Bulletin 4, a Description of the Fossil Fish Remains of the Cretaceous, Eocene and Miocene Formations
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Vertebral Morphology, Dentition, Age, Growth, and Ecology of the Large Lamniform Shark Cardabiodon Ricki
Vertebral morphology, dentition, age, growth, and ecology of the large lamniform shark Cardabiodon ricki MICHAEL G. NEWBREY, MIKAEL SIVERSSON, TODD D. COOK, ALLISON M. FOTHERINGHAM, and REBECCA L. SANCHEZ Newbrey, M.G., Siversson, M., Cook, T.D., Fotheringham, A.M., and Sanchez, R.L. 2015. Vertebral morphology, denti- tion, age, growth, and ecology of the large lamniform shark Cardabiodon ricki. Acta Palaeontologica Polonica 60 (4): 877–897. Cardabiodon ricki and Cardabiodon venator were large lamniform sharks with a patchy but global distribution in the Cenomanian and Turonian. Their teeth are generally rare and skeletal elements are less common. The centra of Cardabiodon ricki can be distinguished from those of other lamniforms by their unique combination of characteristics: medium length, round articulating outline with a very thick corpus calcareum, a corpus calcareum with a laterally flat rim, robust radial lamellae, thick radial lamellae that occur in low density, concentric lamellae absent, small circular or subovate pores concentrated next to each corpus calcareum, and papillose circular ridges on the surface of the corpus calcareum. The large diameter and robustness of the centra of two examined specimens suggest that Cardabiodon was large, had a rigid vertebral column, and was a fast swimmer. The sectioned corpora calcarea show both individuals deposited 13 bands (assumed to represent annual increments) after the birth ring. The identification of the birth ring is supported in the holotype of Cardabiodon ricki as the back-calculated tooth size at age 0 is nearly equal to the size of the smallest known isolated tooth of this species. The birth ring size (5–6.6 mm radial distance [RD]) overlaps with that of Archaeolamna kopingensis (5.4 mm RD) and the range of variation of Cretoxyrhina mantelli (6–11.6 mm RD) from the Smoky Hill Chalk, Niobrara Formation. -
Symmoriiform Sharks from the Pennsylvanian of Nebraska
Acta Geologica Polonica, Vol. 68 (2018), No. 3, pp. 391–401 DOI: 10.1515/agp-2018-0009 Symmoriiform sharks from the Pennsylvanian of Nebraska MICHAŁ GINTER University of Warsaw, Faculty of Geology, Żwirki i Wigury 93, PL-02-089 Warsaw, Poland. E-mail: [email protected] ABSTRACT: Ginter, M. 2018. Symmoriiform sharks from the Pennsylvanian of Nebraska. Acta Geologica Polonica, 68 (3), 391–401. Warszawa. The Indian Cave Sandstone (Upper Pennsylvanian, Gzhelian) from the area of Peru, Nebraska, USA, has yielded numerous isolated chondrichthyan remains and among them teeth and dermal denticles of the Symmoriiformes Zangerl, 1981. Two tooth-based taxa were identified: a falcatid Denaea saltsmani Ginter and Hansen, 2010, and a new species of Stethacanthus Newberry, 1889, S. concavus sp. nov. In addition, there occur a few long, monocuspid tooth-like denticles, similar to those observed in Cobelodus Zangerl, 1973, probably represent- ing the head cover or the spine-brush complex. A review of the available information on the fossil record of Symmoriiformes has revealed that the group existed from the Late Devonian (Famennian) till the end of the Middle Permian (Capitanian). Key words: Symmoriiformes; Microfossils; Carboniferous; Indian Cave Sandstone; USA Midcontinent. INTRODUCTION size and shape is concerned [compare the thick me- dian cusp, almost a centimetre long, in Stethacanthus The Symmoriiformes (Symmoriida sensu Zan- neilsoni (Traquair, 1898), and the minute, 0.5 mm gerl 1981) are a group of Palaeozoic cladodont sharks wide, multicuspid, comb-like tooth of Denaea wangi sharing several common characters: relatively short Wang, Jin and Wang, 2004; Ginter et al. 2010, figs skulls, large eyes, terminal mouth, epicercal but ex- 58A–C and 61, respectively]. -
