ISSN 1211-8788 Acta Musei Moraviae, Scientiae biologicae (Brno) 94: 87–116, 2009

The proposed Úvalský rybník Nature Monument, an important refuge for wetland insects in South Moravia (Czech Republic): A species inventory of Odonata, Heteroptera and Coleoptera (partim) with the first Czech record of Corixa panzeri (Corixidae)

MICHAL STRAKA1, PETR KMENT 2, JAN SYCHRA3 & JAN HELEŠIC 4 1 Department of Botany and Zoology, Masaryk University, Kotláøská 2, CZ-611 37 Brno, Czech Republic; e-mail: [email protected] 2 Department of Entomology, National Museum, Kunratice 1, CZ-148 00 Praha 4, Czech Republic; e-mail: [email protected] 3 Department of Botany and Zoology, Masaryk University, Kotláøská 2, CZ-611 37 Brno, Czech Republic; e-mail: [email protected] 4 Rybáøe 34, CZ-691 53 Tvrdonice, Czech Republic; e-mail: [email protected]

STRAKA M., KMENT P., SYCHRA J. & HELEŠIC J. 2009: The proposed Úvalský rybník Nature Monument, an important refuge for wetland insects in South Moravia (Czech Republic): A species inventory of Odonata, Heteroptera and Coleoptera (partim) with the first Czech record of Corixa panzeri (Corixidae). Acta Musei Moraviae, Scientiae biologicae (Brno) 94: 87–116. – In the course of surveys in the years 2000–2003 and especially in 2009, 162 species of Odonata (21), Heteroptera (50) and Coleoptera (Haliplidae, Noteridae, Dytiscidae, Carabidae, Helophoridae, Hydrophilidae, Hydraenidae, Scirtidae, Dryopidae, Heteroceridae) (91) were recorded within the area of the proposed Úvalský rybník Nature Monument. One species, Corixa panzeri Fieber, 1848 (Heteroptera: Corixidae), was recorded for the first time in the Czech Republic. Six of the recorded species are classified as critically endangered in the Czech Republic: Corixa affinis Leach, 1817, Haliplus furcatus Seidlitz, 1887, H. variegatus Sturm, 1834, Cybister lateralimarginalis (DeGeer, 1774), Hydrophilus atterimus (Eschscholtz, 1882) and Hydraena paganettii Ganglbauer, 1901; five species are endangered: Sympetrum meridionale (Sélys, 1841), Bidessus nasutus Sharp, 1887, Hydrovatus cuspidatus (Kunze, 1818), Pterostichus cursor (Dejean, 1828) and Dryops similaris Bollow, 1936; fourteen are classified as vulnerable and five as near threatened. The occurrence of the endangered species is commented upon and suggestions made concerning the management of this unique locality. Keywords. Odonata, Heteroptera, Coleoptera, species inventory, salt marsh, nature conservation, Moravia, Czech Republic

Introduction The Pálava Protected Landscape Area and Biosphere Reserve in its extended concept, reaching from the limestone hills between Mikulov and Pavlov in the west to the confluence of Morava and Dyje rivers in the east, was the subject of detailed species inventories of terrestrial and aquatic invertebrates (LAŠTÙVKA 1994; ROZKOŠNÝ & VAÒHARA 1995a,b, 1996, 1998, 1999; OPRAVILOVÁ et al. 1999) as well as vertebrates (ØEHÁK et al. 2002) in the 1990’s. However, the environs of the village of Úvaly and the area south of Valtice towards the Austrian boundary were not included at this time. The

87 M. STRAKA et al. area of the Úvalský rybník Pond, hosting very interesting and important plant and animal communities, was overlooked. From the 1980’s onwards at the least, Úvalský rybník Pond (Fig. 1) remained drained and derelict, becoming progressively overgrown with reeds and willow shrubs. In the last two decades, the bulk of the pond has been shallow water, and during the summer it usually dries out, leaving only the deeper main part near the retaining dike. In this period, the occurrence of highly threatened wetland plants and animals has been recorded; e.g. the critically endangered common mare’s-tail (Hippuris vulgaris), water crowsfoot (Batrachium rionii), six frog species and two breeding endangered waterfowl species, the red-crested pochard (Netta rufina) and the garganey (Anas querquedula) (DANIHELKA & GRULICH 2001; D. Horal, pers. comm.). In view of all this, it was registered as a Site of European Community Importance (Evropsky významná lokalita) for the reproduction of the fire-bellied toad (Bombina bombina). However, records of wetland insects from this unique locality were scarce before the surveys. Despite the occurrence of these unique wetland complexes and also despite management recommendations made by nature conservation experts, the pond was inappropriately reconstructed for commercial fish farming in 2005–2006. In 2009 it was filled with turbid water. Littoral vegetation beds developed only sparsely because of reed cutting; high fish densities probably contributed to this scarcity as well. When the pond was reconstructed, a small pool (Fig. 2) was created to the south-west of it to support the amphibian population . The survey, centring upon the aquatic plants and animals of the Úvalský rybník Pond, was performed in 2009 in view of the projected establishment of this site as a Natural Monument. This paper summarizes current knowledge of the aquatic and wetland insects of this locality obtained from the results of an aquatic insect inventory in 2009 and from collections of wetland beetles from the years 2000–2003.

Material and methods In terms of geomorphological classification (DEMEK 1987) the area of Úvalský rybník Pond is part of the Dolnomoravský úval, which is part of the Pannonian lowlands. The area is situated in Valtická pahorkatina about 3 km west of the town of Valtice. The geological substratum is composed of Quaternary fluvial sandy sediments. The soil cover has a high concentration of soluble salt deriving from the Tertiary sediments of the Vienna basin. The Úvalský rybník Pond (48°44′49″N, 16°42′48″E; mapping square 7266) is located in the open valley of the Svodnice, a stream that is part of the River Dyje basin. The altitude of the locality is 200 m a.s.l. Two different bodies of water – the pond and the pool – are currently situated in the area investigated. They were sampled and evaluated individually, apart from adult dragonflies, which were observed, collected and evaluated together over the whole area. In 2009, the vegetation of the pond littoral was dominated by cattails (Typha latifolia and T. angustifolia) and common reed (Phragmites australis), with the bulrush

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Bolboschoenus koshewnikowi in places. The pool littoral was dominated by the cattails Typha angustifolia and T. latifolia. The vegetation of banks was sparse, with some occurrence of Bolboschoenus planiculmis, Cyperus fuscus, Juncus articulatus, J. bufonius, J. compressus, J. inflexus, Mentha verticillata, Plantago uliginosa, Veronica anagalloides and V. catenata. Aquatic insects were collected mostly qualitatively by hand net or kitchen sieve. Horizontal activity traps (BECERRA JURADO et al. 2008) were employed to capture large, actively-swimming species. Adult dragonflies were observed visually and doubtful specimens were caught by sweep net. The locality was visited five times during the 2009 season: 29.iv., 12.vi., 21.viii., 12.ix., and 9.x. Measurements of pH, conductivity, water temperature and dissolved oxygen was performed in the littoral zone of the pool on 9 October 2009 at 5 p.m. using a HACH HQ 40d Multimeter. The very high natural conductivity (1796 µS/cm) of the pool water indicated the occurrence of a considerable amount of soluble salts in the sediments, something typical of the wider surroundings of the locality and which also facilitates the occurrence of the halophilous plants Centaurium pulchellum and Pulicaria dysenterica. The pH was 7.46 and dissolved oxygen stood at 15.34 mg/l, reflected in the rich occurrence of aquatic macrophytes in the pool. Water temperature stood at 17.3°C.

