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Insecta: Dermaptera) with a Check·List of Genera and Species
Rec. zool. Surv. India: 97 (Part-I) : 73-100, 1999 ON THE HIGHER CLASSIFICATION OF ANISOLABIDIDAE (INSECTA: DERMAPTERA) WITH A CHECK·LIST OF GENERA AND SPECIES G. K. SRIVASTAVA* Eastern Regional Station, Zoological Survey of Illdia, ShilJollg INTRODUCTION ~akai (1982) pointed out that family name Anisolabi( di)dae Verhoeff, 1902 has priority 6Verl~arcinophoridae Popham, 1965. Similarly the sub family Anisohibi(di)nae Zacher, 1911 baspriority overCarcinophorinae Hincks, 1954. Accordingly the former are used in the present work. A key for the discrimination of various subfamilies is given. The distinctive characters are based on various morphological character, mainly, on the body shape, shape of thoracic tergites, sternites and the male genitalia. It is proposed to erect a new subfamily Placolabidinae for the reception of Plac.:olabis Bey Bienko, 1959 as its type genus. This genus is very distinct from all t.he known genera of Anisolabidinae in having the median posterior prolongation of 8th sternite in males. Is·olaboidinae has been placed under this family by Srivastava (1996) which was transferred under Spongiphoridae (=Labidae) by Steinmann (1990). An attempt is made here to streamline the taxonomic status of various g~nera of AnisQlabidinae by providing akeyfortheirseparation. The main criteria fortheirdiscrilninatioll is based on the relative length of parameres besides its overall shape. The definition of various genera hOas been enlarged. The genera based on the shape of distal iobes and its associated stnlctures such as chiti nons accessory plates, shape and size of virga and the arrangement of teeth on chitinous pad may not prove t~ be stable characters since these vary intraspecifically. -
Taxonomy of Iberian Anisolabididae (Dermaptera)
Acta Zoologica Academiae Scientiarum Hungaricae 63(1), pp. 29–43, 2017 DOI: 10.17109/AZH.63.1.29.2017 TAXONOMY OF IBERIAN ANISOLABIDIDAE (DERMAPTERA) Mario García-París Museo Nacional de Ciencias Naturales, MNCN-CSIC c/José Gutiérrez Abascal, 2, 28006, Madrid. Spain. E-mail: [email protected] An update on the taxonomy and geographic distribution of Iberian Anisolabididae (Der- maptera) is provided. Former catalogues reported in the Iberian Peninsula three genera of Anisolabididae: Aborolabis, Anisolabis, and Euborellia. A revision of 487 specimens of Iberian and North African Anisolabidoidea permit to exclude the genus Aborolabis from the Iberian fauna, the re-assignation of inland Euborellia annulipes Iberian records to Euborellia moesta, and the exclusion of Aborolabis angulifera from Northwestern Africa. Examination of type materials of Aborolabis mordax and Aborolabis cerrobarjai allows to propose the treatment of A. cerrobarjai as a junior synonym of A. mordax. The diagnostic characters of Euborellia his- panica are included within the local variability found in E. moesta. I propose that E. hispanica should be treated as a junior synonym of E. moesta. Key words: earwigs, systematics, Mediterranean region, Spain, Morocco, NW Africa. INTRODUCTION The Iberian fauna of Dermaptera, including Anisolabididae Verhoeff, 1902, has been the subject of diverse revisionary (Bolívar 1876, 1897, Lapeira & Pascual 1980, Herrera Mesa 1980, Bivar de Sousa 1997) and compilatory works (Herrera Mesa 1999). These revisions together with the monograph of the Fauna of France (Albouy & Caussanel 1990) and the on-line information included in Fauna Europaea (Haas 2010), rendered the image of Dermaptera as a well known group in continental western Europe. -
Phylogeny of Morphologically Modified Epizoic Earwigs Based on Molecular Evidence
When the Body Hides the Ancestry: Phylogeny of Morphologically Modified Epizoic Earwigs Based on Molecular Evidence Petr Kocarek1*, Vaclav John2, Pavel Hulva2,3 1 Department of Biology and Ecology, Faculty of Science, University of Ostrava, Ostrava, Czech Republic, 2 Department of Zoology, Faculty of Science, Charles University in Prague, Prague, Czech Republic, 3 Life Science Research Centre, Faculty of Science, University of Ostrava, Ostrava, Czech Republic Abstract Here, we present a study regarding the phylogenetic positions of two enigmatic earwig lineages whose unique phenotypic traits evolved in connection with ectoparasitic relationships with mammals. Extant