1869 GARB ER I GEYERtCoA. 332 --- GEYERI GRAIVTvG. Bo -- - 01 MAY 1902 FLACCIFOLIA GR IFF I THt Wo 78(KEW 6074) --- GR IFF ITHI I 96(KEW 6094) --t SANGUINEA 138 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

HAENKE, T ------ANTHOXANTHERA ------PRESLI I HALEID. 97 --- IGNOTA HALLrEo 5 80 -- --- 1871 *DE WE YANA 606 -- --_ 1871 HALL IANA 583 01 AUG 1871 *LEPORI NA 60 5 -- --c 1871 OREGONENSI S 583 01 AUG 1871 PHA E OC E PHAL A ------2871 VICARIA ------1872 *WILLDENOVI I HALLIE. AND HARBOURTJOP. 59 1 -- --- 1862 WONPLANDII 61 7 -- --- 1862 HALL II 59 1 -* --- 1862 ILLOTA 587 -- -_- 1862 VIOLkCEA HALLTHOYo 2483 -- JUL-AUG 1901 JACINTOENS IS 9781 25 JUL 1914 TENERAEFORMI S HALLTHOY. AND BABCOCK,HoDo 5472 -- JUL 1904 LANCIFRUCTUS HALL9H. Me AND CHANDLERTHOAO 4716 @I-02 AUG 1903 LEPORINELL A HARPER, RoMo 2109 09 APR 1904 HARPERI 21 59 26 APR 1964 SMALLIANA H ARTMAN T C .V 620 12 APR 1891 CHIHUAHUAENSI S HASSE,H.Eo --- -- JUL 1894 HA SSE I -c- 01 YAY 1886 +ROSEA HAY,G.Uo 84 24 JUL 1900 *CANE SC ENS HAYDENvFo Vo -- - _- -_- 1853 *ARISTATA 580 --* *DOUGLAS1 I 21 -- --- 1853-1854 HAYDENI I ------’ LAEVI-CONICA ------MEEK11 ------NE BRASK ENS IS -- - 20 JUN 1860 RAYNOLDSI I HELLER, A. A. 10820 07 JUN 1913 ABRUPT A 9429 11 AUG 1908 FISSURICOLA 9975 23 JUL I910 HELLERI 9841 16 JUL 1909 INTEGRA 5797 01 JUL I902 LACUNARUM NUMBER 12 139

NUMBER DATE COLLECTED TA XON

HELLER 9 A. A a LO052 20 MAY 1910 LE PTOPODA 71 87 17 AUG 1903 *LU ZU LAEFOL IA 9067 21 JUL 1908 MICROPTERA 71 87 17 AUG 1903 PSEUDO JAPONI C A HENDERSON,LaF. 13 -- _-- 1883 ACUT INELLA 5583 25 JUL 1925 EA ST WOOD IANA _-- -- JUL 1884 INOPS 140 2 31 JUL 1886-20 AUG 1887 PANSA HENRY 9 A. 7860 _- *-- 1855-1888 ICHANGENSIS 5467 t-- LA NC IFOL I A 4266 _-- *LONGICRURI S 10839 --- PRAINI I HENRY 9 J eK. 91 52 04 JUN 1915 ARCTAEFORMIS HERMANNt Fa Ja 13498 28 4UG 1956 ATHABASCENSIS 17059 15 AUG 1961 * 6I PART I TA 6147 05 JUL 1934 X DEAMII 13529 28 AUG 1956 EUR Y STACHYA 13347 15 AUG 1956 INCOND ITA 7985 13 JUL 1936 *INTER I OR 12252 21 AUG 1955 LIHNOPHI LA 5983 15 AUG 1933 PELOCARPA 18120 21 AUG 1962 P LEC T OC ARP A 13453 26 AUG 1956 *PR AT ICOLA 6408 14 AUG 1934 *VULP IN01OE A HOHENACK ER 9 R a F. 943 -- --- 1851 MERCARENSIS HOLLI STERvNa 14 C5 AUG 1911 ATROSQUAMA HOSAKA, E aY a 594 04 JUL 1932 * PLUV ICA HOUGHTON 9 Oa --_ 13 JUL 1832 HOUGHTON IANA HOWE 9 E a C a --- 30 MAY 1887 ROSAEOIDES HOWELL 9 Jafa 14546 11 AUG 1938 D'ANAENS IS 24609 23 JUN 1948 SPECUICOLA 14519 11 AUG 1938 SUBN IGRICANS 35333 06 JUN 1960 TOMPKI NSI HOWELL,J.Te AND STACEYIJ~W~ 13Q42 06 JUN 1937 SONOMENSI S HOWELLvTaJa --- -- MAY 1880 AGC EDE N S 93s 09 MAY 1885 ACUT INA 2994 -- MAY 1886 B RE V ICAUL IS 140 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

HOWELL 9 ToJ 93 7 27 YAY 1885 *MARC IDA --- -- MAY 1880 SPRE TA --- -- YAY 1880 *STY LOS A INGALtS ,To ------TURGESCENS JAMEStEDWIN ------JAMES1 I JEPSONr Wo Lo 4477 20 JUL 1911 JEPSONI I 4476 20 JUL 1911 MAR1 PQSANA JOHNSON t We Mo 594 14 AUG 1967 *MICROPTERA JOHNSTON? IoYe 1505 31 JUL 1917 STENOPTERA JONES?M oE --- 12 JUL 1899 ABOR IG I NUY 834 28 AUG 1878 CRANDALL II --- OR JUL 1909 ELRODI 5345 01 JUN 1874 EPAPILLOSA --- 22 JUL 1881 JONES1 I KELLOGG? A. --- 03 AUG ---- HETERONEURA --f --- KELLOGGI I --- 10 JUL 1870 NERVINA --- _-- SU6 F USCA KELLOGGqA. AND HARFORD, WeGoUo 1073 -- --_ 1868-1869 *FESTIVA 1073 -- --- 1868-1869 HARFORDI I 1@69 -- --_ 1868-1869 *HOOD II 1069 -- --- 1868- 1869 *HOOD1 I YEL SO, L 4967 C1 AUG 1945 ELB E RTAN A 6362 24 JlJL 1948 ERXLEBENIANA 6058 03 JUL 1947 UNCOMPAHGRE KELSOtL AND KELSOr E.Ho 525 08 AUG 193-6 HAG1ANA KENNEDY ,Go Go --- 11 JUN 1899 *VE ST ITA KINASHI T No --- -- JUL 1909 ’ CUNEATA KOYAMApT --- 23 NOV 1958 TA MAK II LEAVENWnRTHt MoCo ------_- 1846 F LACC OSPERMA ------1845 LEAVENWORTHI I LEAVENWORTHI MeC AND HALE 7 D 683 --- HALE I LECHLERqWo 1136 -- OCT 1852 INC I SO-DENTATA NUMBER 12 14 1

NUMBER DATE COLLECTED TAXON

LEIBERG9 JOB. 33 5 26 JUN 1894 PHAEOL EPI S 12 5 10 JUL 1895 PRIONPHYLLA 2556 19 JUL 1896 VAGANS LEMMONTJOG. ------1875 LEMMONI 2 21 JUN 1882 ULTRA LEPAGE t E 32078 30 JUC 1954 X DUMANII 33131 27 JUL 1955 X NUBENS LIPSKY, V. I 2732 19 JUL 1899 PH I L OC R E N A LITVINOV,D.I. 153 -- --- 1897 *STENDPHYLLA LONG,Be F23212 15 JUN 1929 CAESARIENSIS LYALL TOAVID --- -- _-- 1861 *ACUTA YACBRIDF9J.F. AND PAYSON9E.B. 3778 13 AUG 1916 PROPOS ITA YACDANIELS,LeH. 1542 15 MAY 1027 TAHITENSIS MACKEYZIE,K.K. 2676 23 JUN 1907 A€ STIVAL IF ORMI S 2088 10 JUN 1906 CONVOLUTA 4645 26 JUN 1910 CRYPTOLEPI S *-- 30 YAY 1897 MOLE STA 167 -- AUG 1-901 PERGL 0 60s A 9871 -- MAY 1911 RUGOSPERMA YACOUNIJ~M~ 33728 12 JUL 1901 *SC IRPOIOEA MACOUN, JOHN 134Ql 26 JUL 1887 ABLATA s2 14 JUN 1879 ASS INI BOINENS I S 6 01 AUG 1882 *CRAWFORD11 1665 12 AUG 1872 ELEOCHAR IS --- 31 JUL 1891 INCURVIFORMIS 53877 14 AUG 19n2 M I CROC HAE T A 22 04 JUL 1899 *MIR4BILIS --- 31 MAY 1887 PIPERI _-- 31 MAY 188P *PRATENSI S 26624 24 JUN 1901 *STRAYINEA --- 04 JUL 1879 YERANTl CA MAGUI RE Be 16098 05 AUG 1938 INT E RI MUS MAGUIREt 8. AN0 HOLMGSEN9A.H. 21947 16 JUL 1943 *CAMP Y L OC A R P A MAGUIRE t 6. AN0 MAGUIRE ,Re Re 14668 16 AUG 1936 RACHILL IS 142 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUYBER DATE COLLECTED TAXON

MAGUIREt 6.; H0BSONtD.A. AND YAGUIREIR~R. 14013 16 JUL 1936 WERNACULA MANDON t C 1429 --- YANDONIANA YARIE-VTCTORINT(FRERE~

4021 0- JUN 1917 *PAUPERCULA MAR IE-V I CTORIN 9 (FRER E 1 AND ROLL AND-GERYAIN T ( FRER E 1 25767 20 JUL 1926 *INFL AT A MCCALLA t W .C 2348 28 JUL 1899 SC IRP IFORM1 S MEAD, So Be --- -1- ILLI NOENSI S -_- --c MEAD II --- *-- *RICHARDSON II MERRILL t Eo Do 10985 12 AUG 1917 BAMBUSETORUM 6505 -* YAY 1909 * BRU NNE A 6623 -- MAY 1909 YERRILLII 543 -- APR 1910 PYCNOTHYSOS 4730 -- OCT-NOV 1905 SU BT R ANS V ERS A MERTENST CoHo ------LEIOCARPA MOHRTC. --- 26 APR 1897 * STIP ATA YOOREfJo A. AND STEYERMARKTJOA. 3625 25 JUL 1931 MUR ICU L AT A MULLER9C.H. 3520 27 SSP 1939 P E RC 0s TAT A MULLERTC~HOAND MULLERTY~T. 892 28 JUN 1934 MACKENZIANA MUTI S 9 J Co KILLIP NO05715 _- --_ 1760- 1808 *PIRCHINCHENSIS NELSON9 A. 7124 11 JUN 1900 BREVISQUAMA 32 75 02 JUL 1897 FESTI VELLA 73 16 29 JUN 1900 S IMULATA NELSON,A. AND YACBRIDETJ.FO 1533 01 AUG 1911 APODA NELSONTA. AND NELSONTEO 5264 30 AUG 1898 NE LS ON1 I NUTTALL ,T 17 --- NUTTALL II OHHI, Jo 4182 -- JUL 1933 A PODOST ACHY A 329 -- MAR 1933 HATUS I MANA 813 11 AUG 1931 KURI LENS1S 29 04 MAY 1931 RUGATA OLNE Y T S 1 --_ 01 JUL 1867 HORYATHODES NUMBER 12 143

NUMBER DATE COLLECTED TAXON

PAINEt J c A. --_ __- *VAGINA TA PALMERTE. 546 -- JUN 1890 *NUDATA PAR ISH 9 So R 2485 03 JUL 1892 AUSTROMONTANA 3609 -- JUN 1895 MU LTI C OSTA TA 41 44 04-13 ApR 1896 VITREA PARRY qC Co ------18SO BARBARA E PARRY9C.C. AND LEMMONt JOG. 39 6 -- --- 1876 ALMA PAYSONTE~B~AND PAYSONTL-B. 2224 06 AUG 1920 PAYSON1 S PEASEpAoSo AND LONGvB. 2C519 09 JUL 1920 *LASIOCARPA PECK1Co Ho --- -- JUN 1894 *ROSE A 5 -- JUN 1893 *ROSEA --- -- JUN 1894 *ST IPATA --- -- JUN 1892 *TORTA PECK TM. IF. 13 13 AUG 1917 PRAECEPTOR IUM PETELOT CP A. 5325 -- JUL 1930 TR ICHOPHYLLA PETRIEt Do _-- _-- KALOIDE S --- -- JAN 1880 LONGICULMIS PITTIERqH. AND TONDUZrA. 3376 19 JAN 1891 DURANDI I 3381 19 JAN 1891 *LE MANNI ANA PORSIL0,A.E. 173 26 AUG 1937 MORRI SSEYI PORSILD,A.Eo AND P0RSILDqR.T. 5120 02 AUG 1928 *ATROFUSCA 71 1 23 JUN-05 JUL 1926 KOKR INENSIS 713 23 JUN-05 JUL 1926 MELOZITNENSIS PORTER 9 T C --- --_ *QEBILIS --- 26-29 JUN 1871 HALL I I -- - 28 AUG 1871 PORTER1 PREISSTLo 1825 --_ PREI SSI I 1861 -- JUL 1839 PREISSI I PRINGLE TC. Go 4275 03 OCT 1892 AUTUMNAL1 S 4839 19 AUG 1894 AZTECICA 10039 26 AUG 1905 C I LIARI S 4838 25 4UG 1894 FELIPENSIS 144 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