OFR21 a Guide to Fossil Sharks, Skates, and Rays from The
STATE OF DELAWARE UNIVERSITY OF DELAWARE DELAWARE GEOLOGICAL SURVEY OPEN FILE REPORT No. 21 A GUIDE TO FOSSIL SHARKS J SKATES J AND RAYS FROM THE CHESAPEAKE ANU DELAWARE CANAL AREA) DELAWARE BY EDWARD M. LAUGINIGER AND EUGENE F. HARTSTEIN NEWARK) DELAWARE MAY 1983 Reprinted 6-95 FOREWORD The authors of this paper are serious avocational students of paleontology. We are pleased to present their work on vertebrate fossils found in Delaware, a subject that has not before been adequately investigated. Edward M. Lauginiger of Wilmington, Delaware teaches biology at Academy Park High School in Sharon Hill, Pennsyl vania. He is especially interested in fossils from the Cretaceous. Eugene F. Hartstein, also of Wilmington, is a chemical engineer with a particular interest in echinoderm and vertebrate fossils. Their combined efforts on this study total 13 years. They have pursued the subject in New Jersey, Maryland, and Texas as well as in Delaware. Both authors are members of the Mid-America Paleontology Society, the Delaware Valley Paleontology Society, and the Delaware Mineralogical Society. We believe that Messrs. Lauginiger and Hartstein have made a significant technical contribution that will be of interest to both professional and amateur paleontologists. Robert R. Jordan State Geologist A GUIDE TO FOSSIL SHARKS, SKATES, AND RAYS FROM THE CHESAPEAKE AND DELAWARE CANAL AREA, DELAWARE Edward M. Lauginiger and Eugene F. Hartstein INTRODUCTION In recent years there has been a renewed interest by both amateur and professional paleontologists in the rich upper Cretaceous exposures along the Chesapeake and Delaware Canal, Delaware (Fig. 1). Large quantities of fossil material, mostly clams, oysters, and snails have been collected as a result of this activity. -
Evolution of White and Megatooth Sharks, and Evidence for Early Predation on Seals, Sirenians, and Whales
Vol.5, No.11, 1203-1218 (2013) Natural Science http://dx.doi.org/10.4236/ns.2013.511148 Evolution of white and megatooth sharks, and evidence for early predation on seals, sirenians, and whales Cajus G. Diedrich Paleologic, Petra Bezruce 96, Zdice, Czech Republic; [email protected], www.paleologic.eu Received 6 April 2013; revised 6 May 2013; accepted 13 May 2013 Copyright © 2013 Cajus G. Diedrich. This is an open access article distributed under the Creative Commons Attribution License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited. ABSTRACT ments were generally first attributed to “white shark Carcharodon carcharias (Linné, 1758) ancestors”. Con- The early white shark Carcharodon Smith, 1838 troversy has subsequently arisen whether they should be with the fossil Carcharodon auriculatus (Blain- ascribed to the megatooth shark (“Carcharocles”—he- ville, 1818) and the extinct megatooth shark Oto- rein Otodus), or to the white shark (Carcharodon) line- dus Agassiz, 1843 with species Otodus sokolovi age [1]. This controversy is partly a result of non-sys- (Jaeckel, 1895) were both present in the Euro- tematic excavation of single serrated similar looking pean proto North Sea Basin about 47.8 - 41.3 m.y. teeth from many localities around the world, and from ago (Lutetian, early Middle Eocene), as well as in horizons of different ages. DNA studies have at least the Tethys realm around the Afican-Eurasian resolved the general position of the extant form of Car- shallow marine habitats. Both top predators deve- charodon carcharias, placing it between the Isurus and loped to be polyphyletic, with possible two dif- Lamna genera [2,3], without taking into account a revi- ferent lamnid shark ancestors within the Early sion and including of extinct fossil species such as Oto- Paleocene to Early Eocene timespan with Car- dus. -