earwigs (Dermaptera) have traditionally been divided into three suborders: the Hemimerina, Arixeniina, and Forficulina. While the Forficulina are typical, well-known, free-living earwigs, the Hemimerina and Arixeniina are unusual epizoic groups living on molossid bats (Arixeniina) or murid rodents (Hemimerina). The monophyly of both epizoic lineages is well established, but their relationship to the remainder of the Dermaptera is controversial because of their extremely modified morphology with paedomorphic features. We present phylogenetic analyses that include molecular data (18S and 28S ribosomal DNA and histone-3) for both Arixeniina and Hemimerina for the first time. This data set enabled us to apply a rigorous cladistics approach and to test competing hypotheses that were previously scattered in the literature. Our results demonstrate that Arixeniidae and Hemimeridae belong in the dermapteran suborder Neodermaptera, infraorder Epidermaptera, and superfamily Forficuloidea. The results support the sister group relationships of Arixeniidae+Chelisochidae and Hemimeridae+Forficulidae. This study demonstrates the potential for rapid and substantial macroevolutionary changes at the morphological level as related to adaptive evolution, in this case linked to the utilization of a novel trophic niche based on an epizoic life strategy. -
Dermaptera, Forficulidae)
Dtsch. Entomol. Z. 68 (2) 2021, 235–248 | DOI 10.3897/dez.68.68020 The Pyrenean species of Chelidura (Dermaptera, Forficulidae) Pilar Jurado-Angulo1, Yolanda Jiménez-Ruiz1, Mario García-París1 1 Department of Biodiversity and Evolutionary Biology. Museo Nacional de Ciencias Naturales, MNCN-CSIC. c/ José Gutiérrez Abascal, 2. 28006, Madrid. Spain http://zoobank.org/43AD7562-7AF5-426D-A7EF-17225DD9AD98 Corresponding author: Pilar Jurado-Angulo ([email protected]) Academic editor: Susanne Randolf ♦ Received 28 April 2021 ♦ Accepted 6 July 2021 ♦ Published 10 August 2021 Abstract The Pyrenees are inhabited by scattered populations of earwigs of the genus Chelidura Latreille, 1825. There is some controversy about the specific assignment of these populations: while most authors assign them toC. pyrenaica (Gené, 1832), other consider that C. aptera (Mégerlé, 1825) is also present in the Pyrenees. The main objective of this work was to revise the identity and synonyms of Pyrenean Chelidura. Specimens from recent fieldwork and collections (MNCN-CSIC) were used for morphological and molecular studies (cytochrome oxidase 1). All Pyrenean specimens shared similar cox1 sequences, very divergent from those of Alpine C. ap- tera. As a consequence, the variability observed in male cerci morphology from the Pyrenees, ranging from long and slightly curved to short and very curved, corresponded to C. pyrenaica, and the presence of C. aptera in the Pyrenees can be rejected. As previously suggested by Maccagno (1933) and Fontana et al. (2021), the revision of the synonymic list uncovered the misplacement of the name F. simplex Germar, 1825 under the synonymy of C. aptera, while it rather represents a synonym of C. -
Dermaptera: Diplatyidae), Probable Evidence of Contact Between Neotropical and Malagasy Faunas
Zootaxa 3794 (4): 593–599 ISSN 1175-5326 (print edition) www.mapress.com/zootaxa/ Article ZOOTAXA Copyright © 2014 Magnolia Press ISSN 1175-5334 (online edition) http://dx.doi.org/10.11646/zootaxa.3794.4.11 http://zoobank.org/urn:lsid:zoobank.org:pub:EFFE09E5-3F9B-4A58-A56A-97E962323981 Mesodiplatys venado sp. nov. (Dermaptera: Diplatyidae), probable evidence of contact between Neotropical and Malagasy faunas LEONID N. ANISYUTKIN Zoological Institute of the Russian Academy of Sciences, Universitetskaya Emb. 1, 199034 Saint Petersburg, Russia. E-mail: [email protected], [email protected] Abstract A new earwig of the Malagasy genus Mesodiplatys stat. nov., M. venado sp. nov. (Dermaptera: Diplatyidae) is described from Peru (Departament Junin, Satipo Province). The diagnosis and composition of the genus Mesodiplatys stat. nov. are discussed. A detailed morphological description of the new species is given. The possible biogeographical significance of the find as evidence of link between South American and Malagasy fauna is briefly considered. Key words: earwigs, historical biogeography, morphology, Madagascar, Peru, South America Introduction The family Diplatyidae Verhoeff is a well-defined group of