The voucher specimens are deposited in the following collections: JDCD ...... Jakob Damgaard collection, Copenhagen, Denmark (DNA samples) JHTC ...... Jan Helešic collection, Tvrdonice, Czech Republic JSBC ...... Jan Sychra collection, Brno, Czech Republic MMBC ...... Moravian Museum, Brno, Czech Republic MSHC ...... Michal Straka collection, Hustopeèe, Czech Republic PKPC . . . . Petr Kment collection, deposited in National Museum, Prague, Czech Republic

The system and nomenclature for the groups follow: Odonata – ASKEW (1988), Heteroptera – AUKEMA & RIEGER (1995, 1996, 1999, 2001, 2006) and CASSIS & GROSS (1995, 2002), Carabidae – LÖBL & SMETANA (2003), and aquatic Coleoptera – BOUKAL et al. (2007). The categories of threat follow the appropriate chapters of the ‘Red List of Threatened Species in the Czech Republic. Invertebrates’ and its updates: Odonata – DOLNÝ et al. (2007), Heteroptera – KMENT & VILÍMOVÁ (2006), and Coleoptera – BOUKAL et al. (2007), VESELÝ et al. (2006). In the results section, these categories are given in parentheses after the full taxon name, and short comments on the species involved are attached.

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Figs 1–2. 1 – The Úvalský rybník Pond with its underdeveloped littoral macrophyte beds (September 2009). Photo: Jan Sychra. 2 – The newly created pool southwest of the Úvalský rybník Pond, a locality of rich and diverse aquatic insect fauna, including Corixa panzeri Fieber, 1848 new for the Czech Republic (October 2009). Photo: Pavel Sedláèek.

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ODONATA Lestidae Lestes barbarus (Fabricius, 1798) (VU): 12.vi.2009, 5 ♂♂, M. Straka lgt. et det. (MSHC); 21.viii.2009, 4 spec., M. Straka observ. et det. Comment. Lestes barbarus is widely distributed from Europe to India and Mongolia. In Europe it is mainly a Mediterranean species extending northwards to Great Britain and Sweden (ASKEW 1988, DIJKSTRA & LEWINGTON 2006). It probably occurs across whole area of the Czech Republic but its distribution is highly fragmented (DOLNÝ et al. 2007). Since it prefers shallow depressions and meadow pools, it has come to be one of the dragonfly species (together with e.g. Lestes dryas, Aeshna affinis) that is quite tolerant of fluctuating water levels and seasonal habitat drying.

Lestes dryas Kirby, 1890 (VU): 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Comment. A Holarctic species, found in North America as well as in Europe and Asia. In Europe it has a mainly Central and northern European distribution; in the Mediterranean region it is found mainly in the mountains (ASKEW 1988). In the Czech Republic it is distributed in fragmentary fashion across the whole area (DOLNÝ et al. 2007). It is mostly found near small pools and ponds with dense vegetation that typically dry out in the course of summer (DIJKSTRA & LEWINGTON 2006).

Lestes sponsa (Hansemann, 1823): 12.vi.2009, 2 ♂♂ 2 ♀♀, M. Straka lgt. et det. (MSHC).

Lestes virens vestalis Rambur, 1842 (VU): 21.viii.2009, 1 ♂, M. Straka observ. et det. Comment. A Palaearctic species widely distributed in Europe, except in northern areas (ASKEW 1988, DIJKSTRA & LEWINGTON 2006). In the Czech Republic it is most abundant in southern Bohemia and southern Moravia (DOLNÝ et al. 2007).

Lestes viridis (Vander Linden, 1825): 12.vi.2009, 1 ♀, M. Straka lgt. et det. (MSHC).

Sympecma fusca (Vander Linden, 1820) (NT): 29.iv.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC); 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 spec., M. Straka observ. et det. Comment. Widespread throughout southern and Central Europe, extending eastwards to Central Asia (ASKEW 1988, DIJKSTRA & LEWINGTON 2006). Distribution fragmentary over the whole Czech Republic. It prefers warm standing water with rich littoral vegetation (DOLNÝ et al. 2007).

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Coenagrionidae Coenagrion puella (Linnaeus, 1758): 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC).

Enallagma cyathigerum (Charpentier, 1840): 29.iv.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC); 12.vi.2009, 7 spec., M. Straka observ. et det. (2 ♂♂ in MSHC), 21.viii.2009, 10 spec., M. Straka observ. et det.

Ischnura elegans (Vander Linden, 1820): 29.iv.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC); 12.vi.2009, 7 spec., M. Straka observ. et det. (3 ♂♂ in MSHC); 21.viii.2009, 12 spec., M. Straka observ. et det.

Aeshnidae Aeshna affinis Vander Linden, 1823 (VU): 21.viii.2009, 20 spec., M. Straka observ. et det. (1 ♂ in MSHC). Comment. In Europe mostly in the Mediterranean region, further occurs east to Mongolia (ASKEW 1988). Most of the Czech records come from the Moravian lowlands (DOLNÝ et al. 2007). It prefers standing bodies of water that dry up in the course of summer, often overgrown with terrestrial vegetation for the season (DIJKSTRA & LEWINGTON 2006).

Aeshna mixta Latreille, 1805: 21.viii.2009, 1 spec., M. Straka observ. et det.

Anax imperator Leach, 1815: 12.vi.2009, 1 spec., M. Straka observ. et det.; 21.viii.2009, 3 spec., M. Straka observ. et det.

Anax parthenope (Sélys, 1839) (VU): 12.vi.2009, 1 spec., M. Straka observ. et det. Comment. Common in Mediterranean countries; scarce further north, although locally abundant. It has expanded its range since the 1990’s (DIJKSTRA & LEWINGTON 2006); known from the lowlands all over the Czech Republic.

Libellulidae Crocothemis erythraea (Brullé, 1832) (NT): 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Comment. Common and widespread in all of Africa, western Asia and the Mediterranean area. It has steadily expanded its range northwards in recent years (DIJKSTRA & LEWINGTON 2006). In the Czech Republic it can be found in the lowlands, in warm bodies of water with rich littoral vegetation (DOLNÝ et al. 2007).

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Libellula depressa Linnaeus, 1758: 29.iv.2009, 1 spec., M. Straka observ. et det.; 12.vi.2009, 1 spec., M. Straka observ. et det.

Libellula quadrimaculata Linnaeus, 1758: 12.vi.2009, 4 spec., M. Straka observ. et det.

Orthetrum albistylum (Sélys, 1848) (NT): 21.viii.2009, 2 spec., M. Straka observ. et det. Comment. An eastern Palaearctic dragonfly extending from Japan and China through central Asia to the Middle East and Europe (ASKEW 1988). Most of the Czech records are from the Moravian lowlands.

Orthetrum cancellatum (Linnaeus, 1758): 12.vi.2009, 8 spec., M. Straka observ. et det. (1 ♂ in MSHC).

Sympetrum meridionale (Sélys, 1840) (EN): 21.viii.2009, 1 ♂, M. Straka lgt. et det., B. Mocek revid. (MSHC). Comment. In Europe occurs mainly in the Mediterranean area but often migrates elsewhere. In the north it reaches Belgium, southern Germany, Austria and the Czech Republic. Eastwards it reaches as far as Mongolia and southern Siberia. It prefers shallow, still water with rich aquatic vegetation. It is rare in the Czech Republic and can be found only in the warmest lowlands (DOLNÝ et al. 2007).

