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Käferarten Am Wildapfel Heinz Bußler Käferarten am Wildapfel Heinz Bußler Schlüsselwörter: Wildapfel, Streuobstbestände mit Kul- Borken- und Rüsselkäfer turapfelsorten, Käferarten Fünf Borkenkäferarten entwickeln sich auch in Apfel- Zusammenfassung: Der Wildapfel (Malus sylvestris) ist bäumen. Weit verbreitet sind der Große Obstbaum - eine sehr seltene Baumart, deshalb gibt es bisher keine splintkäfer (Scolytus mali) und der Kleine Obst baum- speziellen Untersuchungen der daran vorkommenden splint käfer (Scolytus rugulosus). Im Splintholz toter Käferfauna. In fränkischen Streuobstbeständen mit alten oder geschwächter Partien züchten der Ungleiche Kulturapfelsorten konnten bisher über 100 Käferarten Holzbohrer (Xyleborus dispar), der Asiatische Nutz- nachgewiesen werden. Welche der angeführten Käfer- holzborkenkäfer (Xyleborus germanus) und der Kleine arten auch am Wildapfel vorkommen, ist unbekannt. Der Holzbohrer (Xyleborus saxesenii) Ambrosiapilze, die überwiegende Teil des holzbewohnenden (xylobionten) den Larven als Nahrung dienen. Käferspektrums der Kulturapfelbäume ist in wärmege- tönten Eichen- und Eichenmischwäldern beheimatet. Da Eine häufige und oft sogar schädliche Art ist der Apfel- der Wildapfel seinen Verbreitungsschwerpunkt in diesen blütenstecher (Anthonomus pomorum). Der Rüssel- Waldgesellschaften hat, ist zu vermuten, dass alle darin käfer kommt bereits im März aus dem Winterquartier. vorkommenden Arten auch an ihm potenziell auftreten Im April legt er seine Eier in die Blütenknospen. Die könnten. Larven fressen im Inneren der Knospen, was zur Folge hat, dass sich diese nicht öffnen und später sogar ver- trocknen. Auch die Verpuppung findet innerhalb der vertrockneten Knospe statt. Die neue Käfergeneration Aufgrund der Seltenheit des Wildapfels (Malus syl- schlüpft bereits im Juni, frisst dann noch eine Weile an vestris) in Bayern (Huber und Wurm 2013) gibt es bisher Blättern, bevor die Käfer zur Sommer- und späteren keine speziellen Untersuchungen der Käferfauna die- Winterpause Verstecke im Boden, Laub oder Rinde ser Baumart. In den Jahren 1995 und 1996 erfolgten aufsuchen. Der Apfelblütenstecher ist ein wichtiger jedoch Kartierungen in mittel- und oberfränkischen Schädling im Obstbau, der ohne Bekämpfungsmaß- Streuobstbeständen mit alten Kulturapfelsorten, die nahmen wirtschaftliche Schäden verursachen kann einen gewissen Rückschluss auf das potenzielle Spek- (Rheinheimer und Hassler 2010). Auch fünf holzbesie- trum der Käferarten am Wildapfel erlauben (Bußler delnde Rüsselkäferarten wurden in Apfelbaumtotholz 1997; Schmidl 2000). Die damals erfasste Käferfauna der festgestellt: Phloeophagus lignarius, Stereocorynes Kulturapfelbäume umfasste über 100 Arten, wobei truncorum, Magdalis ruficornis, Magdalis barbicornis keine der Arten monophag an Apfelbäumen lebt. Alle und Magdalis cerasi. Arten entwickeln sich auch in anderen baumförmigen Rosengewächsen (Rosaceae) oder sind Strukturspe- zialisten (zum Beispiel Baumhöhlenbewohner), die an Lauf-, Hirsch-, Pracht- und Bockkäfer keine bestimmte Baumart, sondern an das Auftreten der Struktur gebunden sind. Die schuppige Rinde der Apfelbäume dient einer großen Anzahl von baumbewohnenden (arboricolen) und räuberischen Laufkäfern als Lebensraum. Bisher wurden sieben Arten nachgewiesen: Dromius linearis, Dromius agilis, Dromius quadrimaculatus, Philorhizus melanocephalus, Philorhizus sigma, Philorhizus not- atus und Caladromius spilotus. Vier der sieben heimi- schen Hirschkäferarten können sich ebenfalls im Tot- holz von Apfelbäumen entwickeln, der Balkenschröter (Dorcus parallelipipedus), der Rehschröter (Platycerus LWF Wissen 73 41 Käferarten am Wildapfel caraboides), der Kopfhornschröter (Sinodendron cylin- dricum) (Abbildung 1) und der Hirschkäfer oder Feu- erschröter (Lucanus cervus). Nur zwei Prachtkäfer- arten wurden bisher an Apfelbäumen festgestellt: Anthaxia nitidula und der farbenprächtige Bunte Apfel- Balkenschröter baum-Prachtkäfer (Anthaxia suzannae) (Abbildung 2). (Dorcus parallelipipedus) Diese Art ist aber, auch wenn der deutsche Name das vermuten lässt, ebenfalls nicht streng an Apfelbäume gebunden; Nachweise liegen auch von anderen baum- förmigen Rosengewächsen vor, unter anderem vom Speierling (Sorbus domestica), der Quitte (Cydonia oblonga) und dem Weißdorn (Crataegus spp.). Das