FIELDIANA

Life and Earth Sciences

NO. 8

A Checklist and Key to the Homalopsid Snakes (Reptilia, Squamata, Serpentes), with the Description of New Genera

John C. Murphy Harold K. Voris

Science and Education Science and Education Field Museum of Natural History Field Museum of Natural History 1400 South Lake Shore Drive 1400 South Lake Shore Drive Chicago, Illinois 60605 USA Chicago, Illinois 60605 USA Current address: 15824 Weather Vane Way Plainfield, Illinois 60544 USA E-mail: serpentresearch@gmail. com

Accepted April 28, 2014 Published September 24, 2014 Publication 1567 Associate Editor for this volume was Kenneth Angielczyk.

PUBLISHED BY FIELD MUSEUM OF NATURAL HISTORY Table of Contents

Abstract 1 Historical Background 1 What Are Homalopsid Snakes? 2 Generic Designations, Why So Many Genera? 2 HOMALOPSIDAE INCERTAE SeDIS 4 Common Names and Photographs 4 Methods 4 Accounts for Genera and Species 6 Fangless Homalopsids 6 Brachyorrhos 6 Calamophis 7 Karnsophis 9 Fanged Homalopsids 10 Bitia 10 Cantor ia 10 Cerberus H Dieurostus 13 Djokoiskandarus 14 Enhydris 15 Erpeton 18 Ferania 19 Fordonia 20 Gerarda 20 Gyiophis new genus 21 Heurnia 22 Homalophis 23

Homalopsis • 24 Hypsiscopus 25

Kualatahan new genus » 27 Mintonophis new genus 27 Miralia 28 Myron 29 Myrrophis 30 Phytolopsis 32 Pseudoferania 32 Raclitia 33 Subsessor new genus 33 Sumatranus new genus 34 Taxonomic Keys for the Homalopsidae 37 Acknowledgments 39 Literature Cited 39

List of Figures

Figure 1. Brachyorrhos albus 6 Figure 2. Brachyorrhos gastrotaenius 7 Figure 3. Brachyorrhos raffrayi 7 Figure 4. Brachyorrhos wallacei 8 Figure 5. Calamophis katesandersae 8 Figure 6. Calamophis ruuddelangi 9 Figure 7. Calamophis sharonbrooksae 9 Figure 8. Karnsophis siantaris 10 Figure 9. Bitia hydroides 10 Figure 10. Cantoria violacea H Figure 11. Cerberus australis 12 Figure 12. Cerberus dunsoni 12 Figure 13. Cerberus microlepis 13 Figure 14. Cerberus rynchops 13 Figure 15. Cerberus Schneiderii 14 Figure 16. Dieurostus dussumierii 14 Figure 17. Djokoiskandarus annulatus 15

iii Figure 18. Enhydris chanardi j-j Figure 19. Enhydris enhydris j° Figure 20. Enhydris innominata Yl Figure 21. Enhydris jagorii ^ Figure 22. Enhydris longicauda J° Figure 23. Enhydris subtaeniata jj> Figure 24. Erpeton tentaculatus Figure 25. Ferania sieboldii Figure 26. Fordonia leucobalia fO Figure 27. Gerarda prevostiana 2j Figure 28. Gyiophis maculosa 21 Figure 29. Gyiophis vorisi 22 Figure 30. Heurnia ventromaculata 22 Figure 31. Homalophis doriae 23 Figure 32. Homalophis gyii 23 Figure 33. Homalopsis buccata 24 Figure 34. Homalopsis mereljcoxi 2^ Figure 35. Homalopsis nigroventralis 2^ Figure 36. Homalopsis semizonata 2" Figure 37. Hypsiscopus matannensis 2^ Figure 38. Hypsiscopus plumbea 2^ Figure 39. Kualatahan pahangensis 2^ Figure 40. Mintonophis pakistanicus 2° Figure 41. Miralia alternans 3? Figure 42. Myron karnsi 2" Figure 43. Myron resetari 30 Figure 44. Myron richardsonii 30 Figure 45. Myrrophis bennettii 31 Figure 46. Myrrophis chinensis 31 Figure 47. Phytolopsis punctata 32 Figure 48. Pseudoferania polylepis 33 Figure 49. Raclitia indica 33 Figure 50. Subsessor bocourti 34 Figure 51. Sumatranus albomaculatus 34 Figure 52. Key Plate I 35 Figure 53. Key Plate II 36

List of Tables

Table 1. Scientific names and geographical distribution 3 Table 2. Homalopsid species groups 4

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