Untangling the Dinosaur Family Tree
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Ischigualasto Formation. the Second Is a Sile- Diversity Or Abundance, but This Result Was Based on Only 19 of Saurid, Ignotosaurus Fragilis (Fig
This article was downloaded by: [University of Chicago Library] On: 10 October 2013, At: 10:52 Publisher: Taylor & Francis Informa Ltd Registered in England and Wales Registered Number: 1072954 Registered office: Mortimer House, 37-41 Mortimer Street, London W1T 3JH, UK Journal of Vertebrate Paleontology Publication details, including instructions for authors and subscription information: http://www.tandfonline.com/loi/ujvp20 Vertebrate succession in the Ischigualasto Formation Ricardo N. Martínez a , Cecilia Apaldetti a b , Oscar A. Alcober a , Carina E. Colombi a b , Paul C. Sereno c , Eliana Fernandez a b , Paula Santi Malnis a b , Gustavo A. Correa a b & Diego Abelin a a Instituto y Museo de Ciencias Naturales, Universidad Nacional de San Juan , España 400 (norte), San Juan , Argentina , CP5400 b Consejo Nacional de Investigaciones Científicas y Técnicas , Buenos Aires , Argentina c Department of Organismal Biology and Anatomy, and Committee on Evolutionary Biology , University of Chicago , 1027 East 57th Street, Chicago , Illinois , 60637 , U.S.A. Published online: 08 Oct 2013. To cite this article: Ricardo N. Martínez , Cecilia Apaldetti , Oscar A. Alcober , Carina E. Colombi , Paul C. Sereno , Eliana Fernandez , Paula Santi Malnis , Gustavo A. Correa & Diego Abelin (2012) Vertebrate succession in the Ischigualasto Formation, Journal of Vertebrate Paleontology, 32:sup1, 10-30, DOI: 10.1080/02724634.2013.818546 To link to this article: http://dx.doi.org/10.1080/02724634.2013.818546 PLEASE SCROLL DOWN FOR ARTICLE Taylor & Francis makes every effort to ensure the accuracy of all the information (the “Content”) contained in the publications on our platform. However, Taylor & Francis, our agents, and our licensors make no representations or warranties whatsoever as to the accuracy, completeness, or suitability for any purpose of the Content. -
A Comprehensive Anatomical And
Journal of Paleontology, Volume 94, Memoir 78, 2020, p. 1–103 Copyright © 2020, The Paleontological Society. This is an Open Access article, distributed under the terms of the Creative Commons Attribution licence (http://creativecommons.org/ licenses/by/4.0/), which permits unrestricted re-use, distribution, and reproduction in any medium, provided the original work is properly cited. 0022-3360/20/1937-2337 doi: 10.1017/jpa.2020.14 A comprehensive anatomical and phylogenetic evaluation of Dilophosaurus wetherilli (Dinosauria, Theropoda) with descriptions of new specimens from the Kayenta Formation of northern Arizona Adam D. Marsh1,2 and Timothy B. Rowe1 1Jackson School of Geosciences, the University of Texas at Austin, 2305 Speedway Stop C1160, Austin, Texas 78712, USA <[email protected]><[email protected]> 2Division of Resource Management, Petrified Forest National Park, 1 Park Road #2217, Petrified Forest, Arizona 86028, USA Abstract.—Dilophosaurus wetherilli was the largest animal known to have lived on land in North America during the Early Jurassic. Despite its charismatic presence in pop culture and dinosaurian phylogenetic analyses, major aspects of the skeletal anatomy, taxonomy, ontogeny, and evolutionary relationships of this dinosaur remain unknown. Skeletons of this species were collected from the middle and lower part of the Kayenta Formation in the Navajo Nation in northern Arizona. Redescription of the holotype, referred, and previously undescribed specimens of Dilophosaurus wetherilli supports the existence of a single species of crested, large-bodied theropod in the Kayenta Formation. The parasagittal nasolacrimal crests are uniquely constructed by a small ridge on the nasal process of the premaxilla, dorsoventrally expanded nasal, and tall lacrimal that includes a posterior process behind the eye. -