Sympetrum sanguineum (Müller, 1764): 12.vi.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC); 21.viii.2009, 30 spec., M. Straka observ. et det.

Sympetrum vulgatum (Linnaeus, 1758): 21.viii.2009, 10 spec., M. Straka observ. et det. (3 ♂♂ in MSHC).

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HEMIPTERA: HETEROPTERA Nepidae Nepa cinerea Linnaeus, 1758: Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Ranatra (Ranatra) linearis (Linnaeus, 1758): Pond, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Corixidae Callicorixa praeusta praeusta (Fieber, 1848): Pool, 12.ix.2009, 1 ♂ 1 ♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Corixa affinis Leach, 1817 (CR): Pool, 21.viii.2009, 1 ♂, M. Straka lgt. et det., P. Kment revid. (MSHC). Comment. This species is widely distributed in western, Central, southern and eastern Europe, north Africa, the Near East and Central Asia (JANSSON 1995, PROTIÆ 1998, DAMGAARD 2008). Most of the European records are concentrated in coastal areas (BOSMANS 1982, JANSSON 1986, AUKEMA et al. 2002, WACHMANN et al. 2006), while in Central Europe it maintains an isolated centre of distribution in the Pannonian lowlands (e.g. JANSSON 1986; BRTEK 1953, BAKONYI 1979, BAKONYI & VÁSÁRHELYI 1987, MELBER et al. 1991, BAKONYI et al. 2002, ILIE 2006, RABITSCH 2007).To date there are only two reliable records from the Czech Republic, both from southern Moravia. TEYROVSKÝ (1930) reported a single male from a pool under a railway embankment near Sokolnice (6866) and KMENT (1999) collected a female in Nesyt Pond in Sedlec (7266). The remaining Moravian (ADÁMEK 1976, ADÁMEK et al. 2007) and even Bohemian records (ADÁMEK et al. 2004) are based on misidentifications (Z. Adámek, pers. comm.). All the Moravian records are associated with further localities in southern Slovakia (Danube lowlands – BRTEK 1953; Záhorská nížina lowlands – coll. Moravian Museum, Brno, unpubl.), south-eastern Austria (environs of Neusiedlersee Lake and northern Burgenland – ADLBAUER & HEISS 1980, MELBER et al. 1991, RABITSCH 2007), and the adjacent part of Hungary (BAKONYI et al. 2002). Corixa affinis prefers waters with higher conductivity and salinity, often together with C. panzeri (e.g., HOBERLANDT 1952, JOSIFOV 1961, LANSBURY 1965, BOSMANS 1982, TULLY et al. 1991, SALA & BOIX 2005, KMENT 2006, MARTINOY et al. 2006). According to SAVAGE (1989), it is a rare species in the British Isles, occurring in ponds and pools with a medium content of organic matter on the bottom, a conductivity of 100–10,000 μS/cm, and a pH higher than 6, at altitudes below 300 m a.s.l. In Germany, it has mostly been found in brackish or acid stagnant waters with rich vegetation or in dikes with slowly running water (WACHMANN et al. 2006). All the Moravian records come from areas situated on mineral-rich Tertiary marine sediments where salt-marshes develop. Since it is so tightly associated with endangered salty habitats, this species is considered critically endangered in the Czech Republic (KMENT & VILÍMOVÁ 2006).

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Corixa panzeri Fieber, 1848: Pool, 21.viii.2009, 1 ♂, M. Straka lgt. et det., P. Kment revid. (MSHC); 12.ix.2009, 14 ♂♂ 7 ♀♀ (1 ♂ newly hatched), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JDCD, JSBC, MMBC, MSHC, PKPC). Comment. This species is distributed in western, Central, southern and eastern Europe (Austria, Belgium, Bulgaria, Croatia, Denmark, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Latvia, Liechtenstein, Macedonia, the Netherlands, Norway, Poland, Portugal, Romania, Russia (South European Territory), Slovakia, Spain, Sweden, Ukraine), North Africa (Algeria, Morocco, Tunisia), the Near East and Central Asia (Azerbaijan, Georgia, Iran, Tadjikistan, Turkey) (JANSSON 1986, 1995; PROTIÆ 1998; DOLMEN 2004; for Croatia – P. Kment, unpubl.). Although occurrence in Slovakia is listed by JANSSON (1995), there is no published record from there so far; however, the species has been collected in southern Slovakia by P. Štys (pers. comm.). Like those for C. affinis, most of the European records for C. panzeri concentrate in coastal areas, fading away inland and usually based on single specimens (e.g. WRÓBLEWSKI 1980, BOSMANS 1982, JANSSON 1986, DETTINGER-KLEMM & SONDERMANN 1997, AUKEMA et al. 2002, WACHMANN et al. 2006). In the landlocked parts of Europe, it occurs regularly only in the Pannonian lowlands, with records to date from north-eastern Austria (Neusiedlersee Lake – RABITSCH & ZETTEL 2000, RABITSCH 2007), and Hungary (e.g., JANNSON 1986; BAKONYI 1979, BÁKONY & VÁSÁRHELY 1987, BODA 2008). According to WRÓBLEWSKI (1980) it is rare in Poland. A salinity-tolerant species, it inhabits stagnant waters of various kinds, usually smaller, but with enough aquatic vegetation and sufficient water (at least 0.5 m deep). In Germany, WACHMANN et al. (2006) regarded it as common in saline, oligotrophic or acid stagnant waters with adequate aquatic vegetation, also being found in slowly-running waters. DETTINGER- KLEMM & SONDERMANN (1997) found it in a periodically inundated meadow in Hesse (north-western Germany). JACQUEMIN (2006) collected it from seven different natural brackish localities in Lorraine, and considers the species halophilous. However, he has never found it in the industrial saline waters of Lorraine and Alsace (France). In Belgium, it prefers the dune and polder areas near the coast, with waters naturally rich in nutrients and with abundant submerged vegetation (BOSMANS 1982). According to SAVAGE (1989), it is a rare species in the British Isles, occurring in ponds and pools with medium content of organic matter on the bottom, conductivity ranging from <100 – 10,000 μS/cm, and pH both lower and higher than 6, always occurring at altitudes below 300 m a.s.l. TULLY et al. (1991) observed positive correlations between the occurrence of C. panzeri and (i) high percentage of vegetation cover, (ii) fine bottom substrates, and (iii) increasing concentration of K+ ions. It was also a species typical of the temporary ponds in the study (TULLY et al. 1991). It has also been found in temporary ponds by BOIX et al. (2001; breeding) and SALA & BOIX (2005), as well as in man-made irrigation pools (1069–2595 μS/cm) (ABELLÁN et al. 2006) in the Iberian Peninsula. According to BÁKONY & VÁSÁRHELYI (1987), it is characteristic of the Kiskunság National Park (Hungary), living in both standing and flowing waters, especially in those featuring dense aquatic vegetation and a rich fauna. The habitat information on C. panzeri excerpted from the literature and presented above varies substantially, possibly because accidental

Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 95 M. STRAKA et al. findings of migrating specimens distort the picture. However, at least two preferences become apparent: (i) a preference for slightly saline waters (WRÓBLEWSKI 1980, SAVAGE 1989, BOIX et al. 2001, AUKEMA et al. 2002, ABELLÁN et al. 2006, JACQUEMIN 2006, WACHMANN et al. 2006), and (ii) common occurrence in temporary habitats (TULLY et al. 1991, DETTINGER-KLEMM & SONDERMANN 1997, BOIX et al. 2001). It often occurs together with C. affinis (TULLY et al. 1991, BOIX et al. 2001, KMENT 2006). In the absence of any previous records, it is quite possible that C. panzeri is a new arrival in our fauna. However, in the light of our knowledge of its ecology, its future survival in southernmost Moravia is dependent on the presence of suitable habitats, i.e. pools on a mineral-rich substratum, with rich vegetation and no fish. As this type of habitat is undoubtedly endangered by human pressure, we suggest that C. panzeri be included in our red list as a critically endangered species, as it is in the case for C. affinis.