Geo-Eco-Trop., 2020, 44, 1: 161-174 Osteology and Phylogenetic Relationships of Gregoriopycnodus Bassanii Gen. Nov., a Pycnodon
Geo-Eco-Trop., 2020, 44, 1: 161-174 Osteology and phylogenetic relationships of Gregoriopycnodus bassanii gen. nov., a pycnodont fish (Pycnodontidae) from the marine Albian (Lower Cretaceous) of Pietraroja (southern Italy) Ostéologie et relations phylogénétiques de Gregoriopycnodus bassanii gen. nov., un poisson pycnodonte (Pycnodontidae) de l’Albien marin (Crétacé inférieur) de Pietraroja (Italie du Sud) Louis TAVERNE 1, Luigi CAPASSO 2 & Maria DEL RE 3 Résumé: L’ostéologie et les relations phylogénétiques de Gregoriopycnodus bassanii gen. nov., un poisson pycnodonte de l’Albien marin (Crétacé inférieur) de l’Italie du Sud, sont étudiées en détails. Ce genre fossile appartient à la famille des Pycnodontidae, comme le montre la présence d’un peniculus branchu sur le pariétal. Gregoriopycnodus diffère des autres genres de la famille par son préfrontal court, en forme de plaque et qui est partiellement soudé au mésethmoïde. Au sein de la famille, la position systématique de Gregoriopycnodus est intermédiaire entre celle de Tepexichthys et Costapycnodus, d’une part, et celle de Proscinetes, d’autre part. Mots-clés: Pycnodontiformes, Pycnodontidae, Gregoriopycnodus bassanii gen. nov., ostéologie, phylogénie, Albien marin, Italie du Sud Abstract: The osteology and the phylogenetic relationships of Gregoriopycnodus bassanii gen. nov., a pycnodont fish from the marine Albian (Lower Cretaceous) of Pietraroja (southern Italy), are studied in details. This fossil genus belongs to the family Pycnodontidae, as shown by the presence of a branched peniculus on the parietal. Gregoriopycnodus differs from the other genera of the family by its short and plate-like prefrontral that is partly fused to the mesethmoid. Within the family, the systematic position of Gregoriopycnodus is intermediate between that of Tepexichthys and Costapycnodus, on the one hand, and that of Proscinetes, on the other hand. -
14. Knochenfische (Osteichthyes) 14. Bony Fishes (Osteichthyes)
62: 143 – 168 29 Dec 2016 © Senckenberg Gesellschaft für Naturforschung, 2016. 14. Knochenfische (Osteichthyes) 14. Bony fishes (Osteichthyes) Martin Licht †, Ilja Kogan 1, Jan Fischer 2 und Stefan Reiss 3 † verstorben — 1 Technische Universität Bergakademie Freiberg, Geologisches Institut, Bereich Paläontologie/Stratigraphie, Bernhard-von- Cotta-Straße 2, 09599 Freiberg, Deutschland und Kazan Federal University, Institute of Geology and Petroleum Technologies, 4/5 Krem- lyovskaya St., 420008 Kazan, Russland; [email protected] — 2 Urweltmuseum GEOSKOP, Burg Lichtenberg (Pfalz), Burgstraße 19, 66871 Thal lichtenberg, Deutschland; [email protected] — 3 Ortweinstraße 10, 50739 Köln, Deutschland; [email protected] Revision accepted 18 July 2016. Published online at www.senckenberg.de/geologica-saxonica on 29 December 2016. Kurzfassung Neun Gattungen von Knochenfischen aus der Gruppe der Actinopterygier können für die sächsische Kreide als gesichert angegeben werden: Anomoeodus, Pycnodus (Pycnodontiformes), Ichthyodectes (Ichthyodectiformes), Osmeroides (Elopiformes), Pachyrhizodus (in- certae sedis), Cimolichthys, Rhynchodercetis, Enchodus (Aulopiformes) und Hoplopteryx (Beryciformes). Diese Fische besetzten unter- schiedliche trophische Nischen vom Spezialisten für hartschalige Nahrung bis zum großen Fischräuber. Eindeutige Sarcopterygier-Reste lassen sich im vorhandenen Sammlungsmaterial nicht nachweisen. Zahlreiche von H.B. Geinitz für isolierte Schuppen und andere Frag- mente vergebene Namen müssen als nomina dubia angesehen werden. -