PRINGLE T C G. --- 23 AUG 1881 FRACTA 48 39 19 AUG 1894 FUSCOT I NCT A 3126 08 JUL 1890 INVOLUCRATELLA 40 40 27 AUG 1894 MACROSP ERMA 4842 29 AUG 1894 OAXACANA 8863 02 JUN 1904 PERLONGA 2630 05 JUN 1889 PERSTRICTA 4685 08 JUN 1894 *PI NETORUM 3801 04 AUG 1891 PR 1NGLE I --- 19 AUG 1881 *SCIRPOIDEA 7452 24 JUN 1897 SEATONIANA PYRON, J.H. AND MCVAUGHTR~ 2951 15 MAY 1938 AM PL ISQUAM A RAMOSTM. BUR. SCIo 1434 -- AUG 1906 RAMOSI I RAUPTH. M. AND SOPER 9 J.H. 9534 18 dUL 1939 SOPERI RAY 9 JoD i WOOOTC E : S #IT HI A .C AND EATON ,Re J 10750 26 APR 1961 *FISSA R ICH, W P --- 05 JUN 1894 *TEN ERA R ICH9 WILL IAM WILKES EXPED. 1241 -- --- 1838-1842 LAC1NI ATA RICHARDSON, Jo _-- --- ARCTICA 417 --- BACKANA 1- - --f BACK II OUR I FOL IA FE ST IVA HUOKERANA PARRYANA PODOC ARPA -- - --9 RICHARDSON1 I ROACH, A. W 20 2 10 JUN 1949 DIVERSISTYLIS ROCK,J.F. 9017 -- OCT 1909 KA UA IE NSIS ROSE, J. No 2357 16 AUG 1897 ,MADRENS IS R0SEqJoN.i PAINTER,JoH. AND ROSE, J.S. 90 19 -- 1905 ST ELLAT A RDUSSEAUv J 24989 27 JUL 1926 *OEDERI RUSBY 9H.H. 859 -- --- 1883 RUSBYI RUTH, A. 360 12 AOR 1913 BULBOSTYLIS 45 8 24 APR 1914 ONUSTA --- -- JlJL 1900 RUTH I I NUMBER 12 145

.NUMBER DATE COLLECTED TAXON

RYDBERG9P.A. 23 39 07 AUG 1895 IDAHOA SANDBERG9 J*H* 933 20 AUG 1892 *HINDS1 I SANDBERG9J.H. AND LEIBERGIJ.~~ 194 09 JUN 1893 *NEBRASKENS IS 773 18 AUG 1893 NEUROPHORA

SARTWELL? HOP 0 -- - 7-- AL OPEC0 IDEA 56 --- AQUATIL IS 56 --- *AQUATICIS ------*CEPHALOPHORA 108 -- --- 1848 C OMMU N IS 78 --- *CR IN ITA ------FORMOSA 36 --- INTER IOR 12 --- SARTWELLI I 72 t-- SCABR IOR 35 -v- *STERI LIS 138 --- * TE NTAC ULATA SAVATI ER TL. 1414 -- --- 1866-1 874 MACR OGLOS S A 1404 -- _-- 1866- 1874 *NUTANS 2959 -- --- 1866 -1 874 PLANATA 1413 -- --- 1966-1 874 PODOGYNA SCHAFFNERpJoG. 22 1 -- --- 1877 FUSCOL UTE A 5 46 _- --- 1877 POTOSI NA 546 -- --- 1877 SCHAF FN ER I SCHNEIDERTCO 2738 06 AUG 1914 SCHNE IDER I SCHNEIDER TR. A. 954 25 JUL 1938 *EGGLESTONI I SCHWEINITZ 9LoDo --* --- COSTATA --- q-- GRAC ICL I MA SCOTT 9 ( MISS 1 --- -_ --- 1880 PRAEGRACIL IS SCOULERTJ ------A+ ER TA --- 0-0 COLUMB IANA 296 --- SC OUL ER I SEK IMOTO T Ho --- 15 JUL 1932 HY ME NO DON SHARP9A.J. 45450 29 APR 1945 AT R ACT0 D ES 45 144 07 FEB 1945 QU ICHENS IS SHELDON1E.P. 88 54 09 SEP 1897 SHELDON II 146 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

SMITH9C.P. 1055 24 JUL 1905 MONT EREY ENS IS SYITH9J.De AND TURCKHEIM9H. 659 -- JUN 1885 DONNELL-SM ITHII SMITHtSeJ. AND DUNCAN9W.H. 4872 06 APR 1949 *WILLDENOW1 I STo JOHNvH. 18330 18 AUG 1937 VITIENSIS STeJOHN9He AND FERNALD9M.L. 1C801 16-17 JUL 1914 MISANDROIDES STEELE, EeSe ------0- 1896 ANGUST IOR _-- 23 YAY 1898 MEDITERRANIA --- 0- f-- 1900 ME SOCHUREA STEYERMARK9J.A. 53105 15 JUN 1943 AZU AY AE 48347 07 JUL 1942 CUCHUMATANENSIS 62605 06 MAY 1945 CULMEN ICOLA 49055 18 JUL 1942 HUEHUETECA 55470 11 FER 1944 LARENSI S 5887C 28 SEP 1944 RORAIMENSIS 48 542 14 JWL 1942 STEYERMARKI I 57367 15 JUL 1944 TACH IRENS1 S 5740 1 15 JUL 1944 TAMANA 50 150 06 AUG 1942 TOJQUI ANENS IS 53095 15 JUN 1943 TO RE ADOR A 48334 07 JUL 1942 TUNI MANENS IS 62705 10 MAY 1945 TURUMIQUIRENSIS 48554 14 JUL 1942 VENOSI VAGI NATA SUKSDORF9W.N. 12347 --- *APERTA 12348 15 SEP-23 OCT 1927 *AP ER TA 12359 23 OCT 1927 *APERTA 6864 16 4UG 1909 CUN STANCEANA 51 81 15 JUL 1905 EGREGIA 1284 26 JUN 1386 EU RY C A RP A 11551 21 AUG 1924 *EURYCARPA 12333 22 AUG-05 SEP 1927 * INTERRUPTA 10 249 22 JUN 1919 *NEBRASKENS IS 81 6 02 JUN 1885 OXYC ARPA 1296 13 AUG 1897 PADDOENSIS 1315 -- JUL 1883 S UBOR 8 ICU L AT A 7383 27 AUG 1912 SUKSDORF I I 5259 21 AUG-20 SEP 1905 *SUKSDORFII 5 ULL I VANT 9 W S --- -c1 *ALATA --- --_ CONJUNCTA SVENSON T H 9 K 10469 12 JUL 1939 *oXY L EP IS NUMBER 12 147

,NUMBER DATE COLLECTED TAXON

TAAMpYo Wo 50 2 1-16 APR 1938 SURCULOSA TAKpToWo AND CHOWqWoKo 3202 20 NOV 1926 OBL ANC EOL ATA THURBERt Go 652 --- *GAYANA --- -- JUN 1850 T HUR 6 ER I TORREVT~. _-- 03 AUG 1839 HETEROSTACHYA TOWNSfjNDtC.H.T* AND BARBES9C.M. 157 21 JUL 1899 TOWNSENDI I 1RACYpJ.P. 4547 04 JUL 1914 TRACY I 3783 21 JUL 1912 UNILATERALIS TRACY p So Mo 17 23 MAY 1888 *STRAMINEA TSAI ,HOT. 62809 _- --- 1934 ZIZANIAEFOLI A TSUI p T oM 74 -- YAR-APR 1932 *RUBRO-BRUNNEA TUCKERMANqEo ------AR GVRANTHA --- -- JUN 1864 GL AUCODEA ------*SCOPARIA ---- _--- * SCOPAR IA UN D E R NO 0 0 9 L Y 158 29 JAN 1903 UNDERWOOD1 I VAHL T Jo t-- --- PR AT ENS IS VASEY pGm ------*HIRSUTA VENTURI 9 So 6491 15 FEB 1927 * PHA L ARO I DE S 8650 -- --- 1929 SALTAENSIS VREELANDIF~Y~ 1121 19 AUG 1901 HOLM1 ANA WAREpRo A. t ROLLINS 7s. AND KNIGHftO.Wo 5066 05 JUL 1906 *TR ISPERMA WATERFALL tU.Te 11380 19 APR 1953 L~TEBRACTE AT A HATSONIS. 1231A -- AUG 1869 WANE SC ENS 1246 -- MAY 1868 WATSON I WERDERMANNp E. 1687 -- MAR 1925 WERDERMANNII WHEELER9C.F --- 28 JUN 1890 *TENUI F LORA HIEGANDt Ko Mo AND THOYAS ,CoCo 1915 15 JUN 1914 *AMPHIBOLA 148 SMITHSONIAN CONTRIBUTIONS TO BOTANY

NUMBER DATE COLLECTED TAXON

WILKES EXPLOR. EXPED. --- -* --_ 1838- 1842 HEBE TATA _I------1838-1842 *MICROGLOCH IN -7- -0 --- 1 83 8- 1 842 OL IGANTHA -- --- 1838-1842 WILKESI I WILL IAYST E F --- -- AUG 1900 *VESICARI A WILLI AMS T Em F. 1 CHURCHILL, JeR. AND FERNALOt MoL --- 16 Jut 1900 KATAHDINENSIS WILLIAMS 9 €0 F. i COLLINS7J.F. AND FERNALDTM-L. 12-15 JUL 1905 *DE WE YANA 110 19 JUL 1992 X TRICHINA WILLIAMS ~L.00 13178 06 JUL 1947 ST ANDLEYANA WILLIAMSTR.S. 18 JUN 1899 BONANZENSI S 07 JUN 1893 CONCINNOIDES --- 11 AUG 1894 STANTONENSIS --- 12 JlJN 1899 WILL IAMSII --- 18 JUN 1899 YUKONENS IS WILL IAMS TTa A 2951 19 AUG 1897 PLATYLEPIS WOODtW. A. --- RETROC URVA WOOTON, Eo0. --- 28 JUL 1900 AGROSTOIDES W R I GHT 9 C --- CONF ERTIFLORA -T - MICANS NA NA PAPULOS A PARCIFLORA PICTA WRIGHT1 I

*STRICTA XEROCARPA NUMBER 12 149

GEOGRAPHIC INDEX

COUNTRY STATE TAXON

AFGHANISTAN GR IF F ITHI I S A NGU IN E A ARGENTINA JUJUY *PHALAROIDES SALTA SALTAENSIS TIERRA DEL FUEGO (TER BANKS1I AUSTRAL IA PREISSI I WESTERN AUSTRAL IA PREISSI I BOLIVIA *C L ADO STAC H Y A * JAME S ON I MANDONIANA BRAZIL RIO DE JANEIRO *PURPUREOVAGI NATA BRITISH HONDURAS CAY0 BARTL ETT II CANADA PARRY ANA PODOCARPA RICHARDSON1 I ALBERTA ATHABASCEN S I S ALBERTA ATROSQUAMA ALBERTA EURYSTACHY A ALBERTA INCONDI TA ALBERTA INCURVIFORMI S ALBERTA *PR AT! COLA ALBERTA SC IRPIFORMIS BRITISH COLUMBIA ABLATA BRlTISH COLUMBIA ARCT AE FORM IS BRITISH COLUMBIA PIPER1 BR IT1SH COLUMBI A *PRATENS IS BRIT1SH COLUMBIA *SCIRPOIDEA MANITOBA ASSINIBOINENSIS MAN IT06 A DUTILLYI MAN ITOBA *LONGIROSTRIS NEW BRUNSWICK *ADUSTA NEW BRUNSWICK *C ANESCEN S NEW BRUNSWICK *FOENEA NEW BRUNSW ICK *MIL IARI S NEW B RUN SWI CK *MIRABILIS NEW BRUNSWICK PROJECTA NEW BRUNSWICK *STRAMINEA NEWFOUNDLAND *HORNSCHUCHIANA NEW FOUN DL AND *$IOSTI ANA NEWFOUNDLAND LANGEANA NEWFOUNDLAND *LIVIDA NEW FOUNDLAND MISANDROI DES NEWFOUNDLAND MORRISSEYI NEWFOUNDLAND X NEOFILIPENDULA NEW FOUNDLAND *PALL E SC ENS NE WFOUNDLANO X PSEUDO-FULVA NEWFOUNDLAND *SCOPARI A NEWFOUNDLAND TERRAE-NOVAE NE WFOUNDLAND *VESICARIA 150 SMITHSONIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE TAXON

CANADA NEWFOUNDLAND WIEGANDII NEW FOUNDL AND X XANTHINA NORTHWEST TERRITORIES *ATROFUSCA NORTHWEST TERRITORIES SOPERI NOVA SCOTIA *CUMULATA NOVA SCOTIA *LASIOCARPA NOVA SCOTIA *SCOPARIA ONTAR I0 *STRAMINEA QUEBEC CLIVICOLA QUEBEC *CRAW FORD1.1 QUEBEC *DEWEY ANA QUEBEC X DUMANII QUEBEC X EXSALINA QUEBEC *FLAVA QUEBEC *GARB ER I QUEBEC *GLAREOSA QUEBEC *INFLATA QUEBEC X NEOBIGELOWII QUEBEC X NEOPALEACEA QUEBEC X NUBENS QUEBEC *OEDERI QUEBEC X PATUENSIS QUEBEC *PAUPERCULA QUEBEC X QUEBECENSI S QUEBEC *X SAXENII QUEBEC *VI RIDULA SASKATCHEWAN ARCTIC A SASKATCHEWAN BACKANA S A SK A TC H E WAN BACK1 I SASKATCHEWAN OUR1 FOL IA SASKATCHEWAN ELEOCHAR IS S A SK A TC H E W AN HOOKERANA SASKATCHEWAN XERANTI CA YUKON TERRITORY BONANZENSIS YUKON TERRITORY LEIOPHYLLA YUKON TERRITORY MICROCHAET A YUKON TERRITORY WILLIAMS11 YUKON TERRITORY YUKONENSIS CHILE INCI SO-DENTATA LLANQUIHUE WERDERMANNII MAGALLANES vMICROGLOCH1 N MAGALLAN E S OLIGANTHA CHINA HUPEH ICHANGENSI S HUPEH LANCIFOLIA HUPEH *LONGICRURIS HUPEH AND HONAN CHIKUNGANA KIANGSI KUL INGANA K WANGTUNG B AM BUS ETORUM KWANGTUNG OB LANC E OLA T A NUMBER 12 151