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A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini Sullivan & Lucas, 1999 from the Upper Triassic of New Mexico, USA Martín D. EZCURRA Laboratorio de Anatomia Comparada y Evolución de los Vertebrados, Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Av. Angel Gallardo 470, Buenos Aires (1405) (Argentina) [email protected] Ezcurra M. D. 2006. — A review of the systematic position of the dinosauriform archosaur Eucoelophysis baldwini Sullivan & Lucas, 1999 from the Upper Triassic of New Mexico, USA. Geodiversitas 28 (4) : 649-684. Abstract Eucoelophysis baldwini Sullivan & Lucas, 1999 is represented by several post- cranial elements from the Petrified Forest Formation (Norian), New Mexico, USA. Eucoelophysis Sullivan & Lucas, 1999 was widely considered as a coelo- physoid dinosaur by several authors, but the hindlimb anatomy of this genus clearly indicates that it belongs to neither of these groups. The following fea- tures exclude Eucoelophysis from Neotheropoda: absence of oblique ligament groove on caudal surface of femoral head, femoral medial epicondyle small and smoothly rounded, absence of caudal cleft between medial part of the proximal end of the tibia and fibular condyles, cnemial crest low, and fibular crest absent. Moreover, Eucoelophysis lacks dinosaurian synapomorphic characters, but has a plesiomorphic slightly inturned femoral head that prevents its assignment to Dinosauria. Interestingly, the morphology of the femur of Eucoelophysis is extremely similar to that of the basal dinosauriform Silesaurus opolensis Dzik, Key words Dinosauriformes, 2003 from the Late Triassic of Poland. In order to determine the phylogenetic Coelophysoidea, position of Eucoelophysis, a cladistic analysis was carried out, which depicts Eu- Eucoelophysis, coelophysis as a non-dinosaurian dinosauriform. -
The Anatomy of Asilisaurus Kongwe, a Dinosauriform from the Lifua
THE ANATOMICAL RECORD (2019) The Anatomy of Asilisaurus kongwe,a Dinosauriform from the Lifua Member of the Manda Beds (~Middle Triassic) of Africa 1 2 3 STERLING J. NESBITT , * MAX C. LANGER, AND MARTIN D. EZCURRA 1Department of Geosciences, Virginia Tech, Blacksburg, Virginia 2Departamento de Biologia, Universidade de Sao~ Paulo, Ribeirao~ Preto, Brazil 3Sección Paleontología de Vertebrados CONICET—Museo Argentino de Ciencias Naturales “Bernardino Rivadavia”, Buenos Aires, Argentina ABSTRACT The diagnosis of Dinosauria and interrelationships of the earliest dino- saurs relies on careful documentation of the anatomy of their closest rela- tives. These close relatives, or dinosaur “precursors,” are typically only documented by a handful of fossils from across Pangea and nearly all speci- mens are typically missing important regions (e.g., forelimbs, pelves, skulls) that appear to be important to help resolving the relationships of dinosaurs. Here, we fully describe the known skeletal elements of Asilisaurus kongwe, a dinosauriform from the Middle Triassic Manda Beds of the Ruhuhu Basin of Tanzania. The taxon is known from many disarticulated and partially articulated remains and, most importantly, from a spectacularly preserved associated skeleton of an individual containing much of the skull, pectoral and pelvic girdles, forelimb and hindlimb, and parts of the vertebral column including much of the tail. The unprecedented detail of the anatomy indi- cates that Asilisaurus kongwe had a unique skull that was short and had both a premaxillary and dentary edentulous margin, but retained a number of character states plesiomorphic for Archosauria, including a crocodylian- like ankle configuration and a rather short foot with well-developed meta- tarsals I and V. -
From the Upper Triassic of North-Central New Mexico
Heckert, A.B., and Lucas, S.G., eds., 2002, Upper Triassic Stratigraphy and Paleontology. New Mexico Museum of Natural History and Science Bulletin No. 21. 