Spektrum der Bockkäfer an Apfelbäumen umfasst bisher 13 Arten, darun ter mit dem Kleinen Eichenbock (Cerambyx scopolii) und Anisarthron barbipes zwei gefährdete Arten der Roten Listen. Rehschröter (Platycerus caraboides) Holzpilzbesiedler und Mulmhöhlen- bewohner Von Apfelbäumen ist ein reiches Pilzartenspektrum dokumentiert (siehe Beitrag Blaschke und Nannig). Bei Untersuchungen im Bereich der Mainfränkischen Platten konnte Krieglsteiner 1999 allein 51 Holzzersetzer an der Gattung Malus nachweisen. Häufigste Vertreter waren hier der Zottige Schillerporling (Inonotus hisp i - dus) und die beiden Schlauchpilze Hypoxylon rubigi- nosum und Propolomyces versicolor. Auch bei den Untersuchungen im Biosphärenreservat Rhön wurden Kopfhornschröter 51 Pilzarten am Holz von Apfelbäumen festgestellt (Sinodendron cylindricum) (Krieglsteiner 2004). Die Gilde der holzpilzbesiedelnden Käferarten stellt deshalb circa ein Fünftel des bisher an Apfelbäumen nachgewiesenen Käferspektrums, darun - ter eine Anzahl von gefährdeten Arten. Eine Schlüssel- rolle für die Höhlen- und Mulmhöhlenbildung an Apfel- bäumen spielt der Zottige Schillerporling (Inonotus hispidus). Er besiedelt nur lebende Bäume und ist ein Wundparasit, dessen Myzel eine intensive Weißfäule im Kernholz erzeugt. Die Splint-, Kambial- und Rinden- schicht des Baumes bleibt zu nächst unbeeinträchtigt, so dass die Wasser- und Nährstoffversorgung sowie die Hirschkäfer Standfestigkeit des Baumes lange gewährleistet blei - (Lucanus cervus) ben. Der Zottige Schillerporling ist neben dem Schwe- felporling (Laetiporus sulphurescens) der wichtigste Mulmhöhlenbildner. Bisher konnten vier gefährdete Blatthornkäferarten in Höhlen von Apfelbäumen nach- Abbildung 1: gewiesen werden, der Marmorierte Rosenkäfer (Pro - Heimische Hirsch käfer- tae tia lugubris) (Abbildung 3), der Eremit oder Juch- arten, die sich im Totholz von Apfelbäumen tenkäfer (Osmoderma eremita), Protaetia fieberi und entwickeln können. Gnorimus nobilis. Foto: U. Schmidt, www.kaefer-der-welt.de 42 LWF Wissen 73 Käferarten am Wildapfel Abbildung 2: Bunter Apfelbaum-Prachtkäfer Abbildung 3: Marmorierter Rosenkäfer (Anthaxia suzannae) Foto: H. Bußler (Protaetia lugubris) Foto: H. Bußler Literatur Der Apfelbaum ist Bußler, H. (1997): Die Besiedlung anthropogen geprägter Lebensräume durch xylobionte Käferarten am Beispiel fränki- scher Streuobstbestände. Berichte der ANL 21, S.179–187 aufgeblüht Huber, G.; Wurm, A. (2013): Wildapfel – Baum des Jahres. LWF aktuell 94, S.23 Der Apfelbaum ist aufgeblüht. Krieglsteiner, L. (1999): Pilze im Naturraum Mainfränkische Nun summen alle Bienen. Platten und ihre Einbindung in die Vegetation. Regensburger Mykologische Schriften 9, 905 S. Die Meise singt ein Meisenlied. Der Frühling ist erschienen. Krieglsteiner, L. (2004): Pilze im Biosphären-Reservat Rhön und ihre Einbindung in die Vegetation. Regensburger Mykologi- sche Schriften 12, 770 S. Die Sonne wärmt den Apfelbaum. Rheinheimer, J.; Hassler, M. (2010): Die Rüsselkäfer Baden-Würt- Der Mond scheint auf ihn nieder. tembergs. Verlag Regionalkultur Heidelberg-Ubstadt, Weiher- Neustadt a.d.W., Basel, 944 S. Die kleine Meise singt im Traum Schmidl, J. (2000): Bewertung von Streuobstbeständen mittels die Apfelblütenlieder. xylobionter Käfer am Beispiel Frankens. Naturschutz und Land- schaftsplanung 31, S.357–372 Die Bienen schwärmen Tag für Tag und naschen von den Blüten. Keywords: Wild crab, orchards, beetles Mag sie der Mai vor Hagelschlag Summary: The wild crab (Malus sylvestris) is a rare species, und hartem Frost behüten. so there have been no specific studies of the beetle fauna of this tree. In orchards in Franconia with ancient culture Der Apfelbaum ist aufgeblüht. apple varieties over 100 species of beetles have been detected. Which of the listed beetle species that have Der Winter ist vorbei. been detected in the orchards, also occur on wild crab, is Mit Blütenduft und Meisenlied unknown. The vast majority of saproxylic beetles spec - trum of cultivated apple trees is naturally at home in heat- erscheint der junge Mai. tinted oak and mixed oak forests. Since the wild crab has its center of distribution in these forest communities, it is James Krüss (1926–1997) likely that all species could potentially occur to him. © James-Krüss-Erbengemeinschaft LWF Wissen 73 43.
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