COUNTRY STATE TAXON

CHINA KW ANGTUNG *RUBRO-BRUNNEA KWANGTUN G SCAPOSA KW ANG T UN G SURCULOSA KW ANG TUN G TSOI SZECHWAN FARGESI I YUNNAN PRAfNII YUNNAN PT EROL EPTA YUNNAN SCHNEIDERI YUNNAN Z IZ AN IAE FOL IA COLOMBIA *PIRCHINCHENSIS COSTA RICA DURANDI I *LEMANNIANA CUBA OR IENTE CUBENSIS ECUADOR AZUAY AZUAYAE AZUAY TOREADORA FIJI VITIENSIS FRENCH POLYNESIA TAHITENSIS GREENLAND PRATENS IS GUATEMALA ALTA VERAPAZ DONNELL-SMITH1 I HU EHU ETENAN GO CUCHUMATANENSIS HUEHUETENANGO GUATE MAL ENS IS HUEHUETENANGO HUEHUETECA HU EHU ElE NANGO STEYERMARKII HUEHUETENANGO TOJQU IANENS IS HUEHUETENANGO TUN1 MANENSI S HUEHUETEN ANGO VENOSIVAGINATA JALAPA STANDLEY ANA QUICHE QUICHENSIS HAITI *CUBENS IS *EKMANI I OUEST EKMANI I INDIA MERCARENSI S INDONESIA WEST NEW GUINEA ACROPHI LA WEST NEW GUINEA ERE MOSTACHYA WEST NEW GUINEA MELANOPHORA WEST NEW GUINEA PERILEI A JAMAICA UNOERWOODI I JAPAN HYMENODON MICANS RUGATA AOMORI (PREFECTURE) CMNEAT A HOKKAIDO (PREFECTURE) CONFERTIFLORA HOKKAIDO (PREFECTURE) NANA HOKKAIOO (PREFECTURE) PAPULOSA HOKKAIDO (PREFECTURE) PARCIFLORA HOKKAIDO (PREFECTURE) PICTA KANAGAWA (PREFECTURE) MACROGLOSSA KANAGAWA (PREFECTURE) *NUTANS KANAGAWA (PREFECTURE) PLANATA 152 SMITHSONIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE TAXON

JAPAN TOKYO (PREFECTURE) PODOGYNA KOREA MICR ANTHA MALAYSIA SABAH (TERRITORY) EXPLORATORUM MEXICO CHIAPAS ATR AC TOD E S CHIAPAS CHIAPENSIS CHIHUAHUA CHIHUAHUAENS IS CHIHUAHUA PERCOSTATA CHIHUAHUA TOWNSENDII DURANGO MADRENS IS HIDALGO CILIARIS HIDALGO PERLONGA HIDALGO SEATONI ANA HIDALGO STE LLATA MEXICO AUTUMNAL IS MICHOACAN ARSENI I NUEVO LEON *EGGLESTONII NUEVO LEON MACKENZIANA NUEVO LEON PERSTRICTA OAXACA AZTEC ICA OAXACA FELIPENSIS OA XAC A FUSCOTI NCTA OAXACA MACROSPERMA OAXACA OAX AC AN A OAXACA *PINETORUM PUEBLA CONSPECTA SAN LUIS POTOSI FUSCOLUTEA SAN LUIS POTOSI INVOLUC RAT ELLA SAN LUIS POTOSI POTOS INA SAN LUIS POTQSI PR INGLEI SAN LUIS POTOSI SCHAFFNERI SONORA *GAYANA SONORA THURBERI NEW ZEALAND CANTERBURY (DISTRICT) *C OMAN S CANTERBURY (OISTRICT) PETRIEI NELSON (DISTRICT) DEVIA OTAGO (DISTRICT 1 KALOIDES OTAGO (DISTRICT) LONGICULMIS PAPUA AND NEW GUINEA NORTH -E A ST NEW GUINEA SARAHAKETE NSIS PAPUA (TERRITORY] BREVIS PAPUA (TERRITORY) LAMPROCHLAMYS PERU LIMA HE B E TATA PHILIPPINES BATAAN RHYNCHACHAENIUM BENGUET *BRUNNEA B ENGU ET ELMER1 BENGUET MERRILLII BENGUET SUBTRANSVERSA NEGROS OCCIDENTAL P Y CNO THY SO S PAL AWAN PALAW ANENS IS RIZAL RAMOSI I NUMBER 12 153

COUNTRY STATE TAXON

RYUKYU ISLANDS OKINAWA (PREFECTURE) TAMAK II OK1NA WA ( PREFECTURE 1 TETSUOI SOUTH AFRICA BURCHE LL IANA ST. PIERRE AND MIQUELON FULVESC E NS SW ITZERLAND VALAI S (CANTON 1 *FLAVA TAIWAN APODOSTACHYA FUKIEN HATUSIMANA USA *CANESCENS COLUMBIANA FRANKLIN11 GEYERI JAMES II NUTTALL II PETAS AT A PETRICO SA *UMBELLATA ALABAMA *ST IP ATA ALASKA ANTHOXANTHERA ALASKA C IRC INNATA ALASKA JACOBI-PETER1 ALASKA KOKRINENSIS ALASKA LEIOCARPA ALASKA MELOZITNENSI S ALASKA NIGRICANS ALASKA P ACHY S T ACHY A ALASKA PHYSOCHLAE NA ALASKA PRESLII ARIZONA CURATORIUM AR IZONA *NUDATA ARIZONA RUSBY I ARIZONA SPECUICOLA AR IZONA ULTRA ARKANSAS BUSH1 I ARKANSAS *ROSEA BRITISH COLUMBIA SC OULER I CALIFORNIA ABRAMS II CALIFORNIA ABRUPTA CALIFORN IA AEQUA CALIFCRNIA ALBIOA CALIFORNIA ALMA CALIFORNIA RTHROSTACHYA CALIFORNIA ATHROST ACHY A CAL IFORN IA AUSTROMONTANA CALIFORN IA BARBARAE CALIFORNIA BRAINERCII CALIFORNIA BREWER1 CALI FCRNIA *BRONGNIART II CAL IFORN IA CALIFORNICA CALIFORN IA CINNAMOMEA 154 SMJTHSONIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE TAXON

USA CALIFCRN IA DANAENS I S CALIFORNIA DAVY I CAL IFORN I A DEBILIFORMIS CALIFCRNIA CUDLEY I CALI F ORNI A *ECHINATA CAL IFORN IA *FE ST IVA CALIFORNIA FETA CAL IF ORN IA *FILIFOLlA CAL IFCRN IA FLACC IFOLIA CALIFCRNIP FRACT A CALIFORNIA GRAC I LIOR CALIFCRNIA GYNODYNAMA CALIFCRNIA HARFORDI I CAL IFORN IA HASSEI CAL IFORN IA HAYDENIANA CALIFORNIA HETERONEURA CALIFCRN IA *HOOD II CALIFGRNIA *HOOD II CALIFORNIA INTEGRA CALIFORN IA JACINTOENSIS CALIFCRNIA JEPSONI I CAL IFCRN I A JONES I I CALIFORNIA KELLOGGI I CALIFCRNI A LACI N IATA CALIFORN IA LACUNARUM CALIFORNIA LANC IFRUCTUS CALIFCRNI A LEMMONI CALIFCRNIA LEPORINELLA CALIFGRNIA *LUZULAEF OL I A CALIFORN I A LUZULI NA CAL IFORN IA MAR1 POSANA CALIFCRNIA MENDOC INENS IS CALIFORNIA MONTEREYENSIS CAL IFORNI A MULTICOSTATA CALIFCRNIA NE RV IN A CAL IFORN IA NUDATA CAL IFORNI A OBISPOENSIS CALIFCRN IA PACHY CARPA CALIFORNIA PAUCICOSTATA CALIFORNIA PHYLLOMANI C A CALIFCRNIA PRAEGRACILIS CAL IFORN IA P SEUD 0 J A PONI C A CALIFORNIA OUADR IF IDA CALIFORN IA *OUAORI F IDA CAL IFORN IA SALI NA EFORM IS CALIFORNIA SARTWELL IANA CALIF ORN IA *SC IRPOI DEA CAL IF ORN IA *SCOPARI A CALIFCRNIA SONOMENSIS NUMBER 12 155

COUNTRY STATE TAXON

US A CAL IFORNI A SPEC1 FICA CALIFORNIA STENOPTERA CALIFCRNIA SUB-BRACTEATA CALIFORNIA SUBFUSCA CALIFCRNIA SUBNIGR ICANS CALI FCRNI A TENERAEFORMIS CAL IFCRN IA TOMPKI NSI CALIFCRN IA TRACY I CALIFCRNIA TUMUL ICOLA CALIFCRN IA UNILATERAL1 S CALIFCRNIA VITREA CALIFORNIA WHITNEY I CALIFORNIA WHITNEY I CALIFCRNIA WHITNEY I CALIFORN IA WILKESI I COLOR ADO ARAPAHOENS IS COLORADO *BIPARTITA COLORADO *BONPLANDI I COLOR bD0 CHALC IOLEPI S CO L OR ADO CRANDALLII COLORADO EBENEA COLOR ADO EGGLESTONII COCORADO ELBERTANA COLORADO ELYNO IDE S COLOR ADO ENGELMANN I COLORADO ERXLE BENIANA COLOR ADO *FESTIVA COLOR ADO HAG1ANA COLOR ADO HALL1I COL OR ADO HEPBURNII COLORADO ILLOTA COLOR ADO *MI CROPTERA COLOR ADO NUBICOLA COLORADO PERGLOBOSA COLORADO SAXIMONTANA COLORADO UNCOMPAHGRE COLOR ADO V IOLAC EA DISTRICT OF COLUMBIA ANGUSTIOR DISTRICT OF COLUMBIA MEDITERRAN IA DISTRICT OF COLUMBIA M E$ OC HOR E A FLORIDA BALTZELLII FLORIDA CHAPMAN1 FLORIDA CREBRIFLORA FLORIDA *DIGITALIS FLORIDA *FISSA FLORIDA FLACCOSPERMA FLORIDA MAGNIFOLIA FLORIDA MOHRI ANA FLOR IDA *STELLULATA 156 SMITHSONIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE T AXON

USA FLORIDA *STI PATA FLORIDA VEXANS GEORGIA AMPLI SQUAMA GEORG IA HARPER I GEORGIA SMALLIANA GEORGIA *WI LLDENOWI I HAWAII KAUAIENSIS HAWAI I *PLUVICA HAWAI I *WAHUENSIS IDAHO ABOR IGI NUM IDAHO APODA IDAHO HALL1I IDAHO *HINDS11 IDAHO IDAHOA IDAHO OBOVOI DEA IDAHO PR IONPHYLL A IDAHO PROPOSI TA IDAHO RAYNOLDSII ILLINOI S *HI RSUTA ILLINOIS ILLINOENSI S ILLINCI S MEAD1I ILLINOIS *RICHARDSON11 ILLINOIS SUB1MPRESSA ILLINOIS *TRIBULOIDE S INDIANA *ARTITECTA INDIANA X DEAMII INDIANA LARICINA INDIANA *LAXI FLORA KANSAS MOLESTA LOUIS IANA AMPHI BOLA LOUIS IAN A AUROLENSIS LOUISIANA CRUS-CORVI LOUIS IANA HALE1 LOUISIANA IGNOTA LOUISIANA LEAVENWORTHII LOUIS IANA MACROKOLEA LOUI SIANA PICTA LOUISIANA PTYCHOC ARP A LOUISIANA TURGESCENS MAINE *CRI NITA MAINE *INTERIOR MAINE KATAHDINENS IS MAINE *LA XIFLORA MA INE MERRITT-FERNALDI I MAINE ORONENS IS MAINE *PAUPERCULA MA INE PORTER1 MAINE *SCOPARIA MAINE X TRICHINA NUMBER 12 157