Corixa punctata (Illiger, 1807): Pond, 12.vi.2009, 1 ♀, M. Straka lgt. et det., P. Kment revid. (MSHC). Pool, 21.viii.2009, 5 spec., M. Straka lgt. et det., P. Kment revid. (MSHC); 12.ix.2009, 4 ♂♂ 6 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JDCD, JSBC, PKPC).

Cymatia coleoptrata (Fabricius, 1777): Pool, 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 5 ♂♂ 3 ♀♀ (brachypterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Cymatia rogenhoferi (Fieber, 1864): Pool, 21.viii.2009, 2 ♀♀, M. Straka lgt. et det., P. Kment revid. (MSHC); 12.ix.2009, 24 ♂♂ 11 ♀♀ (6 ♂♂ 1 ♀ newly hatched) 3 larvae (instar 5), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JDCD, JSBC, MSHC, PKPC). Comment. Cymatia rogenhoferi is distributed from Central and southern Europe and northern Africa to India, with the centre of its distributional area in Central Asia (JANSSON 1986, 1995). In Central Europe, it was originally known from rare and scattered records, but seems to have become more common in the last two decades (e.g. KMENT & SMÉKAL 2002), during which it has spread farther towards the north-west, reaching the Netherlands (KELLEHER & VAN DER VELDE 2001), France (ELDER 2002, JACQUEMIN 2006), and England (NAU & BROOKE 2006).

Hesperocorixa linnaei (Fieber, 1848): Pool, 12.ix.2009, 7 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, MSHC, PKPC).

Micronecta (Dichaetonecta) scholtzi (Fieber, 1860): Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 3 ♂♂ (brachypterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Sigara (Sigara) striata (Linnaeus, 1758): Pool, 21.viii.2009, 9 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 16 ♂♂ 19 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JDCD, JSBC, MSHC, PKPC).

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Sigara (Subsigara) falleni (Fieber, 1848): Pool, 12.ix.2009, 5 ♂♂ 2 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Sigara (Vermicorixa) lateralis (Leach, 1817): Pool, 12.ix.2009, 32 ♂♂ 28 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JDCD, JSBC, MSHC, PKPC).

Naucoridae Ilyocoris cimicoides cimicoides (Linnaeus, 1758): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 1 ♂, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (PKPC).

Notonectidae Notonecta (Notonecta) glauca Linnaeus, 1758: Pool, 12.ix.2009, 2 ♂♂ 1 ♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, MSHC, PKPC).

Notonecta (Notonecta) viridis Delcourt, 1909: Pool, 21.viii.2009, 10 spec., M. Straka lgt. et det., P. Kment revid. (MSHC); 12.ix.2009, 5 ♂♂ 3 ♀♀, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Pleidae Plea minutissima minutissima Leach, 1817: Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, numerous adults and larvae, M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (1 spec. in PKPC).

Mesoveliidae Mesovelia furcata Mulsant et Rey, 1852: Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hebridae Hebrus (Hebrus) pusillus pusillus (Fallén, 1807): Pond, 12.vi.2009, 1 ♀, M. Straka lgt. et det., P. Kment revid. (MSHC); 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hebrus (Hebrusella) ruficeps Thomson, 1871: Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Veliidae Microvelia (Microvelia) reticulata (Burmeister, 1835): Pond, 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 2 ♂♂ 4 ♀♀ (apterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

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Gerridae Aquarius paludum paludum (Fabricius, 1794): Pool, 21.viii.2009, 1 spec., M. Straka observ. et det.

Gerris (Gerris) argentatus Schummel, 1832: Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 2 ♀♀ (macropterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (JSBC, PKPC).

Gerris (Gerris) lacustris (Linnaeus, 1758): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Gerris (Gerris) odontogaster (Zetterstedt, 1828): Pool, 12.ix.2009, 1 ♀ (macropterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (PKPC).

Gerris (Gerris) thoracicus Schummel, 1832: Pool, 29.iv.2009, 1 ♀, M. Straka lgt. et det., P. Kment revid. (MSHC); 12.ix.2009, 1 ♀ (macropterous), M. Straka, P. Kment & J. Sychra lgt., P. Kment det. (PKPC).

Miridae Adelphocoris lineolatus (Goeze, 1778): Vegetation around the pool, sweeping, 12.ix.2009, 3 ♀♀, P. Kment lgt. et det. (PKPC). Adelphocoris ticinensis (Meyer-Dür, 1843) (VU): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Comment. Euro-Siberian species; known in the Czech Republic from about a dozen localities in southernmost Moravia and two scattered localities in northern Bohemia (Kokoøínsko PLA and Lužické hory PLA) (STEHLÍK 1973, KMENT et al. 2005). It is known to inhabit wet, swampy and usually warm habitats (SOUTHWOOD & LESTON 1959, WAGNER 1971, RABITSCH 2003). Most Moravian and Slovak localities are wet meadows in warm lowlands (plant associations: Caricetum fuscae, Alopecuretum pratensis, Bromo-Hordeetum murini) (ŠTEPANOVIÈOVÁ 1967, 1991), while the Bohemian locality of Maøenièky is a peat bog surrounded by wet meadow (associations Crepido-Juncetum acutiflori and Lysimachio vulgaris-Filipenduletum) (KMENT et al. 2005). The species is associated with Lathyrus palustris and Lotus uliginosus; adults are also found on Lysimachia vulgaris, Mentha aquatica, and Lythrum salicaria (ELLIS 1939, SOUTHWOOD & LESTON 1959). Additionally, other authors have mentioned Achillea ptarmica (STEHLÍK 1973), Stachys sp. (WAGNER 1971), Gentiana pneumonanthe (STEHLÍK 1973), and Cirsium palustre (RABITSCH 2003) as host plants. The species is categorised as vulnerable in the Czech Republic (KMENT & VILÍMOVÁ 2006).

Lygus pratensis (Linnaeus, 1758): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂ 1 ♀, P. Kment lgt. et det. (PKPC).

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Lygus rugulipennis Poppius, 1911: Vegetation around the pool, sweeping, 12.ix.2009, 2 ♂♂ 4 ♀♀, P. Kment lgt. et det. (PKPC). Notostira elongata (Geoffroy, 1785): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Notostira erratica (Linnaeus, 1758): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Plagiognathus (Plagiognathus) chrysanthemi (Wolff, 1804): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Stenodema (Brachystira) calcarata (Fallén, 1807): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Trigonotylus caelestialium (Kirkaldy, 1902): Vegetation around the pool, sweeping, 12.ix.2009, 2 ♂♂, P. Kment lgt. et det. (PKPC).