First Description of a Tooth of the Extinct Giant Shark Carcharocles
First description of a tooth of the extinct giant shark Carcharocles megalodon (Agassiz, 1835) found in the province of Seville (SW Iberian Peninsula) (Otodontidae) Primera descripción de un diente del extinto tiburón gigante Carcharocles megalodon (Agassiz, 1835) encontrado en la provincia de Sevilla (SO de la Península Ibérica) (Otodontidae) José Luis Medina-Gavilán 1, Antonio Toscano 2, Fernando Muñiz 3, Francisco Javier Delgado 4 1. Sociedad de Estudios Ambientales (SOCEAMB) − Perú 4, 41100 Coria del Río, Sevilla (Spain) − [email protected] 2. Departamento de Geodinámica y Paleontología, Facultad de Ciencias Experimentales, Universidad de Huelva − Campus El Carmen, 21071 Huelva (Spain) − [email protected] 3. Departamento de Geodinámica y Paleontología, Facultad de Ciencias Experimentales, Universidad de Huelva − Campus El Carmen, 21071 Huelva (Spain) − [email protected] 4. Usuario de BiodiversidadVirtual.org − Álvarez Quintero 13, 41220 Burguillos, Sevilla (Spain) − [email protected] ABSTRACT: Fossil remains of the extinct giant shark Carcharocles megalodon (Agassiz, 1835) are rare in interior Andalusia (Southern Spain). For the first time, a fossil tooth belonging to this paleospecies is described from material found in the province of Seville (Burguillos). KEY WORDS: Carcharocles megalodon (Agassiz, 1835), megalodon, Otodontidae, fossil, paleontology, Burguillos, Seville, Tortonian. RESUMEN: Los restos del extinto tiburón gigante Carcharocles megalodon (Agassiz, 1835) son raros en el interior de Andalucía (sur de España). Por primera vez, se describe un diente fósil de esta paleoespecie a partir de material hallado en Sevilla (Burguillos). PALABRAS CLAVE: Carcharocles megalodon (Agassiz, 1835), megalodón, Otodontidae, fósil, paleontología, Burguillos, Sevilla, Tortoniense. Introduction Carcharocles megalodon (Agassiz, 1835), the megalodon, is widely recognised as the largest shark that ever lived. -
Investigating Sexual Dimorphism in Ceratopsid Horncores
University of Calgary PRISM: University of Calgary's Digital Repository Graduate Studies The Vault: Electronic Theses and Dissertations 2013-01-25 Investigating Sexual Dimorphism in Ceratopsid Horncores Borkovic, Benjamin Borkovic, B. (2013). Investigating Sexual Dimorphism in Ceratopsid Horncores (Unpublished master's thesis). University of Calgary, Calgary, AB. doi:10.11575/PRISM/26635 http://hdl.handle.net/11023/498 master thesis University of Calgary graduate students retain copyright ownership and moral rights for their thesis. You may use this material in any way that is permitted by the Copyright Act or through licensing that has been assigned to the document. For uses that are not allowable under copyright legislation or licensing, you are required to seek permission. Downloaded from PRISM: https://prism.ucalgary.ca UNIVERSITY OF CALGARY Investigating Sexual Dimorphism in Ceratopsid Horncores by Benjamin Borkovic A THESIS SUBMITTED TO THE FACULTY OF GRADUATE STUDIES IN PARTIAL FULFILMENT OF THE REQUIREMENTS FOR THE DEGREE OF MASTER OF SCIENCE DEPARTMENT OF BIOLOGICAL SCIENCES CALGARY, ALBERTA JANUARY, 2013 © Benjamin Borkovic 2013 Abstract Evidence for sexual dimorphism was investigated in the horncores of two ceratopsid dinosaurs, Triceratops and Centrosaurus apertus. A review of studies of sexual dimorphism in the vertebrate fossil record revealed methods that were selected for use in ceratopsids. Mountain goats, bison, and pronghorn were selected as exemplar taxa for a proof of principle study that tested the selected methods, and informed and guided the investigation of sexual dimorphism in dinosaurs. Skulls of these exemplar taxa were measured in museum collections, and methods of analysing morphological variation were tested for their ability to demonstrate sexual dimorphism in their horns and horncores. -