COUNTRY STATE TA XON

USA MAINE *TR IS P ERMA MAINE *VE SICARIA MA S S ACHU S ETTS AR GYRAN THA MASSACHUSETTS COLLECTA MASSACHUSETTS G L AUC ODE A MASSACHUSETTS HA LSEYANA MASSACHUSETTS HITCHCOCKIANA MASSACHUSETTS INC@MPERTA MASSACHUSETTS MIRAB ILIS MASSACHUSETTS NOVAE-ANGLIAE MASSACHUS ETTS SCHWEINITZI I MASSACHUSETTS *SCOPARI A MASSACHUSETTS *SCOPARI A MASSA CHU S ETT S SICCATA MAS S A CHU S ETTS TENERA MA S S ACHU SETTS *TENERA MASSACHUSETTS TR ISPERMA MASSACHUSETTS *VESTITA MICHIGAN *ALOPECOIDEA MICHIGAN COOLEY I M ICHIGAN HETEROST AC HYA M ICH IGAN *INTER IOR M ICHIG AN PRA?REA MICHI GAN *RETROCURVA M ICH IGAN *TENUIFLORA MICHIGAN *VULPINUIDEA MINNE SOTA HOUGHTONIANA MISSISSIPPI *STRAMINEA MISSOURI AGGLOMERATA MISSOURI AGGRE GATA MISSOURI EGGERTI I MISSOURI LUNELLIANA MONTANA CONC INNO ID E S MONTANA ELRODI MONTANA HOLM1ANA MONTANA MONTANENSI S MONTANA PL ECTOCARPA MONTANA STANTONENS IS N E BRA SKA *ARI STATA NEBRASKA *DqUGLASI I NEBRASKA LAEVI-CON1 CA NEBRASKA MEEK1I NEBRASKA NEBRASKENSIS NEVADA FISSURICOLA NEVADA HELLERI NEVADA *INTERIOR NEVAOA MICROPTERA NEVADA WATSON1 NEW HAMPSHIRE AENEA 158 SMITHSONIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE TAXON

US A NEW HAMPSHIRE CRAWFORDII NEW HAMPSHIRE *MIRABILIS NEW H AMP SHI RE ORMOST ACHY A NEW HAMPSHIRE *SCOPARI A NEW JERSEY AESTIVALIFORMIS NEW JERSEY BARRATT II NEW JERSEY CAESARIENSIS NEW JERSEY CO NVOL UT A NEW JERSEY CR IST ATA NEW JERSEY CRYPTOLEPI S NEW JERSEY RUGOSPERMA NEW MEXICO AGROSTOIDES NEW MEXICO FENDLERIANA NEW MEXICO GEOPH ILA NEW MEXICO NEOMEXICANA NEW MEXICO PITYOPHILA NEW Y ORK ABDITA NEW YORK A L OPEC 0 IDE A NEW YORK *AMPHIBOLA NEW YORK AQUAT ILIS NEW YORK *AQUATILI S NEW YORK CAREYANA NEW YORK *CEPHALOPHORA NEW YORK COMMUNI S NEW YORK *CRINITA NEW YORK FORMOSA NEW YORK INTERIOR NEW YORK *OLIGOCARPA NEW YORK PLATYPHYLLA NEW YORK RETROCURVA NEW YORK ROSAEOIDES NEW YORK *ROSEA NEW YORK *ROSEA NEW YCRK SARTWELLII NEW YORK SCABR IOR NEW YORK *STERI L IS NEW YORK *ST IPATA NEW YORK *STRI CTA NEW Y ORK SYCHNOCEPHALA NEW YORK *TENTACULAT A NEW YORK *TORTA NEW YORK *VAGI N AT A NEW YORK woo0 I I NEW YORK * X ANTHOC AR P A NEW YORK XEROCARPA NORTH CAROL INA AESTIVALIS NORTH CAROLINA ALATA NORTH CAROLINA BILTMOREANA NORTH CAROLINA BUCKLEY I NUMBER 12 159

COUNTRY STATE T AXON

USA NORTH CAROLINA MISERA NORTH CAROLINA NIGRO-MARGINATA NORTH CAROLINA RUTH1I NORTH CAROLINA *STIPATA NORTH CAROLINA STYLOFLEXA NORTHWEST TERRITORIES FESTIVA OHIO *ALATA OHIO CONJUNCTA OHIO FLACC IDULA OHIO ST EUDEL II OKLAHOMA FISSA OKLAHOMA LATEBRACTEATA OK LAHCMA OKLAHOMENSIS OREGON ACCEDENS OREGON *ACUTA OREGON ACUT INA OREGON ACUTI NELLA OREGON BRACHYPODA OREGON BREVICAULIS OREGON CA MPLYOC AR PA OREGON CUSICKI I OREGON *DEWEYANA OREGON DIVERS1STYLIS OREGON EAST WOOD IAN A OREGON GYMNOCLADA OREGON HALL1ANA OREGON INOPS OREGON *LEPORINA OREGON LEPTOPODA OREGON *MARC I DA OREGON OREGONENSI S OREGON PACHYSTOMA OREGON PANSA OREGON PHAEOCE PHALA OREGON PHAEOLE P IS OREGON SHELDON1I OREGON SPRETA OREGON *STYLOSA OREGON *TERET IUSCULA OREGON VAGANS OREGON VICARIA PENNSYLVANIA *ANGUSTIOR PENNSYLVANIA *AUREA PENNSYLVANI A COSTATA PENNSYLVANIA *DEB1 LIS PENNSYLVANIA GARBER I PENNSYLVANIA GRACILLIMA PENNSYLVANIA *GR ISEA RHODE ISLAND HORMATHODE S 160 SMITHSOXIAN CONTRIBUTIONS TO BOTANY

COUNTRY STATE TAXON

USA SOUTH CAROLINA AUSTRO-CAROLINIANA SOUTH CAROLINA CAROLI NIANA SOUTH DAKOTA HAYOEN II TENNESSEE *OXYLEPIS TENNESSEE PURPURI FERA TENNESSEE ROANENS IS TEXAS BULBOSTY L IS TEXAS MUR ICULATA TEXAS ONUSTA TEXAS RETROFLEXA TEXAS TRIANGULAR1 S TEXAS *WILLDENOVI I TEXAS WRIGHT11 UTAH *CAMPY L OC AR PA UTAH *CANESCENS UTAH EPAPI LLOSA UTAH INTERIMUS UTAH PELOC ARPA UTAH RACHILL IS UTAH *VERNACULA VERMONT rC: INTUMESCEN S VIRGINIA *ABSCONDITA VIRGINIA X ABSCOND IT1FORM1 S VIRGI NIA BAY ARDI VIRGINIA *CR IN1TA VIRGI NI A *CRUS-CORVI VIRGI NIA *DE BIL IS VIRGINIA *DIGITALIS VIRGI NI A *DIGITAL IS VIRGINIA RUGATA WASHINGTON APERTA WASHI NGTON *APERTA WASHINGTON *APERTA WASHINGTON *APERTA WASHINGTON C 0 N ST ANC E ANA WASHI NGTON EGREGIA WASH INGTO N EURYCARPA WASHI NGTON * EURY C AR P A WASHI NGTON *INTERRUPTA WASHINGTON YLENTICULARIS WASHI NGTON MISERABILIS WASHI NGTON *NEBRA SKENS IS WASHI NGTON *NEBRASKENSIS WASHI NGTON NEUROPHORA WASHINGTON OLYMPI CA WASHI NGTON OXYCA RPA WASHI NGTON PADDOENS IS WASHINGTON PRAECEPTORIUM WASHI NGTON SCABRIUSCULA NUMBER 12 161

COUNTRY STATE TA XON

USA WASH INGTON SUBORBICULATA WASHINGTON SUKSDORFII WASHI NGTON *SUKSDORFII WASHI NGTON VIR I01OR WY OM1 NG ALBO-NIGRA WYOMING BR EVISQUAM A WYOMING FE STIVELLA WY OM ING LIMNOPHILA WYOMING NELSON1 I WYOMING PAYSONI S WYOMING PLATYLEPIS WYOMING SIMULATA WYOMING VALLICOLA USSR RUSSIAN SFSR KURILENSIS TADZHIKI STAN PHILOCRENA TURK ESTAN *STENOPHYLLA VENEZUELA BOLIVAR RORAIMENSIS LARA LARENSI S SUCRE CULMENICOLA SUCRE TURUMIQUIRENSIS TACHIRA TACHI RENSI S TACHIRA TAMANA VIET-NAM, NORTH TONKI N TR ICHOP HY LL A WESTEFN SAMOA SAVAI IENSI S 162 SMITHSONIAN CONTRIBUTIONS TO BOTANY

HERBARIUM INDEX

SHEET NO. KIND OF TYPE TAXON

A (ARNOLD ARBORETUM, CAMBRIDGE, MASSACHUSETTS)

ISOTYPE ACROPHI LA ISOTY PE BREVIS ISOTYPE ERE YOST ACHY A I SOTYPE L AMPROCHL AMY S IS OTY P E YEL A NOP HOR A I SOTYPE MERCARENS IS ISOTYPE PER ILEIA ISOTY PE R UBR 0-BRUNN E A VAR ELINEOLATA ISOTY PE SARAWAKETENSIS HDLOTYPE SURCULOSA ISOTYPE TAHITENSIS HOLOTYPE ZIZANI AEFOLIA

CAS (CALIFORNIA ACADEMY OF SCIENCESi SAN FRANCISCO)

242617 ISOTYPE ABORI GI NUM 497554 I SOTYPE ALMA 553879 I SOTY PE ALOPECOIDEA VAR SPARSI-SPI CATA 242962 I SOTY PE APERTA VAR UMBROSA 242960 SYNTYPE APERTA VAR VIRIDANS 242961 SYNTYPE APERTA VAR VIRlDANS 5540 19 ISDTYPE AQUA TIL I S VAR SUBSTRICTA 102030 ISOTYPE ARAPAHOENSIS 43088 1 ISOTYPE BIPARTITA VAR AUSTROMONTANA 383776 TYPE COLLECTION CALIFOSNICA 334353 ISQTY PE C AM PYLOC ARP A SSP AFFINIS 553918 r SOTYPE CHA PMANI 2320 50 ISOTYPE C ILI ARI S 445943 ISOTYPE C ILI ARTS 553874 TYPE FRAGMENT CINNAMOMEA 553913 SYNTYPE COMMUNI S 383554 SYNTYP E C ONJ U NC T A 242987 HOLOTYPE CONSTANCEANA 553883 TYPE COLLECTION CRINlTA VAR MINOR 264973 SYNTYPE CURAT OR IUM 204974 SYNTYPE CURATORIUM 259875 HOLOTY PE DANAENSIS 259874 ISOTY PE DANAENS IS 336835 I SOTYPE DIGITALIS VAR ASYMMETRICA 372834 ISOTYPE DIVERSI STYLI S 351155 I SOTYPE OUR AND1 I 130386 HOLOTYP E E A STWOOD I ANA 2429 57 SYNTYPE EURYCARPA 246772 TYPE COLLECTION E URYCARPA VAR ATTENUATA 40 1490 ISOTYPE E UQ Y STACHY A 234898 ISOTYPE F IS S URICOL A 102307 ISOTYPE FORYOSA NUMBER 12 163

SHEET NO. KIND OF TYPE TA XON

CAS (CALIFORNIA ACADEMY OF SCIENCES, SAN FRANCISCO)

383156 ISOTYPE FORMOSA 103C33 ISOTY PE GRAC ILI OR 353986 I SOTYPE GYNODYNAMA 328017 ISOTYPE HAGIANA 103098 SYNTYPE HOOD II VAR NERVOSA 103098 SYNTYPE HOOD I I VAR NEUROCARPA 553902 TYPE COLLECT ION IGNOTA 404489 ISOTYPE I NCONDITA 2039 10 ISOTYPE lNOPS 348506 I SOTY PE I NT E R I MUS 553999 1 SOTYPE J NTERIOR 272528 ISOTYP E INTERIOR VAR CHARLESTONENSI S 272529 I SOTY PE I NT ER IOR VAR CHARLESTONENSIS 242959 TYPE COLLECTION I NTERRUPTA VAR DISTENTA 477664 I SOTY PE JACOBI-PETER1 384438 ISOTYPE L ATE BRACT EAT A 10 248 1 ISOTYP E LEIOPHYLLA 264341 SYNTYPE LEMANNTANA VAR SIMPLEX 186427 I SOTY PE LE PTOPOOA 136 ISOTYPE LUZ ULAEFOLI A VAR STROBILANTHA 231121 i SOTY PE LUZULAEFOLI A VAR STROBILANTHA 384084 I SOTYPE LUZULINA 264346 I SOTYP E MACKENZI ANA 553885 I SOTYPE MEAD1 I 553875 TYPE FRAGMENT MENDOC INE NS I S 234896 ISOTYPE MICROPTERA 194659 I SOTY PE MURICUL AT A 235733 HOLOTYPE OBI SPOENSIS 237824 IS Of Y PE OBISPOENSIS 237908 ISOTY PE OR I SPOE NS IS 351152 ISOTYPE ONUSTA 239452 ISOTYPE PELOCARPA 369422 I SOTYPE PER COST AT A 155657 ISOTYPE PER LONG A 193005 ISOTYPE P ERL ONGA 383889 ISOTYPE P I TY OPH IL A 416360 ISOTYPE P LEC TOC AR P A 105004 TYPE COLL ECT I ON PRATENSIS 404488 I SOTYPE PRAT IC OL A VAR SUBCORIACEA 102638 ISOTYPE PROPOSI TA 3252 53 ISOTYPE R AC H ILL IS 336836 ISOTYPE RUGATA 3838@1 TYPE COLLECTION SALINAEFORMIS 383407 TYPE COLLECTION SARTWELLII 553975 TYPE COLLECT ION SARTWELLI I 553877 f SOTY PE SCABR IOR 445940 I SOTYPE SEATONIANA 246086 HOLOTY P E SONOMENSI S 161 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TAXON

CAS (CALIFORNIA ACADEMY OF SCIENCES, SAN FRANCISCO)