earwigs (Dermaptera). It is readily recognizable by the following character combination: head with large eyes, presence of well developed post-ocular carinae and a cylindrical abdomen, which is apically widened in males of most taxa (Hincks 1955; Sakai 1982, 1985). Diplatyidae is one of the early derivative groups of earwigs (Haas 1995; Haas & Klass 2003; Engel & Haas 2007) known at least from the Early Cretaceous (Engel 2011). The phylogenetic position of this family is rather obscure because links among the most archaic families of earwigs (Karschiellidae, Diplatyidae and Pygidicranidae) are unclear and inclusion of these families in Protodermaptera Zacher is based on symplesiomorphies (Grimaldi & Engel 2005). -
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46 Rec. zool. Sura. India Genus. Acrania Burr, 1915 Distribution: INDIA (Tamil Nadu, Himachal 1915. Acrania Burr. J.R. micro Soc., 1915: 432, 436 (Type Pradesh, West Bengal and Kashmir). Pygidicrana picta Guerin-Meneville) Elsewhere: Pakistan, Uganda and Zaire. 1993. Acrania; Srivastava, Rec. zool. Sura. India, 92(1-4): 44. Remarks : This species is recorded for the first time from Kashmir. 3. Acrania fletcheri (Bharadwaj and Kapoor, 1967) 5. Euborellia annulata (Fabricius, 1793) 1967. Cranopygia jletcheri Bharadwaj and Kapoor, Bull. Ent.,8(2) : 1(0"; India: Meghalaya; Shillong, 5000 1793. Forficula annulata Fabricius, Ent. Syst., II : 4 (sex; ft.) Americae meridionale). 1914. Kalocrania picta (nec.Guerin-Meneville); Burr, Rec. 1867. Anisolabis stali; Scudder, Proc. Boston soc. Nat. Hist., Indian Mus., 8(2): 136 (lO",l~, Kobo). 18: 308. 1993. Acrania jletcheri; Srivastava, Rec. zool. Sura. India, 1910. Borellia stali Burr, Fauna of British India, Dermaptera: 92(1-4): 45. 88. Material examined: INDIA: Manipur: Imphal, Material Examined : INDIA: Chennai, Elliots 16 miles North on Dimapur Road, Alt., 3500 ft., Beach, Radio Station, 10", 18.9.1961. 14.x.1945 (Major M.L. Roonwal). Measurements: (in mm): Male Measurements: (in mm): Male Length of body 10mm Length of body 20 Forceps l.5mm. Length of forceps 5.5 Distribution : INDIA (Kerala, Tamil Nadu, Distribution: INDIA: Meghalaya, (Shillong and Karnataka, Maharashtra, Orissa, West Bengal, Manipur) (Imphal district). Manipur and Lakshadweep islands). Elsewhere: Not yet recorded. Elsewhere: Sri Lanka, Pakistan and China. Remarks : It is reported for the first time from Sub family BRACHYLABIDINAE the state of Manipur. Genus Metisolabis Burr, 1910 Super family ANISOLABOIDEA 1910. -
André Nel Sixtieth Anniversary Festschrift
Palaeoentomology 002 (6): 534–555 ISSN 2624-2826 (print edition) https://www.mapress.com/j/pe/ PALAEOENTOMOLOGY PE Copyright © 2019 Magnolia Press Editorial ISSN 2624-2834 (online edition) https://doi.org/10.11646/palaeoentomology.2.6.1 http://zoobank.org/urn:lsid:zoobank.org:pub:25D35BD3-0C86-4BD6-B350-C98CA499A9B4 André Nel sixtieth anniversary Festschrift DANY AZAR1, 2, ROMAIN GARROUSTE3 & ANTONIO ARILLO4 1Lebanese University, Faculty of Sciences II, Department of Natural Sciences, P.O. Box: 26110217, Fanar, Matn, Lebanon. Email: [email protected] 2State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China. 3Institut de Systématique, Évolution, Biodiversité, ISYEB-UMR 7205-CNRS, MNHN, UPMC, EPHE, Muséum national d’Histoire naturelle, Sorbonne Universités, 57 rue Cuvier, CP 50, Entomologie, F-75005, Paris, France. 4Departamento de Biodiversidad, Ecología y Evolución, Facultad de Biología, Universidad Complutense, Madrid, Spain. FIGURE 1. Portrait of André Nel. During the last “International Congress on Fossil Insects, mainly by our esteemed Russian colleagues, and where Arthropods and Amber” held this year in the Dominican several of our members in the IPS contributed in edited volumes honoring some of our great scientists. Republic, we unanimously agreed—in the International This issue is a Festschrift to celebrate the 60th Palaeoentomological Society (IPS)—to honor our great birthday of Professor André Nel (from the ‘Muséum colleagues who have given us and the science (and still) national d’Histoire naturelle’, Paris) and constitutes significant knowledge on the evolution of fossil insects a tribute to him for his great ongoing, prolific and his and terrestrial arthropods over the years. -
(Insecta: Dermaptera) with a Check-List of Genera and Species