Nabidae Nabis (Nabis) ferus (Linnaeus, 1758): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Nabis (Nabis) pseudoferus pseudoferus Remane, 1949: Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂, P. Kment lgt. et det. (PKPC).

Anthocoridae Orius (Heterorius) majusculus (Reuter, 1879): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂ 2 ♀♀, P. Kment lgt. et det. (PKPC). Orius (Orius) niger (Wolff, 1811): Vegetation around the pool, sweeping, 12.ix.2009, 2 ♂♂ 1 ♀, P. Kment lgt. et det. (PKPC).

Lygaeidae Nysius senecionis senecionis (Schilling, 1829): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC).

Cymidae Cymus claviculus (Fallén, 1807): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂ 1 ♀, P. Kment lgt. et det. (PKPC). Cymus melanocephalus Fieber, 1861: Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂ 1 ♀, P. Kment lgt. et det. (PKPC).

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Artheneidae Chilacis typhae (Perris, 1857): Vegetation around the pool, in flower stem of Typha sp., 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC).

Oxycarenidae Oxycarenus (Euoxycarenus) pallens (Herrich-Schaeffer, 1850): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂ 1 ♀, P. Kment lgt. et det. (PKPC).

Piesmatidae Parapiesma quadratum (Fieber, 1844): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀ (macropterous), P. Kment lgt. et det. (PKPC).

Pyrrhocoridae Pyrrhocoris apterus (Linnaeus, 1758): Úvalský rybník Pond, retaining dike, on stem of Tilia sp., 12.ix.2009, thousands of spec., P. Kment observ. et det.

Rhopalidae Corizus hyoscyami hyoscyami (Linnaeus, 1758): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♀, P. Kment lgt. et det. (PKPC). Stictopleurus punctatonervosus (Goeze, 1778): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂, P. Kment lgt. et det. (PKPC).

Pentatomidae Carpocoris (Carpocoris) purpureipennis (DeGeer, 1773): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂, P. Kment lgt. et det. (PKPC). Dolycoris baccarum (Linnaeus, 1758): Vegetation around the pool, sweeping, 12.ix.2009, 1 ♂, P. Kment lgt. et det. (PKPC).

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COLEOPTERA Haliplidae Haliplus (Haliplidius) obliquus (Fabricius, 1787): Pool, 12.ix.2009, 4 ♂♂ 5 ♀♀, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Haliplus (Haliplus) fluviatilis Aubé, 1836: Pond, 21.viii.2009, 4 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 5 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 6 ♂♂ 3 ♀♀, M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (JSBC, MSHC).

Haliplus (Haliplus) furcatus Seidlitz, 1887 (CR): Pond, 29.iv.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Comment. This Euro-Siberian species occurs especially in sun-exposed shallow wetlands and natural pond littorals (HOLMEN 1987). It is very rare in the Czech Republic; all recent records come from Moravia, e.g. from Znojemsko (BOUKAL et al. 2007).

Haliplus (Haliplus) immaculatus Gerhardt, 1877: Pond, 21.viii.2009, 6 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 6 ♂♂ 5 ♀♀, M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (JSBC, MSHC).

Haliplus (Haliplus) ruficollis (DeGeer, 1774): Pond, 29.iv.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC); 21.viii.2009, 4 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 15 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 6 ♂♂ 11 ♀♀, M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (JSBC, MSHC).

Haliplus (Liaphlus) variegatus Sturm, 1834 (CR): Pool, 29.iv.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Comment. The occurrence of this western Palaearctic species is associated with small water bodies or wetlands, such as fens and peat-bogs, with rich growths of Chara and mosses and clear water. In the Czech Republic it has been found in only small populations, in both Bohemia and Moravia. This species is sensitive to habitat degradation (HOLMEN 1987, BOUKAL et al. 2007).

Haliplus (Neohaliplus) lineatocollis (Marsham, 1802): Pool, 12.ix.2009, 2 ♀♀, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Peltodytes caesus (Duftschmid, 1805): Pond, 21.viii.2009, 3 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

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Noteridae Noterus clavicornis (DeGeer, 1774): Pond, 29.iv.2009, 3 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 ♂, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Dytiscidae Acilius sulcatus (Linnaeus, 1758): Pool, 21.viii.2009, 1 spec., M. Straka observ. et det.

Agabus (Gaurodytes) bipustulatus (Linnaeus, 1767): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC), 12.vi.2009, 7 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 4 spec., M. Straka lgt. et det. (MSHC).

Agabus (Agabus) undulatus (Schrank, 1776): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 15 spec., M. Straka lgt. et det. (MSHC).

Agabus (Gaurodytes) unguicularis (Thomson, 1867): Pool, 29.iv.2009, 1 ♀, M. Straka lgt. et det. (MSHC).

Bidessus nasutus Sharp, 1887 (EN): Pool, 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC). Comment. This species has a Pontic distribution, occurring from Austria and the Czech Republic across the Balkan Peninsula, Ukraine and southern Russia to Turkey, Israel, Kazakhstan, and Turkmenistan (NILSSON 2008). Recent records in the Czech Republic are confined to southern Moravia, in a few localities along the River Dyje (BOUKAL et al. 2007), e.g. in the Lednické rybníky Ponds and Slanisko u Nesytu National Nature Reserves (ŠŤASTNÝ et al. 1999), which are situated only about 3.2 km north of the Úvalský rybník Pond. Bidessus nasutus prefers smaller and warmer water bodies with a sandy or clayey bottom (BOUKAL et al. 2007).

Colymbetes fuscus (Linnaeus, 1758): Pond, 12.vi.2009, 14 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 10 spec., M. Straka lgt. et det. (MSHC).

Cybister (Cybister) lateralimarginalis lateralimarginalis (DeGeer, 1774) (CR): Pond, 29.iv.2009, 6 spec., M. Straka observ. et det. (1 ♂ in MSHC). Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC). Comment. A widespread Palaearctic species, distributed from North Africa to Great Britain and southern Sweden, and eastwards to Kashmir and China. In Denmark and southern Sweden it is most frequently collected in small lakes with dense marginal vegetation (NILSSON & HOLMEN 1995). In the Czech Republic it prefers larger bodies of water such as ponds and flooded quarries with dense littoral vegetation (BOUKAL et al. 2007). Recently known from only about four localities in Bohemia (HAMET et al. 2002; J. Hájek, pers. comm.), more common in South Moravia.

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Dytiscus circumflexus Fabricius, 1801 (VU): Pool, 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Comment. Mainly a European species, distributed from North Africa to Great Britain and southern Sweden, eastwards to the Near East and Turkey. In Fennoscandia this species is mainly found in nutrient-rich ponds, ditches and small lakes with clay, silt or sand bottoms and at least some vegetation, situated in open country (NILSSON & HOLMEN 1995). It probably occurs throughout the Czech Republic but is very rare everywhere (BOUKAL et al. 2007).

Dytiscus marginalis marginalis Linnaeus, 1758: Pool, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC).

Graphoderus austriacus (Sturm, 1834): Pond, 12.vi.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Graphoderus cinereus (Linnaeus, 1758): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 ♂, M. Straka lgt. et det. (MSHC).

Graptodytes bilineatus (Sturm, 1835) (NT): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC). Comment. A widespread Palaearctic species known from most of Europe eastwards through Turkey to Mongolia and Yakutsk in eastern Siberia (NILSSON & HOLMEN 1995). It prefers small bodies of water with dense vegetation and a muddy bottom. Recent Czech records are confined to southern Moravia (BOUKAL et al. 2007).