(Chondrichthyes, Elasmobranchii) from the Upper Cretaceous of South America
A ptychodontid (Chondrichthyes, Elasmobranchii) from the Upper Cretaceous of South America Paulo M. BRITO Departamento de Biologia Animal e Vegetal, Universidade do Estado do Rio de Janeiro, rua São Francisco Xavier, 524, Rio de Janeiro (Brazil) [email protected] Philippe JANVIER UMR 8569 du CNRS, Département Histoire de la Terre, Muséum national d’Histoire naturelle, 8 rue Buffon, F-75231 Paris cedex 05 (France) [email protected] Brito P. M. & Janvier P. 2002. — A ptychodontid (Chondrichthyes, Elasmobranchii) from the Upper Cretaceous of South America. Geodiversitas 24 (4) : 785-790. ABSTRACT An articulated assemblage of chondrichthyan teeth remain, previously described by Larrazet (1886) as selachian dermal scutes from the Carboniferous of New Granada, is re-described as a Ptychodus sp. and turns out to be Cretaceous in age, presumably from the La Luna Formation of Colombia or Venezuela. This Ptychodus species seems to differ from all the other known ptychodon- KEY WORDS tids by its homodontous dentition, tooth morphology and ornamentation, Chondrichthyes, however we prefer not to erect a new taxon here until tooth morphology Ptychodontidae, Upper Cretaceous, morphoclines become better known and help in the understanding of the South America. phylogenetic relationships among chondrichthyans. RÉSUMÉ Un ptychodontidé (Chondrichthyes, Elasmobranchii) du Crétacé supérieur d’Amérique du Sud. Une plaque dentaire de Chondrichthyes, précédemment décrite par Larrazet (1886) comme des plaques dermiques d’un Sélacien du Carbonifère de la Nouvelle -
Upper Cretaceous Chondrichthyes Teeth Record in Phosphorites of the Loma Gorda Formation•
BOLETIN DE CIENCIAS DE LA TIERRA http://www.revistas.unal.edu.co/index.php/rbct Upper Cretaceous chondrichthyes teeth record in phosphorites of the • Loma Gorda formation Alejandro Niño-Garcia, Juan Diego Parra-Mosquera & Peter Anthony Macias-Villarraga Departamento de Geociencias, Facultad de Ciencias Naturales y Exactas, Universidad de Caldas, Manizales, Colombia. [email protected], [email protected], [email protected] Received: April 26th, 2019. Received in revised form: May 17th, 2019. Accepted: June 04th, 2019. Abstract In layers of phosphorites and gray calcareous mudstones of the Loma Gorda Formation, in the vicinity of the municipal seat of Yaguará in Huila department, Colombia, were found fossils teeth of chondrichthyes, these were extracted from the rocks by mechanical means, to be compared with the species in the bibliography in order to indentify them. The species were: Ptychodus mortoni (order Hybodontiformes), were found, Squalicorax falcatus and Cretodus crassidens (order Lamniformes). This finding constitutes the first record of these species in the Colombian territory; which allows to extend its paleogeographic distribution to the northern region of South America, which until now was limited to Africa, Europe, Asia and North America, except for the Ptychodus mortoni that has been described before in Venezuela. Keywords: first record; sharks; upper Cretaceous; fossil teeth; Colombia. Registro de dientes de condrictios del Cretácico Superior en fosforitas de la formación Loma Gorda Resumen En capas de fosforitas y lodolitas calcáreas grises de la Formación Loma Gorda, en cercanías de la cabecera municipal de Yaguará en el departamento del Huila, Colombia, se encontraron dientes fósiles de condrictios; estos fueron extraídos de la roca por medios mecánicos, para ser comparados con las especies encontradas en la bibliografía e identificarlos. -