Rec. zool. Surv. India: 97 (Part-I) : 73-100, 1999 ON THE HIGHER CLASSIFICATION OF ANISOLABIDIDAE (INSECTA: DERMAPTERA) WITH A CHECK·LIST OF GENERA AND SPECIES G. K. SRIVASTAVA* Eastern Regional Station, Zoological Survey of Illdia, ShilJollg INTRODUCTION ~akai (1982) pointed out that family name Anisolabi( di)dae Verhoeff, 1902 has priority 6Verl~arcinophoridae Popham, 1965. Similarly the sub family Anisohibi(di)nae Zacher, 1911 baspriority overCarcinophorinae Hincks, 1954. Accordingly the former are used in the present work. A key for the discrimination of various subfamilies is given. The distinctive characters are based on various morphological character, mainly, on the body shape, shape of thoracic tergites, sternites and the male genitalia. It is proposed to erect a new subfamily Placolabidinae for the reception of Plac.:olabis Bey Bienko, 1959 as its type genus. This genus is very distinct from all t.he known genera of Anisolabidinae in having the median posterior prolongation of 8th sternite in males. Is·olaboidinae has been placed under this family by Srivastava (1996) which was transferred under Spongiphoridae (=Labidae) by Steinmann (1990). An attempt is made here to streamline the taxonomic status of various g~nera of AnisQlabidinae by providing akeyfortheirseparation. The main criteria fortheirdiscrilninatioll is based on the relative length of parameres besides its overall shape. The definition of various genera hOas been enlarged. The genera based on the shape of distal iobes and its associated stnlctures such as chiti nons accessory plates, shape and size of virga and the arrangement of teeth on chitinous pad may not prove t~ be stable characters since these vary intraspecifically. -
Big Wigs and Small Wigs: the Roles of Size, Sex and Shelter in Spatial Distribution Patterns in the Maritime Earwig Anisolabis Maritima
Big wigs and small wigs: the roles of size, sex and shelter in spatial distribution patterns in the maritime earwig Anisolabis maritima Nicole Hack1,2, Vikram Iyengar¹,3 Blinks/NSF REU/BEACON 2013 Summer 2013 Contact Information: Nicole Hack Ecology and Evolutionary Biology Department University of California Santa Cruz 1156 High St. Santa Cruz, CA 95064 [email protected] Keywords: Anisolabis maritima, maritime earwig, cohabitation, sex and size 1 Friday Harbor Laboratories, University of Washington, Friday Harbor, Washington 98250 2 Department of Ecology and Evolutionary Biology, University of California Santa Cruz, Santa Cruz, CA 95064 3 Department of Biology, Villanova University, Villanova, PA 19085 Hack 1 Abstract Animal aggregations can occur for a variety of abiotic factors, such as resource limitation, or biotic factors including sexual selection and predator-prey interactions. Although it is challenging to determine the underlying mechanism of such grouping behavior, we conducted experiments in which we examined the interactions and distribution patterns among pairs of the maritime earwig Anisolabis maritima (Order Dermaptera). This insect, found in aggregations under beach debris around the world, is sexually dimorphic regarding its most distinctive feature in that females have straight posterior forceps/pinchers whereas males have asymmetrical, curved forceps. We placed pairs of individuals varying in sex and size and monitored their distribution with and without shelter at 15 min, 12 h and 24 h to determine the roles that these factors may play in spatial patterns and gain insight into the mating system. Overall, we found that females were less likely to cohabitate than males, and they were more tolerant of males than other females. -
Ecological Considerations for Development of the Wildlife Lake, Castlereagh
Ecological considerations for development of the Wildlife Lake, Castlereagh Total Catchment Management Services Pty Ltd August 2009 Clarifying statement This report provides strategic guidance for the site. Importantly this is an informing document to help guide the restoration and development of the site and in that respect does not contain any matters for which approval is sought. Disclaimer The information contained in this document remains confidential as between Total Catchment Management Services Pty Ltd (the Consultant) and Penrith Lakes Development Corporation (the Client). To the maximum extent permitted by law, the Consultant will not be liable to the Client or any other person (whether under the law of contract, tort, statute or