Hydaticus (Hydaticus) seminiger (DeGeer, 1774): Pond, 29.iv.2009, 3 spec., M. Straka lgt. et det. (MSHC).

Hydroglyphus geminus (Fabricius, 1781): Pond, 21.viii.2009, 6 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 5 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 4 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 7 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Hydroporus angustatus Sturm, 1835: Pool, 12.ix.2009, 3 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Hydroporus palustris (Linnaeus, 1761): Pond, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hydroporus planus (Fabricius, 1781): Pool, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hydroporus striola (Gyllenhal, 1826): Pool, 12.ix.2009, 1 ♀ 1 spec. newly hatched, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

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Hydrovatus cuspidatus (Kunze, 1818) (EN): Pond, 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 4 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC). Comment. A western Palaearctic species known from most of Europe except for Great Britain and Scandinavia, from North Africa, and Asia eastwards to Uzbekistan (NILSSON 2008). The centre of its occurrence in the Czech Republic is southern Moravia; it is known from only two localities in Bohemia (HÁJEK et al. 2001, BOUKAL et al. 2007). More localities have been noted in the Bøeclavsko region, e.g. from Boøí les (ŠŤASTNÝ et al. 1999) and from the River Kyjovka in Mikulèice (P. Komzák, pers. comm.). This species largely prefers older, exposed and unmanaged water bodies with densely overgrown littoral zones (HÁJEK et al. 2001).

Hygrotus (Coelambus) confluens (Fabricius, 1787): Pool, 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Hygrotus (Coelambus) impressopunctatus (Schaller, 1783): Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 2 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Hygrotus (Hygrotus) decoratus (Gyllenhal, 1810): Pool, 12.ix.2009, 3 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Hygrotus (Hygrotus) inaequalis (Fabricius, 1777): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hyphydrus ovatus (Linnaeus, 1761): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC).

Ilybius ater (DeGeer, 1774): Pond, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Ilybius fuliginosus fuliginosus (Fabricius, 1792): Pool, 9.x.2009, 1 spec., J. Sychra lgt. et det. (JSBC).

Ilybius subaeneus Erichson, 1837: Pond, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Laccophilus minutus (Linnaeus, 1758): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 3 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 5 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Laccophilus poecilus Klug, 1834 (VU): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

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Comment. This species ranges through most of the Palaearctic region except for the most northern areas. In Scandinavia it is known from brackish water (NILSON & HOLMEN 1995). In the Czech Republic it prefers shallow bodies of water, warm and with dense vegetation. (BOUKAL et al. 2007). In Bohemia very rare, common only in southern Moravia.

Rhantus (Rhantus) consputus (Sturm, 1834): Pond, 12.vi.2009, 8 spec., M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC).

Rhantus (Rhantus) frontalis (Marsham, 1802): Pool, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Rhantus (Rhantus) suturalis (MacLeay, 1825): Pond, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Pool, 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 ♂, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Carabidae Acupalpus (Acupalpus) maculatus (Schaum, 1860) (VU): Under vegetation around the pond, 27.v.2001, 1 ♂, J. Helešic lgt., R. Láska det. (JHTC). Comment. Western Palaearctic species, very rare in Bohemia and Moravia. It prefers non-shaded wetland shores and saline habitats with vegetation in lowlands (HÙRKA 1996).

Acupalpus (Acupalpus) parvulus (Sturm, 1825): Under vegetation around the pond, 21.v.2001, 1 ♂, J. Helešic lgt., R. Láska det. (JHTC); 1.v.2003, 1 ♀, J. Helešic lgt., R. Láska det. (JHTC); 9.v.2003, 2 ♂♂ 5 ♀♀, J. Helešic lgt., R. Láska det. (JHTC). Agonum (Agonum) gisellae Csiki, 1931 (= A. angustatum (Dejean, 1828)): Derelict pond, 18.v.2002, 2 ♂♂, J. Helešic lgt., R. Láska det. (JHTC). Agonum (Agonum) duftschmidi J. Schmidt, 1994: Derelict pond, 1994: 2.vi.2000, 3 ♂♂ 4 ♀♀, J. Helešic lgt., R. Láska det. (JHTC).

Agonum (Agonum) lugens (Duftschmid, 1812): Derelict pond, 2.vi.2000, 1 ♂, J. Helešic lgt. et det. (JHTC); 27.v.2001, 4 ♂♂ 3 ♀♀, J. Helešic lgt. et det. (JHTC); 9.v.2003, 1 ♀, J. Helešic lgt. et det. (JHTC).

Agonum (Agonum) versutum Sturm, 1824: Derelict pond, 1.v.2003, 1 ♂, J. Helešic lgt. et det. (JHTC).

Agonum (Europhilus) fuliginosum (Panzer, 1809): Derelict pond, 27.v.2001, 2 ♀♀, J. Helešic lgt. et det. (JHTC).

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Badister (Baudia) collaris Motschulsky, 1844: Derelict pond, 2.vi.2000, 2 ♂♂, J. Helešic lgt., R. Láska det. (JHTC).

Badister (Baudia) dilatatus Chaudoir, 1837: Derelict pond, 2.vi.2000, 2 ♂♂ 2 ♀♀, J. Helešic lgt., R. Láska det. (JHTC). Bembidion (Diplocampa) assimile Gyllenhal, 1810: Derelict pond, 27.v.2001, 12 spec. J. Helešic lgt., R. Láska det. (JHTC). Bembidion (Diplocampa) fumigatum (Duftschmid, 1812): Derelict pond, 27.v.2001, 2 ♂♂, J. Helešic lgt. et det. (JHTC). Bembidion (Eupetedromus) dentellum (Thunberg, 1787): Derelict pond, 27.v.2001, 2 ♂♂, J. Helešic lgt. et det. (JHTC), 1.v.2003, 1 ♂ 1 ♀, J. Helešic lgt. et det. (JHTC). Bembidion (Emphanes) tenellum tenellum Erichson, 1837: Derelict pond, 2.vi.2000, 1 spec., J. Helešic lgt., R. Láska det. (JHTC). Blethisa multipunctata multipunctata (Linnaeus, 1758) (VU): Derelict pond, 2.vi.2000, 1 ♂ 1 ♀, J. Helešic lgt. et det.(JHTC); 27.v.2001, 2 ♂♂, J. Helešic lgt. et det. (JHTC); 1.v.2003, 2 ♂♂ 4 ♀♀, J. Helešic lgt. et det. (JHTC). Comment. Circumpolar species, only sporadic occurrence in the Czech Republic. Usually found on wet shores covered in rich vegetation (HÙRKA 1996). Chlaenius (Chlaeniellus) nigricornis (Fabricius, 1787): Derelict pond, 2.vi.2000, 1 ♂, J. Helešic lgt. et det. (JHTC), 1.v.2003, 2 ♂♂ 5 ♀♀, J. Helešic lgt. et det. (JHTC). Chlaenius (Chlaeniellus) tristis tristis (Schaller, 1783): Derelict pond, 2.vi.2000, 5 ♂♂ 2 ♀♀, J. Helešic lgt. et det. (JHTC); 27.v.2001, 1 ♂, J. Helešic lgt. et det. (JHTC). Chlaenius (Chlaenites) spoliatus spoliatus (P. Rossi, 1792): Derelict pond, 27.v.2001, 1 ♀, J. Helešic lgt. et det. (JHTC). Oodes gracilis A. Villa et G. B. Villa, 1833 (VU): Derelict pond, 27.v.2001, 1 ♂, J. Helešic lgt., R. Láska det. (JHTC). Comment. This western Palaearctic species is rare in both Bohemia and Moravia. It prefers naturally preserved non-shaded shores and wetlands and it is considered as a halophilous species (HÙRKA 1996).