246636 I SOTY PE SONOMENSIS 232289 SYN TYPE SPECIFICA 342553 HOLOTYPE SPEC UIC OL A 342552 I SOTY PE SPECUICOLA 162423 I SOTYPE SUB1 MPRESSA 259816 HOLOTYPE SURNIGRICANS 152864 I SOTY PE SUKSDORFIX 243333 I SOTYPE SUKSDORF I I 553943 TYPE COLLECT ION T EN1AC UL A T A VAR 4LTIOR 428953 HOLOTYPE TOMPKINSI 429306 I SOTYPE TOMPKI NSI 351161 I SOTY PE TOWNSEND I I 348507 ISOTYPE VER NACULA VAR HOBSONII

DS ( DUDLEY HERBAR IUY, STANFORD UNIVERSITY, STANFORD, CALIFORNIA)

149709 ISOTYPE ABORIGI NUM 55317 I SOTYPE ARRAMSI I 64125 I SOTY PE ABRUPT A 109019 I SOTYPE ARAPAHOENS IS 489410 HOLOTY P E AUSTROYONTANA 49734 TYPE COLLECTION CALIFORNICA 269649 1 SOTYPE C ONSTANC E ANA 14561 9 HOLOTY P E OUDLEY I 629609 I SOTYPE OUDL EY I 144009 I SOTYPE EA STWOOD I ANA 284598 ISOTY PE EGREGEA 149706 I SOTYPE ELRODI 145620 HDLOTY P E GRACILIOR 49500 ISOTYPE GYNO DY NA MA 490408 ISOTYPE GYNOOY NAMA 278190 ISOTYPE INTERIOR VAR CHARLESTONENSI S 75003 HOLOTY PE J AC 1: NTOENS IS 490443 I SOTYPE LEPORI NELLA 13923 I SOT YP E LEPTOPODA 76794 I SOTY PE LUZllL I NA §4832 I SOTYPE MENDOC I NE N S I S 489409 HOLOTY PE YULTICOSTATA 171453 I SOTYPE NESRASKENSIS VAR ERUCAEFORM IS ISOTYPE NUOATA VAR FIRMIOR 270930 ISOTYPE OBISPOENSIS 490462 SYNTYPE PANSA 49738 TYPE COLLECTION PAUCICOSTATA 55002 SYNTYPE QUADR IFIDA 293480 TYPE COLLECTION SAL€NAEFORMIS 490735 I SOTYPE SCABR IUSC UL A 258275 ISOTY PE SONOMENSIS NUMBER 12 165

SHEET NO. KIND OF TYPE TAXON

DS (DUDLEY HERBARIUM9 STANFORD UNIVERSITY, STANFORD9 CALIFORNIA)

374718 I SOTYPE SONOMENSI S 337970 I SOTY P E SPECUI COL A 83850 ISOTY PE STE NOPTE R A 171455 ISOTYPE S UK S DORF I J 269641 f SOTY P E SUKSDORF II 269625 SYNTYPE SUKSDORFII VAR OVALIS 63991 I SOTYPE ULTRA 64032 I S OTY PE ULTRA

F (FIELD MUSEUM OF NATURAl HISTORY 9 CHICAGO, ILLINOIS 1

186491 ISOTYPE ABRAMS I I 206585 TYPE MATERIAL ACUTINA 32699 ISO TYP E ALOPECOIDEA 32700 ISOTYPE ALOPEC0 I D E A 56916 I SOTYPE ALOPECOIDEA 349624 ISOTYP E ALOP ECOIDE A 373673 ISOTY PE ALOPECOIDEA 373679 I SOTYPE ALOPECOI DE A 1464064 TYPE MATERIAL APODOSTACHYA 264169 f SOTYPE AUTUMNAL IS 12661 84 TYPE MATERIAL AZUAYAE 999642 TYPE MATERIAL BARTLETTI I 314869 ISOTYPE BONPLANDI I VAR MINOR 456934 ISOTYPE BONPL4NDIl VAR MINOR 1620435 IS OTY PE CHIAPENSIS 49642 I SOTYPE CHIHUAHUAENS IS 2020 21 ISUTYP E C ILIARIS 26304 TYPE MATERIAL CREBRIFLORA 1 I28952 HOLOTYPE CUCHUMATANENSI S 1266170 HO L D TYP E CULMENI COLA 1406416 TYPE YATERI AL CUNEATA 751055 ISOTYPE X DEAMII 455703 TYPE COLLECT ION DEW EYANA VAR SPARSIFLORA 1429766 TYPE COLLECTION DEHEYANA VAR SPARSIFLORA 122779 TYPE MATERIAL FESTlVA VAR DECUMBENS 314892 SY NTY PE HALL 1 I 456958 SYNTYPE HALL II 176870 ISOTYPE HARPER1 141 1493 TYPE MATERIAL HATUSIMANA 2831 19 TYPE MATERIAL HELLERI 1128957 HOLOTYPE HUE HUE TEC A 1463659 TYPE MATERIAL HYYENODON 314869 I SOTY PE ILLOTA 456934 f SCITY PE ILLOTA 1076930 ISOTY PE INT ER I OR VAR CHARLESTONENSI S 263394 TYPE COLLECTION INVOLUCRATELLA 166 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TA XON

F (FIELD MUSEUM OF NATURAL HISTORY, CHICAGO, ILLINOIS)

1406403 TYPE MATERIAL KURILENSI S 129242 SYNTYPE L ACU NARUM 1566419 SYNTYPE LAC UNARUM 1481645 ISOTYPE L ANGEANA 55470 HOLOTY PE L ARENS I S 267758 I SQTYPE LA XI FLORA VAR LEPTONERVI A 455706 TYPE COLL ECT ION L EPORINA VAR AMERICANA 1425899 TYPE COCLECTiON LEPORINA VAR AMERICANA 206587 TYPE COLLECTION MARC IDA VAR OEBILIS 866418 I SOTY PE OBISPOENSIS 455736 S YN TYPE OREGONENS IS 96129 TYPE YATER IAL OXYCARPA 211365 TYPE YATERIAL OXY CARPA 223512 TYPE Y4TERIAL PADOOENSI S 1471 48 9 TYPE Y4TERIAL P ADDOENS I S 309086 TYPE YATERIAL PAUC I COST AT A 178542 I SOTYPE PER LONG A 455706 TYPE COLLECT ION PHAEOCEPHALA 1425899 TYPE COLLECTION P HA E OC E PHAL A 755322 TYPE MATERIAL P HY SOC HL A E N A 1G5551 TYPE COLLECT ION PR I NGL E I 1607711 TYPE COLLECTION PRI NGLE I 215918 TYPE COLLECTION PSEUDOJAPONIC A 1263854 HOLOTYPE RORA IMENS IS 1489429 ISOTYPE RUGATA 1463953 TYPE MATERIAL RUGATA 30885 TYPE M4TERIAL S AL INAE FORM IS 210109 TYPE COLLECTION SCI RPOIDE A VAR GIGAS 464432 SY NTY PE SCOPAR IA VAR SUBTURBI NATA 907841 I SOTYPE SONOMENSIS 1252385 HOL OTYP E S TA NDLE YANA 1129096 HOLOTYPE STEY ERYARK II 813737 TYPE COLLECTION STI PATA V AR L A EV IVAG INAT A 1263 85 8 HO L 0 TY P E TACHIRENSI S 1263857 HOLOTYPE TAYANA 1129085 TYPE MATERIAL TOJQUIANENSIS 1129086 TYPE MATERIAL TOJQUIANENSIS 1266183 TYPE MATERIAL TORE ADORA 1128966 TYPE TUN1 MANENSI S 12661 50 HOLOTY PE TURUMIQUIRENS IS 1129094 HOLOTY9 E V E NOS I V AG IN AT A 89120 TYPE YATERIAL VITREA 309085 SY NTY P E WHITNEY I 309086 SYNTYPE WHITNEY I NUMBER 12 167

SHEET. NO. KIND OF TYPE TAXON

GH (GRAY HERBARIUM, HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS)

ISON EOTY P E ABLATA I SOTY PE A B SC ON0 ITA VAR ROSTECLATA HOLOTYPE X ABSC ONDI T IFORM IS ISOTY PE X A BSCONO IT IFOR M IS SY NTY PE ACCEDENS TYPE COLLECTION AC UTA VAR PALLIDA ISOTYPE ACUTINA SYNTYPE ADUSTA VAR GLOMERATA SYNTYPE AENEA TYPE CQLL ECT ION AEQlJA ISOTYPE AESTIVACIFORMIS HOLOTYPE ALATA VAR FERRUGINEA I SOTYPE AL OPEC 0 IDE A I SOTYPE AL OPEC 0 I DE A VAR SPARS I-SPI CATA HOLOTYP E AMPHIBOLA VAR TURGIDA TYPE ANGUSTIOR VAR GRAC ILENTA SYNTYPE APERTA ISOTYPE AQUAT IL IS VAR SUB STR ICTA ISOTY PE ARAPAHOENS IS HDLOTYPE ARG YR AN THA ISOTYPE ARGYRANTHA HOLQTYPE ARI STATA V A R LONGO-L ANCEOL ATA I SOTYPE ARSENII HOLOTY PE ART ITECTA VAR SUBTILIROSTRIS SYNTYPE ASSINIROINENSIS TYPE COLLECTION AZTECICA TYPE COLLECT ION BACKANA SYNTYPE BACK1 I HOLOTYP E BARBARA€ ISOTYPE BARTLETT II HOLOTYPE BAYARDI ISOTYPE B ILTMOREANA HOLOTY P E BONPLANDI I VAR MINOR I SOTYPE SREWERI ISOTY PE B UR C HELL I AN A ISOTYPE CAESARIENS IS PARATYPE CANESCENS VAR DISJUNCTA ISOTY PE CANE SCEN S VAR SPHAEROSTACHYA TYPE COLLECTION CAROL INI AN A TYPE CEPHALOPMORA VAR MAXIMA SY NTY PE CHALC IOLEP IS HOLOTY PE C ILI ARIS TYPE COLLECTION CI NNAMOYEA ISOTYPE C IRC INNATA HOLOT Y PE CLIVICOLA HOLOTYPE COLLECTA SY NTY PE CONJUNCTA HOLOTYPE C OOCEY I 168 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET Nn. KIND OF TYPE TA XON

GH (GRAY HERBARIUM, HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS)

SYNTYPE CRA WFORD I I SYNTYPE CRAWFORDX I VAR VIGENS TYPE C REBRIFLORA HOLQTYP E CRINITA VAR BREVICRINIS SYNTYPE CRINITA VAR SIMULANS I SOTYPE C Rf STATA ISOTYDE CRUS-CORVI YOLOTY PE C RUS-CO RV I VAR VIRGINIANA HOLOTYPE C UYULATA FOR SOLUTA I SOT Y PE DANAENS IS I SOTYPE DAVY I HOLOTYPE DERILIS VAR INTERCURSA HOLOTY P E DEB ItIS VAR PUBERA ISOTYPE DEVIA HOLDT Y P E DEUEYANA VAR COLLECTANEA TYPE COLLECTION DEWEY ANA VAR SPARSIFLORA HOLOTYPE DIGITALIS VAR ASYMMETRICA HOL OTY P E DIGITAL IS VAR MACROPODA TYPE YATERI AL DOUGLAS11 VAR DENSISPICATA I SOTYPE X DUMANII ISOTYPE DUT ILLY I I SOTYPE EASTWOODI ANA HOLOT Y P € ECHINATA VAR ORMANTHA I SOTY P E EKMANI I I SOTY PE EKYANI I VAR HOTTENSIS TYPE Y4TERIAL EL9ERTANA I SOTY PE ELEOCHARIS I SOTYPE ELYNOI DES HOLOTYPE ENGE LMANNI TYPE YATE RIAL ERXLEBENI ANA I SOTYPE EURY STACHYA HOLOTY PE E XP L OR ATORUM TYPE COLLECTION FEL I PENS IS ISDTYPE FESTIVELLA HDLOTY PE FETA I SOTYPE FILIFOLIA VAR EROSTRATA HOLCITYPE F ISSA VAR ARISTATA HOLOTY PE FLACCOSPERMA HOLOTYPE FLAVA VAR GASPENSIS ISOTYPE FLAVA VAR RECTIROSTRA ISOTY PE FOENEA VAR PERPLEXA I SOTYPE FORMOSA ISOTYP E FRANKLIN11 I SOTYPE GARBERI HOLOTYPE GARBERI VAR BIFARIA HOLOTY PE GLAREOSA VAR AYPHIGENA I SOTY PE GLAUCODEA I SOTYPE GR4CILIOR NUMBER 12 169

SHEET. NO. KIND OF TYPE TA XON

GH (GRAY HERBARIUYv HARVARD UNIVERSITY, CAMBRIDGE, MASSACHUSETTS)