Dr. Kenshu Shimada: Peer-Reviewed Published Articles [Last Updated 04/26/2019]
Dr. Kenshu Shimada: Peer-Reviewed Published Articles [last updated 04/26/2019] [* = undergraduate student; ** = graduate student] 2019 [105] Shimada, K. In press. A new species and biology of the Late Cretaceous 'blunt-snouted' bony fish, Thryptodus (Actinopterygii: Tselfatiiformes), from the United States. Cretaceous Research. [104] Cronin, T. J.*, and K. Shimada . In press. New anatomical information on the Late Cretaceous bony fish, Micropycnodon kansasensis (Actinopterygii: Pycnodontiformes), from the Niobrara Chalk of western Kansas, U.S.A. Transactions of Kansas Academy of Science. [103] Pimiento, C., J. L. Cantalapiedra, K. Shimada , D. J. Field, and J. B. Smaers. 2019. Evolutionary pathways towards shark gigantism. Evolution, 73(3):588–599. 2018 [102] Johnson-Ransom, E. D.*, E. V. Popov, T. A. Deméré, and K. Shimada. 2018. The Late Cretaceous chimaeroid fish, Ischyodus bifurcatus Case (Chondrichthyes: Holocephali), from California, USA, and its paleobiogeographical significance. Paleontological Research, 2(4):364-372. [101] Guinot, G., S. Adnet, K. Shimada , C. J. Underwood, M. Siversson, D. J. Ward, J. Kriwet, and H. Cappetta. 2018. On the need of providing tooth morphology in descriptions of extant elasmobranch species. Zootaxa, 4461(1):118–126. [100] Jacobs, P. K.*, and K. Shimada . 2018. Ontogenetic growth pattern of the extant smalltooth sandtiger shark, Odontaspis ferox (Lamniformes: Odontaspididae)— application from and to paleontology. Journal of Fossil Research, 51(1):23–29. [99] Guzzo, F.*, and K. Shimada . 2018. A new fossil vertebrate locality of the Jetmore Chalk Member of the Upper Cretaceous Greenhorn Limestone in north-central Kansas, U.S.A. Transactions of the Kansas Academy of Science, 121(1-2):59-68. -
Copyrighted Material
06_250317 part1-3.qxd 12/13/05 7:32 PM Page 15 Phylum Chordata Chordates are placed in the superphylum Deuterostomia. The possible rela- tionships of the chordates and deuterostomes to other metazoans are dis- cussed in Halanych (2004). He restricts the taxon of deuterostomes to the chordates and their proposed immediate sister group, a taxon comprising the hemichordates, echinoderms, and the wormlike Xenoturbella. The phylum Chordata has been used by most recent workers to encompass members of the subphyla Urochordata (tunicates or sea-squirts), Cephalochordata (lancelets), and Craniata (fishes, amphibians, reptiles, birds, and mammals). The Cephalochordata and Craniata form a mono- phyletic group (e.g., Cameron et al., 2000; Halanych, 2004). Much disagree- ment exists concerning the interrelationships and classification of the Chordata, and the inclusion of the urochordates as sister to the cephalochor- dates and craniates is not as broadly held as the sister-group relationship of cephalochordates and craniates (Halanych, 2004). Many excitingCOPYRIGHTED fossil finds in recent years MATERIAL reveal what the first fishes may have looked like, and these finds push the fossil record of fishes back into the early Cambrian, far further back than previously known. There is still much difference of opinion on the phylogenetic position of these new Cambrian species, and many new discoveries and changes in early fish systematics may be expected over the next decade. As noted by Halanych (2004), D.-G. (D.) Shu and collaborators have discovered fossil ascidians (e.g., Cheungkongella), cephalochordate-like yunnanozoans (Haikouella and Yunnanozoon), and jaw- less craniates (Myllokunmingia, and its junior synonym Haikouichthys) over the 15 06_250317 part1-3.qxd 12/13/05 7:32 PM Page 16 16 Fishes of the World last few years that push the origins of these three major taxa at least into the Lower Cambrian (approximately 530–540 million years ago).