189 THE TYPE LOCALITY OF BELODON BUCEROS COPE, 1881, A PHYTOSAUR (ARCHOSAURIA: PARASUCHIDAE) FROM THE UPPER TRIASSIC OF NORTH-CENTRAL NEW MEXICO SPENCER G. LUCAS, ANDREW B. HECKERT, KATE E. ZEIGLER and ADRIAN P. HUNT lNew Mexico Museum of Natural History, 1801 Mountain Road NW, Albuquerque, NM 87104-1375 Abstract-Here we establish the stratigraphic and geographic provenance of the "Belodon" buceros Cope. The holotype, originally collected by David Baldwin in 1881, is an incomplete phytosaur skull discov ered near "Huerfano Camp" in north-central New Mexico. This skull is the first phytosaur skull de scribed from the American West, and its precise provenance has never been established. Baldwin's use of the term Huerfano Camp may refer to Orphan Mesa, an isolated butte just south of Arroyo Seco. Fossils collected by Baldwin, and our subsequent collections from Orphan Mesa, are from a fossilifer ous interval high in the Petrified Formation of the Chinle Group. These strata yield a tetrapod fauna including the aetosaur Typothorax coccinarum and the phytosaur Pseudopalatus, both index taxa of the Revueltian (early-mid Norian) land-vertebrate faunachron. "Belodon" buceros is correctly referred to Pseudopalatus buceros (Cope) and is an index taxon of the Revueltian land-vertebrate faunachron. Keywords: phytosaur, Belodon, Pseudopalatus, Revueltian INTRODUCTION and its exact provenance has never been ascertained. Here, we establish the geographic and stratigraphic provenance of the ho In the Fall of 1874, Edward Drinker Cope (1840-1897) trav lotype of Belodon buceros and discuss its taxonomic position and eled through parts of north-central New Mexico and the San Juan biostratigraphic significance. -
The Origin and Early Evolution of Dinosaurs
Biol. Rev. (2010), 85, pp. 55–110. 55 doi:10.1111/j.1469-185X.2009.00094.x The origin and early evolution of dinosaurs Max C. Langer1∗,MartinD.Ezcurra2, Jonathas S. Bittencourt1 and Fernando E. Novas2,3 1Departamento de Biologia, FFCLRP, Universidade de S˜ao Paulo; Av. Bandeirantes 3900, Ribeir˜ao Preto-SP, Brazil 2Laboratorio de Anatomia Comparada y Evoluci´on de los Vertebrados, Museo Argentino de Ciencias Naturales ‘‘Bernardino Rivadavia’’, Avda. Angel Gallardo 470, Cdad. de Buenos Aires, Argentina 3CONICET (Consejo Nacional de Investigaciones Cient´ıficas y T´ecnicas); Avda. Rivadavia 1917 - Cdad. de Buenos Aires, Argentina (Received 28 November 2008; revised 09 July 2009; accepted 14 July 2009) ABSTRACT The oldest unequivocal records of Dinosauria were unearthed from Late Triassic rocks (approximately 230 Ma) accumulated over extensional rift basins in southwestern Pangea. The better known of these are Herrerasaurus ischigualastensis, Pisanosaurus mertii, Eoraptor lunensis,andPanphagia protos from the Ischigualasto Formation, Argentina, and Staurikosaurus pricei and Saturnalia tupiniquim from the Santa Maria Formation, Brazil. No uncontroversial dinosaur body fossils are known from older strata, but the Middle Triassic origin of the lineage may be inferred from both the footprint record and its sister-group relation to Ladinian basal dinosauromorphs. These include the typical Marasuchus lilloensis, more basal forms such as Lagerpeton and Dromomeron, as well as silesaurids: a possibly monophyletic group composed of Mid-Late Triassic forms that may represent immediate sister taxa to dinosaurs. The first phylogenetic definition to fit the current understanding of Dinosauria as a node-based taxon solely composed of mutually exclusive Saurischia and Ornithischia was given as ‘‘all descendants of the most recent common ancestor of birds and Triceratops’’. -
A Phylogenetic Analysis of the Basal Ornithischia (Reptilia, Dinosauria)
A PHYLOGENETIC ANALYSIS OF THE BASAL ORNITHISCHIA (REPTILIA, DINOSAURIA) Marc Richard Spencer A Thesis Submitted to the Graduate College of Bowling Green State University in partial fulfillment of the requirements of the degree of MASTER OF SCIENCE December 2007 Committee: Margaret M. Yacobucci, Advisor Don C. Steinker Daniel M. Pavuk © 2007 Marc Richard Spencer All Rights Reserved iii ABSTRACT Margaret M. Yacobucci, Advisor The placement of Lesothosaurus diagnosticus and the Heterodontosauridae within the Ornithischia has been problematic. Historically, Lesothosaurus has been regarded as a basal ornithischian dinosaur, the sister taxon to the Genasauria. Recent phylogenetic analyses, however, have placed Lesothosaurus as a more derived ornithischian within the Genasauria. The Fabrosauridae, of which Lesothosaurus was considered a member, has never been phylogenetically corroborated and has been considered a paraphyletic assemblage. Prior to recent phylogenetic analyses, the problematic Heterodontosauridae was placed within the Ornithopoda as the sister taxon to the Euornithopoda. The