I SOTY PE GRACILLIMA HOLOTY PE GRI SEA VAR RIGIDA HOLOTYPE GUATEMALENSI S TYPE COLL ECT ION HALL IANA SYNTYPE HALL II HOLOTYPE HALSEYANA YO LOTYP E HARPER I SYNTYPE HAY DENI AN A HOLOTYP E HERE TATA SYNTYPE HEPBURN II TYPE YATERI AL H EfE RON EUR A HOLOTYP E HITCHCOCKI ANA TYPE COLLECT ION HOOK ERANA SYNTYPE H OR MATHODES HOLOTY PE HORNSCHUCHI ANA VAR LAURENTI ANA HOLOTYPE HOST IANA VAR LAURENTIANA HOLOTY PE TLLOTA ISOTYPE I NC I SO-DE NTATA ISOTY PE I NCURV IFORM IS HOLOTY P E INFLATA VAR ANT ICOSTENS IS YOLOTY P E INOPS I SOTY PE INTERIMUS I SOTYPE INTERIOR VAR CHARLESTONENS IS HOLOTY PE INTERIOR VAR JOSSELYNI I HOLOTYPE INTERIOR VAR KEWEENAWENSIS HOLOTYPE I NTUME SCENS FOR VENTRIOSA I S OTY PE INVOLUCRATELLA I SOTYDE KALOIDES HOLOTYPE KATAHDINENSIS SYNTYPE KELLOGGI I ISOTYPE KOKRINENSIS HOLOt Y P E LAEVI-CONICA HOLOTYPE LANGEANA HOLOTVPE LASIOCARPA VAR AMERICANA ISOTYPE I- ATEBRACTEATA HOtOTYPE LAX IFLORA VAR LEPTONERVIA HOLOTYPE LAX1FLORA VAR SERRULATA ISOTYPE L E IOCARPA ISOTYPE LEIOPHY LLA TYPE COLLECTION LEMHONI TYPE COLLECTION L EPOSINA VAR AMER ICANA ISOTYPE LEPORINELLA HOLOTYP E LIVIDA VAR RUFINAEFORMIS ISOTYPE LONG ICULM IS TYPE COLLECTION LUZULINA HOLOTYPE MACKENZIANA HOLOTY PE MEAOX I TYPE CDLLECTION MEEK1I 170 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TA XON

GH (GRAY HERBARIUM, HARVARD UNIVERSITYI CAMBRIDGE, MASSACHUSETTS)

I SOTYPE MELOZITNENSIS H0LC)TYPE YENDOCINENSIS HOLOTYPE MERR ITT-FERNALDI I ISOTY P E ME S OCHOR E A I SOTYPF M I C ROC H AE 1A HOLOTYPE MIL1 ARI S VAR AUREA HOLOTY P E MIRABILIS SY NTY PE MIRABIL IS VAR PERLONGA SYNTYPE MIRABILIS VAR TINCTA HOLOTY PE M IS ANDRO IDES HOLOTYP E MONTEREYENSIS I SOTYP E NEL SON1 I ISOTYPE X NEOBIGELOW I I HOLO TY PE X NEOFILIPENDULA I SOTYP E X NEOPALEACEA HOL OTYP E NERV INA ISOTYPE NEUROPHORA I SOTYP E NIGRICANS ISOTYPE NOVAE-ANGL I AE I SOTYPE X NUBENS ISOTYPE NU5 ICOLA H OLOTY PE NUTTALL I I I SOTYPE OAXACANA ISOTYPE OBISPOENSIS I SOTYPE OBOVOI DEA HOLOTY P E OL I G UC A R P A VAR LATIFOLIA HOLOTY PE ORMOST ACHY A HOLOTYP E ORONENSI S ISOTYPE P AC H Y ST AC HYA !SOTYPE PADDOENSIS I SOTYPE PALAWANENSI S HOLOTY PE PALL ESCENS VAR NEOGAEA HOLOTYPE PAPULOSA ISDTYPE X PATUENSIS HOLOTY PE P AUP ERCULA VAR BREVISQUAMA YOLOTY PE P AUP ERCUL A VAR PALLENS I SOTYP E PAYSONI S HOLOTY PE P ER L ONGA I SOTYPE PERSTRI CTA I SOTYPE PETRIE I TYPE COLLECTION PHA EOC E PHAL A HOLOTYPE PHY I, LUM AN I C A ISOTYPE P I NE TOR UM VAR ELATIOR I SOTYPE P IPERI TYPE MATERIAL PLATYPHYLLA TYPE VAT ER IAL P 0 DOCAR PA HOLOT Y PE PORTER1 I SOTYP E POTOSINA NUMBER 12 17 1

SHEET NO. KIND OF TYPE TAXON

GH (GRAY HERBARIUM9 HARVARD UNIVERSITY, CAMBRIDGE9 MASSACHUSETTS)

HOLOTY PE PRAEGRACIL IS H OLOT Y QE PRA IREA ISOTYPE PRATENSI S VAR FURVA SY NTY PE PRE ISS I I TYPE COLLECTION BKI NGLEI ISOTYPE PROJECTA T SOTY PE PROPOSITA HOLO TYP E X PSEUDO-FULVA ISOTY P E X QUEBECENSIS SYNTYPE RAYNOLDSI I HOLOTYPE R E T R OC UR VA ISOTYPE R ETROCURVA VAR COPULATA TYPE YATERI AL R ICHARDSON II HOLOTYP E RICHARDSON II FOR EXSERTA HOLOTYP E ROS EA VAR ARKANSANA I SOTYPE RURRO-BRUNNEA VAR ELINEOLATA HCILOTYPE RUGATA HOLOT Y P E S ARTW ELL I ANA ISOTYPE X SAXENII NM. FERRUGINEA ISOTYPE SCHAFFNERI HOLOTY P E S C H N E I 0 E R I HOLOTYPE SCHWEINITZII HOLOTY P E SCOP AR 1 A VAR CONDENSA SY NTY PE SCOPARl A VAR FULVA HOLOTVPE SCOPARI A VAR MONILIFORMIS HOLOTY PE SCOPAR IA FOR PERACUTA HOLOTYPE SCOP AR I A V A R SU BTUR BINAT A HOLOTYP E SC OPARI A VAR TESSELLA A ISOTYPE S EATONIANA ISOTYPE SONOMENSIS HOL OTY P E SOPERI ISOTYPE SPECUICOLA TYPE COLLECTION SPRETA TYPE COLLECT ION STIPATA VAR LAEVIVAG NATA SYNTYPE STRAMINEA VAR CUMULATA SYNTY PE STRAMINEA VAR ECHINODES SYNTY PE STYLOSA VAR VIRENS ISOTY PE S UR -BR ACT EAT A ISOTYPE SUB FUSC A ISOTYPE SUK SDORF II ISOTYPE SYCHNOC EPHALA YOLOTYPE TENERA HOLOTY PE TENERA VAR RICH11 I SOTYPE T ENU IFL OR A VAR SETACEA HOLD TYP E TERRAE-NOVAE HOLOTYPE T HU RBE R I HOLOTYPE X TRICHINA HOLD TY P E TRICHOPHYLLA 172 SMITHSOKIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TAXON

GH (GRAY HERBARIUM, HARVARO UNIVERSITY, CAMBRIDGE, MASSACHUSETTS)

I SOTY PE TRIS PERM A VAR B ILL INGS II 1 SOTY PE TSOI ISOTY PE TURUYIQUIRENSIS HOLOTYPE UMBELLATA VAR VICINA ISOTYPE UNCOMPAHGRE HOLOTY PE VAG INATA VAR ALTO-CAUL1 S HOLOTYPE VALL ICOLA ISOTYPE VENOSIVAGINATA ISOTY PE V ERN ACUL A VAR HOBSONII HOLO TYP E VESICARIA VAR JEJUNA HOLOTYP E VES I CAR IA VAR LAURENTIANA HOLOTY PE VESTITA VAR KENNEDY1 H0LI)TY PE VICARIA ISOTYPE VIRIDULA FOR PYGMAEA HOLOTYPE VULP INOIDEA VAR PYCNOCEPHALA HOLOTYPE WA T SON1 ISOTYPE W I EGBND I I TYPE COLLECTION WILLDENOVI I V AR PAUC I FLORA HOL0 TY P E WOOD I1 HOLOTY PE X XANTHINA SYNTYPE X E RANTI C A

JEPS (JEPSON HERBAP IUMt UNIVERSITY OF CALIFORNIA, BERKELEY)

2511 ISOTYPE DAVY I 4013 ISOTYPE INTERIOR VAR CHARLE STONENSI S 20008 I SOTY PE JEPSONI I 19722 I SOTYPE MAR1 POSANA

MO (MISSOURI BOTANICAL GARDEN9 ST. LOUIS)

TYPE COLLECTION ABSCOND I TA VAR ROSTELLATA 130648G ISOTY PE X ABSCOND ITI FORM IS SYNTYPE ACCEDENS TYPE MATERIAL AESTIVALIFORMIS TYPE VAT ER I AL AESTIVALIS TYPE COLLECT1 ON AGGLOMERATA TYPE COLLECTION AQUATILI S VAR SUBSTRICTA SYNTYPE ATHR OST ACHY A 1834152 HOLOTYPE AUSTRO-C AROL I N I ANA ISOTYPE A UTVMN4L I S TYPE COLL ECT I ON AZT ECICA 1611724 SYNTYPE BANKS1 I ISOTYPE BONPLANDI I VAR MINOR 710112 TYPE COLLECTION BUL BOSTY L IS TYPE COLLECTION CALI FORNICA NUMBER 12 173

SHEET NO. KIND OF TYPE TAXON

YO (YISSOURI BOTANICAL GARDEN, ST. LOUIS 1

SYNTYPE CHALCIOLEPIS ISOTYPE C ILI AR I S 1108572 I SOTYPE C R U S-C ORV I VAR VIRGINIANA TYPE MATERIAL DE6 ILIFORM IS 1306423 I SOTYPE 0 I G ITALI S VAR ASYMMETRICA 1129747 r SOTYPE 0 IG I TAL I S VAR MACROPODA I SOTY P E DUDLEY I TYPE YATERI AL ELYERI TYPE COLLECTION ECYNOIDE S I SOTY PE EPAP ILLOSA 181 6497 TYPE COLLECTION FENDLERIANA I SOTYPE F ISSA I SOTY P E GRAC ILIOR TYPE MATERIAL GY MNOCLA DA SYNTYPE HALL II TYPE YATERIAL HAY DEN I I I SOTYPE ILLOTA 1816496 TYPE COLLECTION INTERIOR 114838 1 I SOTY P E INTERI OR VAR CHARLESTONENSI S 1190731 ISOTYPE INTERIOR VAR CHARLESTONENSIS 1201697 ISOTYPE INTERIOR VAR CHARLESTONENSIS TYPE MATERIAL I NV OLUC RAT ELLA SYNTYPE LACUN4RUM 1692 174 I SOTY P E LATEBRACTEATA I SOTY PE LAYIFLORA VAR CEPTONERVI A TYPE COLLECTION LEPORINA VAR AMERICANA TYPE YATERIAL LUZUL INA I SOTY PE MEAD I I I SOTYPE MENDOCI NENSIS 952735 TYPE COLLECT ION N EBRA SK ENS I S VAR ERUCAEFORMI S TYPE YATERI AL NUDATA TYPE COLLECTION 0 A XACA N A 1220830 TYPE WATERIAL 0 BOVO IDE A TYPE MATERI AC OKLAHOMENSIS TYPE MATERIAL P A UC I CO STAT A TYPE MATERIAL PER GLOBOSA I SOTYPE PERLONGA TYPE COLLECTION PHA E OC E PH 4LA ISOTYPE P HY L LOMAN IC A 9208 15 TYPE COLLECTION PHY S OCHL A E N A TYPE MATERIAL PRAINI I 2002968 SYNTYPE PRE ISSI I TYPE COLLECTION PRI NGLEI 1260436 I SOTYPE RUB RO-B RUNNE A VAR ELINEOLATA 1306478 I SOTY PE RUGATA TYPE MATERIAL SALI NAEFORMIS TYPE YATERIAL SAX1MONTANA I SOTY PE SCABRIUSCULA 174 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TAXON

MO (MISSOURI BOTANICAL GARDEN, ST. LOUIS)

2002967 TYPE COLLECTION SCAPOSA TYPE COLLECTION SPR ETA TYPE COLLECTION STIPATA VAR LAEVIVAGINATA SY MT Y P E STR AM I NE A VAR CUMULATA TYPE COLLECT ION STYLOFL EXA SYNTYPE STYLOSA VAR VIRENS 95212 TYPE COLLECTION SUK SDORF I I TYPE COLLECT ION VIOL ACEA I SOTYPE VITREA SYNTYPE WH I TNEY I SYNTY P E WHITNEY I SY NT Y PE WHITNEYI

NY (NEW YORK BOTANICAL GARDEN, NEW YORK CITY)

TYPE ABDI TA IS ONE OTY P E ABLATA I SOTY PE ABORI GI NUM HOL OT Y P E ARRAMSI I HOLOTY PE ABRUPTA TYPE COLLECTION A 6SC OND ITA VAR ROSTELLATA ISOTYPE X ARSCONDITIFORMIS TYPE COLLECT ION AClJT INA TYPE COLLECTION AEQUA TYPE YATER IAL AESTIVALIS TYPE A GGREG AT A SYNTYPE ALATA TYPE COLLECTION ALO I DA ISOTYPE ALYA I SOTYPE ALOPECOIDEA I SOTYP E ALOPECOIDEA VAR SPARSI-SPI CATA TYPE COLLECTION AMP H 100 L A TYPE COLLECTION ANGUSTI OR I SOTYPE APODA TYPE COLLECT ION AQUATIL IS I SOTYPE ARAPAHOENS IS TYPE ARCTAEFORYI S TYPE COLLECTION ARC1 ICA I SOTYPE ARSENII SYNTYPE ATHROSTACHYA SYNTYPE ATHROST ACHY A ISOTYPE A TQ AC TODE S IS OTY PE A TP. 0 S Q U A M A I SOTY PE AUREA VAR ANDROGYNA TYPE COLLECTION AUROLENSI S ISOTYPE AZUAYAE SYNTYPE BAC KII NUMBER 12 175