Pterostichus (Argutor) cursor (Dejean, 1828) (EN): Derelict pond, 2.vi.2000, 5 ♂♂ 7 ♀♀, J. Helešic lgt. et det. (JHTC); 27.v.2001, 3 ♂♂ 8 ♀♀, J. Helešic lgt. et det. (JHTC). Comment. This halophilous species is known from the south-eastern part of Central Europe and from southern Europe. In the Czech Republic it has been recorded only from Moravia, where it is very rare and local. It prefers bodies of water with very wet, vegetation-covered shores, wetlands and saline habitats in lowlands (HÙRKA 1996). Before the fishpond was brought back into commercial use, the study locality hosted numerous population of this species.

106 Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 Odonata, Heteroptera, Coleoptera in the proposed Úvalský rybník Nature Monument

Pterostichus (Melanius) elongatus elongatus (Duftschmid, 1812) (VU): Derelict pond, 2.vi.2000, 1 ♀, J. Helešic lgt., R. Láska det. (JHTC); 27.v.2001, 7 ♂♂ 2 ♀♀, J. Helešic lgt., R. Láska det. (JHTC). Comment. This species is distributed from north-western Africa to southern Europe and the south-eastern part of central Europe. In the Czech Republic it is very rare and local with only few localities in southern Moravia, e.g. Bøeclav – Kanèí obora (Èudan, pers. comm.), Rakvice–Trkmanský dvùr (J. Helešic, pers. observ.). It occurs on decaying vegetation in wetlands and on saline marshland shores (HÙRKA 1996).

Stenolophus (Stenolophus) skrimshiranus Stephens, 1828: Derelict pond, 2.vi.2000, 1 ♂ 1 ♀, J. Helešic lgt. et det. (JHTC).

Helophoridae Helophorus (Rhopalohelophorus) granularis (Linnaeus, 1761): Pool, 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Helophorus (Rhopalohelophorus) minutus Fabricius, 1775: Pool, 29.iv.2009, 6 spec., M. Straka lgt. et det. (MSHC).

Hydrophilidae Anacaena limbata (Fabricius, 1792): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 3 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 3 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Berosus (Enoplurus) frontifoveatus Kuwert, 1888 (VU): Pool, 12.ix.2009, 3 ♂♂ 5 ♀♀, M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC). Comment. A Palaearctic species which is distributed from the Netherlands to Central Asia. It prefers shallow, warm bodies of water with dense vegetation. Sometimes considered as halophilous (HEBAUER & KLAUSNITZER 2000). In the Czech Republic, it is common in Moravia and southern Bohemia (BOUKAL et al. 2007).

Berosus (Berosus) signaticollis (Charpentier, 1825): Pond, 29.iv.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 2 ♂♂, M. Straka lgt. et det. (MSHC). Coelostoma orbiculare (Fabricius, 1775): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Cymbiodyta marginella (Fabricius, 1792): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (MSHC).

Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 107 M. STRAKA et al.

Enochrus (Enochrus) melanocephalus (Olivier, 1792): Pool, 12.ix.2009, 2 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Enochrus (Lumetus) bicolor (Fabricius, 1792): Pool, 29.iv.2009, 6 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 2 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Enochrus (Lumetus) quadripunctatus (Herbst, 1797): Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.vi.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Enochrus (Lumetus) testaceus (Fabricius, 1801): Pond, 21.viii.2009, 3 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Enochrus (Methydrus) coarctatus (Gredler, 1863): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Helochares obscurus (O. F. Müller, 1776): Pond, 29.iv.2009, 3 spec., M. Straka lgt. et det. (MSHC); 9.x.2009, 1 ♀, J. Sychra lgt. et det. (JSBC). Pool, 29.iv.2009, 14 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 10 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC, MSHC).

Hydrobius fuscipes (Linnaeus, 1758): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC).

Hydrochara caraboides (Linnaeus, 1758): Pond, 21.viii.2009, 8 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Hydrochara flavipes (Steven, 1808): Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 2 spec., M. Straka, P. Kment & J. Sychra lgt., J. Sychra det. (JSBC).

Hydrophilus atterimus (Eschscholtz, 1822) (CR): Pond, 29.iv.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Comment. A Palaearctic species distributed from northern and Central Europe to Siberia and areas around the Caspian Sea. It mostly occurs in larger ponds with broad, densely overgrown littoral zones (HEBAUER & KLAUSNITZER 2000). Most recent records come from southern Moravia. It is sensitive to habitat disturbance since the larvae depend on a succession of differently sized snails to complete their development (BOUKAL et al. 2007).

Laccobius (Dimorpholaccobius) bipunctatus (Fabricius, 1775): Pond, 21.viii.2009, 1 ♀, M. Straka lgt. et det. (MSHC).

108 Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 Odonata, Heteroptera, Coleoptera in the proposed Úvalský rybník Nature Monument

Laccobius (Laccobius) minutus (Linnaeus, 1758): Pond, 29.iv.2009, 2 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 3 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 ♂ 1 ♀, M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (MSHC).

Limnoxenus niger (Gmelin, 1790) (NT): Pond, 29.iv.2009, 1 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Comment. Central and southern Europe, widespread in the Mediterranean (from Spain to Greece), eastwards to Siberia. It lives in stagnant eutrophic waters rich in aquatic vegetation (HANSEN 1987). Locally common in central and southern Moravia but very rare in Bohemia (BOUKAL et al. 2007).

Hydraenidae Hydraena (Hydraena) paganettii Ganglbauer, 1901 (CR): Pool, 29.iv.2009, 6 spec., M. Straka lgt. et det. (MSHC). Comment. A Ponto-Pannonian species, distributed from Central and south-eastern Europe to Turkey, Azerbaijan, Lebanon and Israel (HANSEN 1998, JÄCH 2004). In recent times, the only Czech records have been from lowland localities in southern Moravia (BOUKAL et al. 2007).

Limnebius atomus (Duftschmid, 1805): Pond, 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 4 spec., M. Straka lgt. et det. (MSHC), 21.viii.2009, 1 ♂, M. Straka lgt. et det. (MSHC).

Limnebius papposus Mulsant, 1844: Pond, 21.viii.2009, 2 ♀♀, M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 4 ♀♀, M. Straka lgt. et det. (MSHC), 21.viii.2009, 1 ♂, M. Straka lgt. et det. (MSHC).

Ochthebius (Asiobates) minimus (Fabricius, 1792): Pond, 21.viii.2009, 1 ♀, M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Ochthebius (Ochthebius) pusillus Stephens, 1835: Pool, 29.iv.2009, 6 spec., M. Straka lgt. et det. (MSHC), 21.viii.2009, 2 spec., M. Straka lgt. et det. (MSHC).