heterodontosaurids have also been considered as the basal member of the Cerapoda (Ornithopoda + Marginocephalia), the sister taxon to the Marginocephalia, and as the sister taxon to the Genasauria. To reevaluate the placement of these taxa, along with other basal ornithischians and more derived subclades, a phylogenetic analysis of 19 taxonomic units, including two outgroup taxa, was performed. Analysis of 97 characters and their associated character states culled, modified, and/or rescored from published literature based on published descriptions, produced four most parsimonious trees. Consistency and retention indices were calculated and a bootstrap analysis was performed to determine the relative support for the resultant phylogeny. The Ornithischia was recovered with Pisanosaurus as its basalmost member. -
A New Crested Theropod Dinosaur from the Early Jurassic of Yunnan
第55卷 第2期 古 脊 椎 动 物 学 报 pp. 177-186 2017年4月 VERTEBRATA PALASIATICA figs. 1-3 A new crested theropod dinosaur from the Early Jurassic of Yunnan Province, China WANG Guo-Fu1,2 YOU Hai-Lu3,4* PAN Shi-Gang5 WANG Tao5 (1 Fossil Research Center of Chuxiong Prefecture, Yunnan Province Chuxiong, Yunnan 675000) (2 Chuxiong Prefectural Museum Chuxiong, Yunnan 675000) (3 Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences, Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences Beijing 100044 * Corresponding author: [email protected]) (4 College of Earth Sciences, University of Chinese Academy of Sciences Beijing 100049) (5 Bureau of Land and Resources of Lufeng County Lufeng, Yunnan 650031) Abstract A new crested theropod, Shuangbaisaurus anlongbaoensis gen. et sp. nov., is reported. The new taxon is recovered from the Lower Jurassic Fengjiahe Formation of Shuangbai County, Chuxiong Yi Autonomous Prefecture, Yunnan Province, and is represented by a partial cranium. Shuangbaisaurus is unique in possessing parasagittal crests along the orbital dorsal rims. It is also distinguishable from the other two lager-bodied parasagittal crested Early Jurassic theropods (Dilophosaurus and Sinosaurus) by a unique combination of features, such as higher than long premaxillary body, elevated ventral edge of the premaxilla, and small upper temporal fenestra. Comparative morphological study indicates that “Dilophosaurus” sinensis could potentially be assigned to Sinosaurus, but probably not to the type species. The discovery of Shuangbaisaurus will help elucidate the evolution of basal theropods, especially the role of various bony cranial ornamentations had played in the differentiation of early theropods. -
University of Birmingham a New Dinosaur With
University of Birmingham A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil Marsola, Julio; Bittencourt, Jonathas; Butler, Richard; Da Rosa, Atila; Sayao, Juliana; Langer, Max DOI: 10.1080/02724634.2018.1531878 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Marsola, J, Bittencourt, J, Butler, R, Da Rosa, A, Sayao, J & Langer, M 2019, 'A new dinosaur with theropod affinities from the Late Triassic Santa Maria Formation, South Brazil', Journal of Vertebrate Paleontology, vol. 38, no. 5, e1531878. https://doi.org/10.1080/02724634.2018.1531878 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 25/07/2018 This is the accepted manuscript for a forthcoming publication in Journal of Vertebrate Paleontology. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. -
Comparative Bone Microstructure of Three Archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina
Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina JORDI ALEXIS GARCIA MARSÀ, FEDERICO L. AGNOLÍN, and FERNANDO E. NOVAS Marsà, J.A.G., Agnolín, F.L., and Novas, F.E. 2020. Comparative bone microstructure of three archosauromorphs from the Carnian, Late Triassic Chañares Formation of Argentina. Acta Palaeontologica Polonica 65 (2): 387–398. The Chañares Formation exhibits one of the most important archosauriform records of early Carnian ecosystems. Here we present new data on the palaeohistology of Chañares archosauriforms and provide new insights into their paleobiology, as well as possible phylogenetically informative traits. Bone microstructure of Lagerpeton chanarensis and Tropidosuchus romeri is dominated