SHEET NO. KIND OF TYPE TA XON

NY (NEW YORK BOTANICAL GARDEN, NEW YORK CITY)

COTY P E BALTZELLII I SOTY PE BARBARAE TYPE COLLECTION BARRATTI I TYPE BARTLETTI1 TYPE B ILTMOREANA TYPE BOYANZENSI S TYPE COLLECT ION RRE V I CAUL IS HOCOTY P E RREVISQUAMA SYNTY PE BRONGNI ART1 I VAR DENSA TYPE COLL ECT ION BUCKL EY I TYPE COLLECTION BUSHI I TYPE CALSARIENSI S TYPE COLLECTION CAL I FORN ICA HOLOTYPE CAMPY LOCARP A SSP AFFINIS ISOTYPE CANE SCENS VAR DUBIA TYPE COLLECT ION CANESCENS WAR SPHAEROSTACHYA HOLO TYPE CAREYANA TYPE COLLECTION CAROL? NIANA SYNTYPE CHALC IOLEP IS I SOTYPE CHI APENSIS HOLOT YP E CHIHUAHUAENSI S TYPE CHIKUNGANA I SOTYPE CILIARIS HOLOTYPE COLUMBI ANA TYPE CONC INNOIDES ISOTYPE CONSTANCE ANA TYPE CONVOLUTA TYPE COLLECTION COSTATA TYPE COLLECTION CRANDALLI I TYPE CR EBR I FLORA I SOTY PE CRUS-CORVI VAR VIRGINIANA TYPE CRYPTOLEPIS I SOTY PE CURENS IS TYPE COLLECTION CU6 ENS IS VAR FLACCIDA SYNTYPE CUSICKII ISOTYPE DAVY I TYPE COLLECTION DEWEYANA V AR SPARS I FLORA TYPE COLLECT ION DIGITALI S VAR GLAUCA ISOTYPE D IGIT AL IS VAR MACROPODA ISOTYPE DUDLEY I SYNTYPE DUR I FOL I A TYPE EBENEA TYPE COLLECTION EGGERTI I I SOTY P E EGGLESTONII TYPE EGREGTA TYPE MATERIAL EKMANI I ISDTYPE ELRODl HOLOTY PE €PAP ILLOSA 176 SMITHSONIAN CONTRIBUTIONS TO BOTANY

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ISOTYPE EPAPI LLOSA I SOTY PE EXPLORATORUM TYPE MATERIAL FARGESII TYPE COLLECTION FELIPENS1 S TYPE FENDLER IANA SYNTYPE f EST IVA TYPE COLLECTION FESTIVA VAR DECUMBENS TYPE YATERIAL FESTIVA VAR STRICTA TYPE FESTIVELLA HOLOTYPE F TSSA I SOTY PE F ISSA VAR ARISTATA TYPE FLACCI DULA ISOTYPE F LACC 0s PE RMA TYPE f RANKLINI I TYPE FULVE SC ENS TYPE YATER IAL FUSCOLUTEA TYPE FUSC Of INCT A TYPE GAYANA VAR HYALINA TYPE COLL ECT ION GEYERI ISOTY PE GLAREOS A VAR AYPHIGENA TYPE COLLECTION GRIFFITHII 1S OTY PE GY NOC L AD A ISOTYPE GY N ODY N AM A SYNTYPE HALE I SYNTYPE HALLII SYNTYPE HALL II ISOTYPE HALSEYANA HO LOT Y P E H AR FORD I I ISOTY PE HARPER1 TYPE COLLECTION HASSEI TYPE HELLERI TYPE HETEROSTACHYA ISOTYPE HINDS11 VAR BREVIGLUMA TYPE COLLECT ION HIR SUT A VAR CUSPIDATA TYPE HOL M IAN A SYNTYPE HOOD I I VAR NERVOSA SYMTYPE HOOD I I VAR NEUROCARPA TYPE COLLEC TION H OOKERANA ISOTYPE HORNSCHUCHI ANA VAR LAURENTIANA HOLOTY PE HOUGHTONI ANA TYPE COLLECTION IGNOTA TYPE COLLECTION ILLINOENS IS TYPE INCOMPERT A HOLOTYPE INCURVIFORMI S I S OTY PE INOPS HOL0 TYPE INT EGRA TYPE I NTERIM US ISOTYPE INTER IOR NUMBER 12 177

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NY (NEW YORK BOTANICAL GARDEN, NEW YORK CITY)

ISOTYPE INTERI OR VAR CHARL ESTON ENS IS I SOTYP E INTERIOR VAR JOSSELYNI I ISOTY PE I NT E R IOR VAR KEWEENAWENSI S TYPE COLLECTION I NVOLUCRATELLA YO LO TYP E JAYESII HOCOTY PE JEPSONI I SYNTYPE JON€ SI I I s OTY PE K ATAHDINENSIS TYPE KUL I NGANA TYPE LAC INI A TA SYNTY P E L ACUNARUM I SOTYPE LANC I FRUCTUS TYPE LARICI NA ISOTY Pf LAX I FLORA VAR LEPTONERVI A I SOTYPE LAX1FLORA VAR SERRULATA TYPE COLLECTION LEAVENWORTHI I I SDTY PE LENT ICULARI S VAR PAULL IFRUCTUS TYPE COLLECTION L E PORT N A VAR AMERICANA TYPE LEP TOPODA TYPE COLLECTION L ONGI ROSTRIS VAR MICROCYSTIS TYPE LUNELLI ANA TYPE COLLECT ION LUZULIYA COTYPE MAC ROKOL E A TYPE MAC ROSPERMA ISOTYPE MADRENSIS I SOTYPE MANDONI ANA TYPE MARC I DA VAR DEBILIS HOLOTYPE MAR I POSANA ISOTYPE MEADII. TYPE WED1 TERRANI A ISOTYPE MENDOC INENSIS TYPE YATERI AL MERRILLII TYPE M 1C ROPTE R A ISOTYPE MISANOROIDES ISOTYPE MISERA HOLOTYP E MOHR IANA HOLOTYPE MOLESTA SYNTY P E MONTANENSIS TYPE MATERIAL NANA IS ClTY P E NEBRASKENSIS ISOTYPE NEBRASKENS IS VAR ERUCAEFORM IS I SOTYPE NEBRASKENSI S VAR ULTRIFORMIS HOLOTYPE NELSON1 I I SOTYPE X NEOBIGELOWI I TYPE COLLECTION NEOYEXICANA I SOTY PE NEUROPHOR A TYPE COLLECTION NIGRO-MARGINATA 1 SOTY PE NUTTALL I I 178 SMITHSONIAN CONTRIBUTIONS TO BOTANY

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I SOTYPE OBI SPOENS IS TYPE OEDERI VAR ROUSSEAUIANA TYPE OKLAHOMENSIS SYNTYPE OLY MPICA TYPE ONUSTA SYNTYPE OREGONENS IS ISOTYPE ORONENS IS TYPE COLLECTION OXYL EP I S VAR PUBESCENS I SOTYPE PADDOENSIS TYPE MATERIAL PALA WANENS IS SYNTYPE PANSA I S OTY P E PAPULOSA TYPE COLLECTION PARRYANA TYPE COLL ECT ION P AUC ICOSTATA HOLOTY PE PEL OC AR P A HOLOTYPE PERGLOBOSA I S OT Y PE PERLONGA TYPE PER STR 1 CT A TYPE PE TA SATA TYPE PETR ICOSA TYPE COLLECTION PHA EOC E PHA L A I SOTYP E PHAEOCEPI S TYPE MATERIAL PHILOCRENA ISOTYPE PHYLLOMAN! CA TYPE COLLECTION PHY SOCHLAENA TYPE COLLECTION PICTA ISOTYPE PINETORUM VAR ELATIOR TYPE MATERIAL P ITYOPHILA HOLOTYPE PLA TYLE P I S HOLOTYPE PRAECE PTOR I UM TYPE C3LL ECT ION PR I NGLE I TYPE COLLECTION PRIONPHY LLA TYPE COLLECTION PROJEC TA HOLOTY PE PROPOS ITA TYPE COLLECTION PSEUDOJ APON I C A TYPE COLLECTION PTYCHOC ARP A TYPE COLLECT ION P UR P UR I FERA SYNTYPE QUADRIFIDA SYNTYPE QUADRI F IDA VAR LENIS I SOTY PE X QUEBECENSIS TYPE RACHI LL IS TYPE RETROFLEXA COTYPE RHYNCHACHAENIUM I SOTYPE RORAIMENSIS ISOTYPE R OS AEO I OES T SOTYPE ROSEA VAR ARKANSANA TYPE ROS EA VAR PUSILLA TYPE ROSEA VAR STAMINATA NUMBER 12 179

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NY (NEW YORK BOTANICAL G4RDENv NEY YORK CITY)

TYPE RUB RO-B RUNNEA VAR ELINEOLATA I SDTY PE RUGATA TYPE RUGOSPERMA TYPE RUSBYI HOLOTY 0 E RUTHI I TYPE COLLECTION SALI NAEFORMIS TYPF COLLECTION SANGUI NEA TYPE COLLECTION SARTWELLII I SOTYPE SAVAIIENSIS TYPE MATERIAL SAXIYONTANA TYPE SCARRI USCUL A TYPE SCIRPIFORMIS TYPE COLLECTION SCIRPOIDEA VAR GIGAS TYPE COLLECTION SC I RPO I DE A VAR STENOCHLAENA TYPE COLLECTJON SCfIPARI A VAR MINOR I SOTY PE SCOPAR I A VAR SUBTURBINATA I: SOTYPE SCOPARI A VAR TESSELLATA HOLOTYPE SC OULERI ISOTY PE SHELDONI I TYPE SHELDONI I TYPE COLLECT ION S ICCATA HOLOTY P E SIMULAT9 1 SOTYPE SONOMENSI S ISOTY PE SPEC U ICOLA TYPE STANTONENS IS I SOTYPE STE L LATA fY PE STELLULATA VAR CONFERTA TYPE STENOPTER A ISOTYPE STERILI S VAR EXCELSIOR TYPE COLLECT ION STEUDEL I I TYPE COLLECTION STI PATA VAR LAEV IVAGINATA TYPE COLLECTION STIPATA VAR SUBSECUTA ISOTYPE STIPATA VAR UBERIOR SYNTYPE STQAMINEA VAS CUMULATA SYNTYPE STRAMINEA VAR RENIFORMIS TYPE COLLECTION STRICTA VAR XEROCARPA TYPE COLLECTION STYL OFL EX A HDLOTYPE SUS-RRACTEATA TYPE SURORBI CULATA TYPE Y4TERIAL SUSTPANSVERSA ISOTYPE SUK SDORF 1 I SY NTY PE SUKSDORF II VAR OVALIS HDLOTYPE T4MAKI I HOLOTY PE T ENERAE FORM IS SYNTYPE TERETIUSCULA VAR AMPLA ISOTYPF TOMPKI NSI ISOTYPE TOR EADORA TYPE COLLECTION TORTA VAR STAMINATA 180 SMITHSONIAN CONTRIBUTIONS TO BOTANY

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ISOTYPE TOWNSENDI I TYPE TRACY1 I SOTYPE TRI ANGULAR1 S ISOTYPE TR I SPERYA TYPE TSOI HOLOTYP E TUMULICOLA TY PF TURGESCENS I SOTY PE TURUMIQUIRENS IS TYPE UNDERWOOD1 I TYPE UN I L AT ER At I S I SOTY PE VAGANS HOLOTYPE VEQ N AC UL A VAR HOBSONII TYPE COLLECT ION WERDERMANNI I SYNTYPE WHI TNEY I SYNTYPE WHITNEYT SYNfYPE WHITNEY I TYPE COLLECTION WILKESI I TYPE WILLIAYSII TYPE COLLECTION WRIGHT1 I TYPE X A N THOC A R P A VAR ANNECTANS SYNTYPE XERANT ICA TYPE COLLECTION XEROCARPA TYPE YUYONENSI S uc (UNIVERSITY OF CALIFORNIA, BERKELEY)

9@5439 HOLDTYPF A PODA 905436 HOLOT Y PE AR AP AH0 ENS I S 50814 HOLOTYPE DAVY I 9 100 20 HOLOTYPE INTERIOR VAR CHARLESTONENS IS 127723 HOLOTY PE LANC IFRUCTUS 55234 HOLRTYPE LEPORINELLA 1098 ISOTYPE MENDQC I NENS I S 319673 HOLOTY PE OBL ANCE OL AT A 10 60 HOLO TYPE PACH YC A RP A 90 5434 HOLOTYPE PAY SON1 S 835699 I SOTY PE SONOMENS I S 9c5433 HOLOTYPE SUBIMPRESSA 90 544 1 HOCOTY PE TRIBULO IDES VAR SANGAMONENSI S us (U. S. NATIONAL HERBARIUM, SYITHSONIAN INSTITUTIONV WASHINGTON) 2003161 TYPE COLLECTION ABSCQNDITA VAR ROSTELLATA 2003299 I SOTYPE X ABSCONDITIFORMIS 25 I64 TYP’E COLLECT ION ACUTXNA 817087 TYPE COLLECTION ACUf INA NUMBER 12 181

SHEET NO0 KIND OF TYPE. TAXON us (U. So NATIONAL HERBARIUM, SYITHSONIAN INSTITUTIONt WASHINGTON)