otherwise) for any loss, claim, demand, cost, expense or damage arising in any way out of or in connection with, or as a result of reliance by any person on: • the information contained in this document (or due to any inaccuracy, error or omission in such information); or • any other written or oral communication in respect of the historical or intended business dealings between the Consultant and the Client. Notwithstanding the above, the Consultant's maximum liability to the Client is limited to the aggregate amount of fees payable for services under the Terms and Conditions between the Consultant and the Client. Any information or advice provided in this document is provided having regard to the prevailing environmental conditions at the time of giving that information or advice. The relevance and accuracy of that information or advice may be materially affected by a change in the environmental conditions after the date that information or advice was provided. -
ARTHROPODA Subphylum Hexapoda Protura, Springtails, Diplura, and Insects
NINE Phylum ARTHROPODA SUBPHYLUM HEXAPODA Protura, springtails, Diplura, and insects ROD P. MACFARLANE, PETER A. MADDISON, IAN G. ANDREW, JOCELYN A. BERRY, PETER M. JOHNS, ROBERT J. B. HOARE, MARIE-CLAUDE LARIVIÈRE, PENELOPE GREENSLADE, ROSA C. HENDERSON, COURTenaY N. SMITHERS, RicarDO L. PALMA, JOHN B. WARD, ROBERT L. C. PILGRIM, DaVID R. TOWNS, IAN McLELLAN, DAVID A. J. TEULON, TERRY R. HITCHINGS, VICTOR F. EASTOP, NICHOLAS A. MARTIN, MURRAY J. FLETCHER, MARLON A. W. STUFKENS, PAMELA J. DALE, Daniel BURCKHARDT, THOMAS R. BUCKLEY, STEVEN A. TREWICK defining feature of the Hexapoda, as the name suggests, is six legs. Also, the body comprises a head, thorax, and abdomen. The number A of abdominal segments varies, however; there are only six in the Collembola (springtails), 9–12 in the Protura, and 10 in the Diplura, whereas in all other hexapods there are strictly 11. Insects are now regarded as comprising only those hexapods with 11 abdominal segments. Whereas crustaceans are the dominant group of arthropods in the sea, hexapods prevail on land, in numbers and biomass. Altogether, the Hexapoda constitutes the most diverse group of animals – the estimated number of described species worldwide is just over 900,000, with the beetles (order Coleoptera) comprising more than a third of these. Today, the Hexapoda is considered to contain four classes – the Insecta, and the Protura, Collembola, and Diplura. The latter three classes were formerly allied with the insect orders Archaeognatha (jumping bristletails) and Thysanura (silverfish) as the insect subclass Apterygota (‘wingless’). The Apterygota is now regarded as an artificial assemblage (Bitsch & Bitsch 2000). -
Insect Classification Standards 2020
RECOMMENDED INSECT CLASSIFICATION FOR UGA ENTOMOLOGY CLASSES (2020) In an effort to standardize the hexapod classification systems being taught to our students by our faculty in multiple courses across three UGA campuses, I recommend that the Entomology Department adopts the basic system presented in the following textbook: Triplehorn, C.A. and N.F. Johnson. 2005. Borror and DeLong’s Introduction to the Study of Insects. 7th ed. Thomson Brooks/Cole, Belmont CA, 864 pp. This book was chosen for a variety of reasons. It is widely used in the U.S. as the textbook for Insect Taxonomy classes, including our class at UGA. It focuses on North American taxa. The authors were cautious, presenting changes only after they have been widely accepted by the taxonomic community. Below is an annotated summary of the T&J (2005) classification. Some of the more familiar taxa above the ordinal level are given in caps. Some of the more important and familiar suborders and families are indented and listed beneath each order. Note that this is neither an exhaustive nor representative list of suborders and families. It was provided simply to clarify which taxa are impacted by some of more important classification changes. Please consult T&J (2005) for information about taxa that are not listed below. Unfortunately, T&J (2005) is now badly outdated with respect to some significant classification changes. Therefore, in the classification standard provided below, some well corroborated and broadly accepted updates have been made to their classification scheme. Feel free to contact me if you have any questions about this classification.