by fibro-lamellar tissue and dense vascularization. On the other hand, Chanaresuchus bonapartei is more densely vascularized, but with cyclical growth characterized by alternate fibro-lamellar, parallel-fibered and lamellar-zonal tissues. Dense vascularization and fibro-lamellar tissue imply fast growth and high metabolic rates for all these taxa. These histological traits may be tentatively interpreted as a possible adaptative advantage in front of Chañares Formation environmental conditions. Key words: Archosauromorpha, Lagerpeton, Tropidosuchus, paleobiology, paleohistology, Mesozoic, South America. Jordi Alexis Garcia Marsà [[email protected]] and Fernando E. Novas [[email protected]], Labora- torio de Anatomía Comparada y Evolución de los Vertebrados, -
Dinosaurs British Isles
DINOSAURS of the BRITISH ISLES Dean R. Lomax & Nobumichi Tamura Foreword by Dr Paul M. Barrett (Natural History Museum, London) Skeletal reconstructions by Scott Hartman, Jaime A. Headden & Gregory S. Paul Life and scene reconstructions by Nobumichi Tamura & James McKay CONTENTS Foreword by Dr Paul M. Barrett.............................................................................10 Foreword by the authors........................................................................................11 Acknowledgements................................................................................................12 Museum and institutional abbreviations...............................................................13 Introduction: An age-old interest..........................................................................16 What is a dinosaur?................................................................................................18 The question of birds and the ‘extinction’ of the dinosaurs..................................25 The age of dinosaurs..............................................................................................30 Taxonomy: The naming of species.......................................................................34 Dinosaur classification...........................................................................................37 Saurischian dinosaurs............................................................................................39 Theropoda............................................................................................................39 -
University of Birmingham the Earliest Bird-Line Archosaurs and The
University of Birmingham The earliest bird-line archosaurs and the assembly of the dinosaur body plan Nesbitt, Sterling; Butler, Richard; Ezcurra, Martin; Barrett, Paul; Stocker, Michelle; Angielczyk, Kenneth; Smith, Roger; Sidor, Christian; Niedzwiedzki, Grzegorz; Sennikov, Andrey; Charig, Alan DOI: 10.1038/nature22037 License: None: All rights reserved Document Version Peer reviewed version Citation for published version (Harvard): Nesbitt, S, Butler, R, Ezcurra, M, Barrett, P, Stocker, M, Angielczyk, K, Smith, R, Sidor, C, Niedzwiedzki, G, Sennikov, A & Charig, A 2017, 'The earliest bird-line archosaurs and the assembly of the dinosaur body plan', Nature, vol. 544, no. 7651, pp. 484-487. https://doi.org/10.1038/nature22037 Link to publication on Research at Birmingham portal Publisher Rights Statement: Checked for eligibility: 03/03/2017. General rights Unless a licence is specified above, all rights (including copyright and moral rights) in this document are retained by the authors and/or the copyright holders. The express permission of the copyright holder must be obtained for any use of this material other than for purposes permitted by law. •Users may freely distribute the URL that is used to identify this publication. •Users may download and/or print one copy of the publication from the University of Birmingham research portal for the purpose of private study or non-commercial research. •User may use extracts from the document in line with the concept of ‘fair dealing’ under the Copyright, Designs and Patents Act 1988 (?) •Users may not further distribute the material nor use it for the purposes of commercial gain. Where a licence is displayed above, please note the terms and conditions of the licence govern your use of this document.