27286 HOLOTYPE ACUTI NELLA 4401 79 TYPE COLLECTION A GGL OME RA T A 617798 TYPE AGROSTOIDES 694342 TYPE AGRO STOI DE S 858947 TYPE COLLECT ION AL 60-M I GR A 2231424 TYPE AYPLISQUAMA 865058 TYPE ANTH OXANTHERA 1G30011 TYPE COLL ECT ION ARSENII 2265958 HOLOTY PE ATHABASCENS IS 319165 SYNTYPE A THR OSTACHY A 2133192 TYPE ATRACTODES 2096188 I SOTY PE ATROFUSCA VAR DECOLORATA 622651 HOLOTY P E ATROSQUAMA 817295 YOLOTYPE AUTUMNAL IS 1933437 ISOTYPE A ZUAYA E 2373748 ISOTYPF BAY8USETORUY 1232938 SVNTYPE BANKSI I 415269 TYPE COLLECTION 6 R A C HY POD A 964504 TYPE COLLECT ION BRA I NERD I I 711129 TYPE BRUNNEA VAR SUBTEIOGYNA 587668 TYPS COLLECTION RULBOSTYL I S 29741 TYPE COLLECTION CAL I FORN ICA 319268 TYPE COLLECTION CAJ_I FORNICA 6909 37 TYPE CDLLECTION C A Y P L YOC A R P A 1885701 ISOTYPE CAM PYLOCARP A SSP AFFINIS 368814 SUN TYP E CHALCIOLEPIS 28433 ISCTYPE C HA P MANI 2460272 HOLr3TY PE CHIAPENS IS 306281 I SOTYPE CHIHUAHUAENS I S 462090 (SOTYPE CILIARIS 28457 TYPE COLLECT ION CINNAMOYEA 319228 TYPE COLLECTION CI NNAMOYE A 350c77 TYPE COLLECTION CC A DO S T AC HY A VAR MAXIMA 1839933 I SOTY PE CLIVICOLA 20 388 2 2 TYPE COLLECTION COMANS VAR STRICTA 2723 5 TYPE MAT ERIAL CONFERT IFLORA 1032323 HOLOTYPE CONSPECTA 278555 I SOTYPE CRINITA VAR BREVICRINIS 278555 SYNTYPE CRINITA VAR SIMULANS 1682487 I SOTYPE C RUS -C 0 RV I VAR VIRGINIANA 13026G2 TYPE CDLLECT ION CUBENSIS 17657C.O I SOTY PE DANAENS IS 2003164 I SOTYPE DEBILIS VAR INTERCURSA 2003133 ISOTYPE DIGITAL IS VAR ASYMYETRICA 969128 TYPE COLLECTION DIGITAL IS VAR GLAUCA 176 11 5 1 ISOTYPE DIGITAL1S VAR MACROPODA 817314 TYPE COLLECTION DONNELL-SYITHI I 2176489 1 SOTYPE X DUMANII 182 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIVD OF TYPE TA XON us (U. S. NATIONAL HERBARIUM, SMITHSONIAN INSTITUTION, WASHINGTON)

579795 TYPE MATERIAL DURANDI I 857864 TYPE EGGLEST'ONII 2466328 HOLOTYPE EGGLESTONI I VAR FESTIVELLIFORMIS 1411790 COTYPE EKMANI I 1414090 TYPE COLLECTION EKYAN I I VAR HOTTENSI S 854950 TYPE YATERIAL ELMER1 1531248 TYPE YATERIAL FLRODl 368818 TYPE COLLECTION ELY NO IDES 270933 I SOTYPE EPAPILLOSA 2 2659 59 HOLDTYPE EURYSTACHYA 2 176495 I SOTY PE X EXSALINA 1123660 I SOTYPE FARGESI I 2 449 5 0 6 ISOTY PE F ISSA VAR ARISTATA 468192 TYPE FLACCIFOLIA 8 178 10 HOLOTYPE FRACTA 397187 TYPE COLLECT ION FUSCOLUTEA 251773 TYPE COLLECTION FUSCOTI NCT A 817237 TYPE COLLECTIQN F USC OT I NC T A 6352 5 TYPE MATERIAL GARBERI 660800 TYPE GEOPHI LA 319177 ISClTYPE GRACILIOR 29651 SYNTYPE HALL II 28685 I SOTYPE HAR.fORDI I 509004 ISOTYPE HELLERI 28206 TYPE COLLECTION HETERONEURA 802160 TYPE MATERIAL ICHANGENSIS 235568 TYPE COLLECTION I DA HOA 235569 TYPE COLLEC TION IDAHOA 2265956 HOLOTYP E I NC OND I TA 1872574 ISOTYPE INTERIMUS 1733722 I SOTYPE X NTERIOR VAR CHARLESTONENSIS 605797 I SOTYPE INTERIOR VAR JOSSELYNI I 16970 5 7 ISOTYPE INTFRIOR VAR KEWEENAWENSI S 30661 TYPE COLLECTION I NVOLUC RATELLA 82 5890 TYPE YATERIAL JAME SON I VAR SUBFULVA 132 5847 I SOTY PE KAT AH01NENS IS 2074700 TYPE MATERIAL KAUAIENSIS 430229 SYNTYPE LACUNARUM 801132 SYNTYPE LANC IFOLI A 193201 5 I SOTYPE LARENS! S 538796 HOLOTY PE LEIOPYYLLA 29211 TYPE COLLECTION LEYMONI 2231577 HOLOTYPE LIYNOPH I LA 800846 SYNTYPE L ON G ICQ UR I S VAR HENRY1 964080 TYPE COLLECTION LUZUL INA 1746479 I SOTYPE MACKENZ IANA 27238 TYPE MATERIAL. M ACR OGL 0 S S A 301267 TYPE MAORENS IS NUMBER 12 183

SHEET. Nile KIND OF TYPE TAXON us (U. S* NATIONAL HERBARIUM, SYITHSONIAN INSTITUTION, WASHINGTON)

969118 TYPE YATERIAL MAGNIFOLI 4 1789621 ISOTYPE MELOZITYENSIS 29453 I SOTYPE MENDOC INENS IS 711171 TYPE YATER IAL MERR ILL1 I 27281 TYPE MATERIAL MICANS 2501314 I SOTYPE MICRANTYA 30h9 5 I SOTYPE MICROGLOCHIN SSP FUEGINA 2 543 897 HOLOTY PE M ICROPT E RA VAR CRASSINERVIA 886422 HOL13 TYPE MISERABILI S 2133195 ISOTYPE MOHR IANA 23257 SYNTYPE MONTANE NS I S 2095886 I SOTYPE MORRI SSEYI 2728C TYPE M4TERI 4L NANA 1438017 TYPE COLLECTION NEBRASKENS IS V AR ERUCAE FORM IS 2433719 I SOTYPE X NEDRIGELOWI? 2 176496 I SOTY PE X NEOPALEACEA 286861 I SOTYPE NERVINA 31277 TYPE YATERIAL NUTANS VAR JAPONICA 251772 TYPE COLLECT ION OAXACANA 817656 TYPE COLLECTION OAX 4CANA 16781 88 ISOTYP E OB I SPOENSI S 30695 1 SOTYPE OL I GANTHA 504456 I SOTYPE ONUSTA 27292 TYPE YATER IAL OXYC4RPA 415172 TYPE PAC HY STOM A 529528 TYPE YATERIAL PADDOENSI S 872860 TYPE Y4TERIAL PALA WAN ENS IS 31344 ISOTYPE PAPULOSA 27275 TYPE MATERIAL PARCIFLORA 2133207 TYPE PERCOSTATA 461358 I SOTYPE PERLONGA 154583 1 TYPE P HA L ARO I DE S VAR PARVULA 31374 TYPE Y4TERIAL PICTA 15638 11 TYPE PIRCHINCHENSIS VAR SIMPLEX 660821 TYPE PITYOPHILA 27269 TYPE YATERI AL PLANAT A 24202 76 HOLOTYPE P L E C TOC A RPA 2074725 TYPE YATERIAL PLUVICA VAR KOOLAUENSI S 27270 TYPE YAYERI AL POD OGY NA 458 108 TYPE YATERIAL PRAINII 2265957 HOI.OTY PE PRAT ICOLA VAR SUBCORIACEA 865056 TYPE MATERIAL PRESLI I 817724 TYPE CDLLECTION PR INGLE I 1123683 ISOTY PE PTEROL EPTA 1282178 TYPE PUR P UREOVAG IN ATA VAR ITATIAIAE 1398 8 3C TYPE M4TERIAL PYCNOTHY SOS 243371 8 IS OTY P E X QUEBECENSIS 2133193 TYPE QUICHENS IS 184 SMITHSONIAN CONTRIBUTIONS TO BOTANY

SHEET NO. KIND OF TYPE TA XON us (U. So NATIONAL HERBARIUM, SMITHSONIAN INSTITUTION, WASHINGTON)

1872576 TYPE MATERIAL RACHILL IS 626608 TYPE YATERI AL R AMOS I I 2133190 HOLOTYPE ROANENSI S 1754487 ISOTYPE RUBRO-BRUNNEA VAR ELINEOLATA 200 3 132 ISOTY PE RUG AT A 30267 TYPE COLLECTION P. use yr 29888 TYPE COLLECT I3N SAL INAEFORY IS 319226 TYPE COLLECTION SALI NAEFORMIS 154575 2 TYPE SACTAENSI S 2176493 I SOTY PE X SAXENII NM. FERRUGINEA 528631 ISOTYPE SCABRIUSCULA 305734 TYPE COCC ECT ION SEATONIANA 528495 TYPE COLLECTION SHELDON II 511177 TYPE COLLECTION SMALLIANA 1736782 IS CITY PE SONOMENSIS 30329 SYNTYPE SPECIFICA 20063 86 ISOTYPE S PECUIC OL A 452499 HOLOTY PE STELLATA 616142 TYPE YATERI AL STENOPHY L L A VAR DESERTORUM 96909 1 TYPE COLLECTION STI PATA VAR MAXIMA 28683 ISOTYPE SUB- BR ACTE ATA 1765699 I SOTYPE SlJRNIGRI CANS 710428 TYPE MATERIAL SUB TR A N SVE R S A 143792 6 TYPE COLLECTION S UKS DOR F I I 1932033 ISDTYPE TACHIRENSIS 20923 56 TYPE MATERIAL TETSUOI 2604282 ISOTYPE TOYPKINS I 568126 YOLOTYPE TOWNSEND1 I 1675120 TYPE Y4TER IAL TSOI 1933688 I SOTYPE TURUMIQUIRENSIS 1872573 ISOTYPE VE R N AC UL A VAR HORSONII 20 50647 I SOTYPE VES ICARIA VAR LAURENTIANA 2 23 14%5 HOLOTYPE VEX AM 886234 TYPE VI? IDI OR 19678 19 ISOTY PE VITIENSIS 279151 I SOTYPE VITREA 2074653 TYPE COLLECTION WAHUENSIS VAR RUBIGINOSA 2133191 TYPE COLLECT ION WILLDENOWI I V A R M E G A R R H Y NC H A 2G50636 ISOTYPE X XANTHINA Bibliography

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Shaw, E. A. Stafleu, Frans A. 1971. The Gray Herbarium Card Index. Taxon, 20:333- 1967. Taxonomic Literature. xxi-546 pages. Utrecht: In- 336. ternational Bureau for Plant Taxonomy and No- Shetler, Stanwyn G. menclature. 1969. The Herbarium: Past, Present, and Future. Pro- Stafleu, F. A. (chairman), E. G. Voss (secretary), and mem- ceedings of the Biological Society of Washington, bers of Editorial Committee 82:687-758. [Flora North America Report, Number 1972. International Code of Botanical Nomenclature. 28.1 Regnum Vegetabile, 82: 1-426. 1972. Information Systems and Data Banking. Pages 469- Stearn, William T. 497 in Radford, A. E., W.C. Dickison, C. R. Bell, 1966. Botanical Latin. xiv +566 pages. London: Thomas and coiitributors, Vascular Plant Systematics. vi+ Nelson and Sons. 543 pages. Chapel Hill, North Carolina: University United States National Bureau of Standards of North Carolina Student Stores. 1970. Federal Information Processing Standards Publica- Shetler, S. G., J. H. Beaman, M. E. Hale, L. E. Morse, J. J, tion (FIPS Pub) 10: Countries, Dependencies and Crockett, and R. A. Creighton. Areas of Special Sovereignty. 23 pages. Washington, 1971. Pilot Data Processing Systems for Floristic Informa- D.C.: U.S. Department of Commerce, National tion. Pages 275-310 in J. L. Cutbill (editor), Data Bureau of Standards. Processing in Biology and Geology (The Systematics United States National Herbarium Association Special Volume Number 3). xv-l-346 [1965.] Index to Principal Collections Represented in the pages. London and New York: Academic Press. U. S. National Herbarium. 74 pages, unpublished, [Flora A;orth America Report, Number 26.1 processed. Washington, D.C.: Smithsonian Institu- Shetler, Stanwyn G. (Program Director) and Harriet R. tion, Department of Botany. Meadow (Systems Development Manager) Willis, J. C. 1971. Flora North America: A Comprehensive Program 1966. A Dictionary of Flowering Plants. Seventh edition, of Biological Research, Information Systems De- revised by H. K. Airy Shaw. xxii+l214+liii pages. velopment, and Data Banking Concerned with the Cambridge, England: Cambridge University Press. Vascular Plants of North America North of Mexico. Wood, C. E., Jr., R. S. Cowan, and G. Buchheim Flora North America Report, 61:vii-k 1-124-l-ap- 1963. Botanical Nomenclature, Punched Cards, and pendices A-I. Machines. Taxon, 12:2-12.