Ochthebius (Ochthebius) bernhardi Jäch & Delgado 2008 (VU): Pond, 21.viii.2009, 1 spec., M. Straka lgt. et det. (MSHC). Pool, 29.iv.2009, 6 spec., M. Straka lgt. et det. (MSHC); 21.viii.2009, 26 spec., M. Straka lgt. et det. (MSHC); 12.ix.2009, 1 ♀, M. Straka, P. Kment & J. Sychra lgt., M. Straka det. (JSBC). Comment. JÄCH (1992) revised the Ochthebius marinus group and concluded that the species known as O. viridis Peyron, 1858 represents a species complex consisting of at least two species, referred to as O. viridis sp. 1 and O. viridis sp. 2 (JÄCH 1992). BOUKAL et al. (2007) stated that the populations of O. viridis from the Czech Republic belong to the more widespread species that also occurs inland (= O. viridis sp. 1 sensu JÄCH 1992).

Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 109 M. STRAKA et al.

Recent revision (JÄCH & DELGADO 2008) of the O. viridis species complex corrected the earlier view and showed that all Central European populations of the species complex belong to a species with Ponto-Mediterranean distribution, described as new in that paper: O. bernhardi Jäch et Delgado, 2008. The species is confined in the Czech Republic to warm and shallow drains, pools, and small lakes in southern Moravia (BOUKAL et al. 2007). We keep the same conservation status (vulnerable) for O. bernhardi as for “O. viridis” (BOUKAL et al. 2007). It is very unlikely that true O. viridis viridis Peyron, 1858 occurs in the Czech Republic; nevertheless, all previous records of the O. viridis species complex should be revised.

Scirtidae Cyphon laevipennis Tournier, 1868: Pool, 29.iv.2009, 1 ♂, M. Straka lgt. et det. (MSHC), 12.vi.2009, 5 spec., M. Straka lgt. et det. (MSHC).

Dryopidae Dryops similaris Bollow, 1936 (EN): Pool, 12.vi.2009, 1 ♂, M. Straka lgt. et det. (MSHC). Comment. A Palaearctic species, widely distributed in Europe. In Central Europe it prefers shallow bodies of water in the lowlands, with rich aquatic vegetation (WIÊŹLAK 1986, BOUKAL et al. 2007). In recent times, the only Czech records come from a few localities in southern Moravia.

Heteroceridae Heterocerus fenestratus (Thunberg, 1784): Pond, 21.viii.2009, 6 spec., M. Straka lgt. et det. (MSHC).

Discussion The survey of 2000–2003 and especially that of 2009 recorded 162 taxa of dragonflies (21), true bugs (50) and wetland beetles (91) at the study site. Most of the recorded taxa belong to common euryoecious species (e.g. Ischnura elegans, Sigara falleni, S. striata, Notonecta glauca, Noterus clavicornis, Agabus bipustulatus or Hydrobius fuscipes) or inhabitants of warmer lowland wetlands with rich vegetation (e.g. Aeshna affinis, Notonecta viridis, Hydrovatus cuspidatus, Rhantus suturalis or Berosus frontifoveatus). The bare clayey bottom of the recently created pool also supported populations of some pioneer species (e.g. Libellula depressa, Cymatia rogenhoferi, Bidessus nasutus, Dytiscus circumflexus or Hygrotus confluens). The most interesting records, however, were the Atlanto-Mediterranean species predominantly distributed along the coasts of Europe with their rare occurrence in Central Europe limited to the Pannonian lowland (Corixa affinis, C. panzeri, Ochthebius bernhardi). These species

110 Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 Odonata, Heteroptera, Coleoptera in the proposed Úvalský rybník Nature Monument often prefer sites with higher conductivity and salinity, which is consistent with the measured conductivity value of the pool (1796 µS/cm). Altogether 30 of the recorded species are included in the Red List of threatened species in the Czech Republic or its updates. Six species are considered critically endangered: Corixa affinis, Haliplus furcatus, H. variegatus, Cybister lateralimarginalis, Hydrophilus atterimus and Hydraena paganettii; five species are categorised as endangered: Sympetrum meridionale, Bidessus nasutus, Hydrovatus cuspidatus, Pterostichus cursor and Dryops similaris; fourteen as vulnerable and five as near threatened. For the water boatman Corixa panzeri, the species here first recorded for the Czech Republic, we suggest its inclusion in the critically endangered category, i.e. the same status as C. affinis, alongside which it often occurs. Although the area of the pond is many times larger that of the recently created pool, the 2009 survey revealed that, in comparison with the pool, only about three-quarters of the aquatic species appeared there. The reason is clear. The commercial restoration of the pond in 2006 was to enable intensive fish farming. In 2009 the pond had an obviously dense stock of fish, including large carp, and the littoral macrophyte beds were only sparsely developed. Nevertheless, some rare species were found even in the degraded littoral zones (e.g. Haliplus furcatus, Cybister lateralimarginalis or Hydrophilus atterimus), evidence of the great ecological potential of the pond. However, this site will be further threatened by such inappropriate fish farming and associated progressive habitat degradation. For the conservation of populations of rare species, today probably only residual from the period previous to the restoration of fish farming (DANIHELKA & GRULICH 2001; D. Horal, pers. comm.), a change in pond management practice will be necessary, including (i) either reduction of the fish stock or use of the pond merely to raise carp fry; (ii) prohibition of manuring and intensive fish feeding; (iii) avoiding mass cutting of macrophyte beds; and (iv) periodical partial drainage in summer. Suitable management is necessary to preserve the attractiveness of the pool for rare aquatic insects. It is important to avoid the introduction of fish and to prevent massive invasion of shade-inducing plants such as reeds or willows. Periodical restoration by heavy machinery and the eventual creation of new pools will become necessary. However, the management of the pool must be congruent with the management of the surrounding meadowland, which hosts populations of endangered halophilous plants. Further surveys of the locality, targeted mainly on wetland insects, are also to be recommended; the pool and its surroundings have great ecological potential, especially for halophilous species. The proposed Úvalský rybník Nature Monument is a very valuable natural wetland locality of supraregional importance. Its geographical position on the south-eastern border of the Czech Republic, in the Pannonian lowland and within reach of the larger saline marshlands in the surroundings of the Neusiedlersee Lake in Austria make this locality a very important refuge for lowland, wetland, halophilous insect fauna far more common in southern Moravia in past times. Moreover, lowland, shallow, sun-exposed waters that are only lightly stocked with fish are very rare today, a trend associated with intensification of fish-farming and reclamation of small lowland wetlands in the second

Acta Mus. Moraviae, Sci. Biol. (Brno), 94, 2009 111 M. STRAKA et al. half of the 20th century. The pool at the Úvalský rybník pond, unique for its uncommon abiotic parameters, is a good example of how sensitive management may come to enable threatened halophilous species of aquatic insects to survive in South Moravian conditions. Species inventories and subsequent appropriate management of similar localities in the salt-marsh areas of southern Moravia are necessary for the survival of some of these species, such as Corixa affinis, C. panzeri and Ochthebius bernhardi.

Acknowledgements We are indebted to David Horal (Agency for Nature Conservation and Landscape Protection of the ÈR, Brno) and Jiøí Danihelka (Masaryk University, Brno) for supplying invaluable information. Jiøí Hájek, Martin Fikáèek (National Museum, Prague), Miroslav Papáèek (University of South Bohemia, Èeské Budìjovice), and David Boukal (Institute of Entomology, Czech Academy of Science, Èeské Budìjovice) kindly commented on the manuscript. The investigation was initiated and supported by the Department of the Environment of the South Moravia Region and also by a grant from the Ministry of Culture of the Czech Republic MK00002327201 to the National Museum, Prague, and from the Ministry of Education of the Czech Republic MŠMT ÈR 0021622416 to Masaryk University, Brno.

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