DOCSLIB.ORG

New Material of Trematochampsa Oblita (Crocodyliformes

Total Page：16

File Type：pdf, Size：1020Kb

Download full-text PDF Read full-text
New Material of Trematochampsa Oblita (Crocodyliformes a... .x . (21:1: , . .. 5:1 ..\xtvy. .fimhfi... ,1; vi. m. 3 1% .3....s L1 . 3 .3 .: I. , i ‘1‘? IA(- .V. .2. I: .1 33?? ,1. in). 2:... .- .38. :. ..Y . t: .5 in“... It Ham”. 32; 2 5r. z! a . l1 :t»5 3m. .Ibubfii ¢ . ‘ , ,1 1,22% .51.. 3| l 1‘ .I fit: 34$..ka91%.! Liv— :t ‘5 ‘5! .i AH. ‘13. , I; .q .1 . ‘ 7 !. .........u...a..... .1 , . 1.... .134 . a? 33.354. , . 1-K. .3.» .lihfi . .l\l \ l .x . a: .3 4... :7. .. .‘ iii: 4.: .9 5:4: .3 ‘3 a 3...! I... 3.x : O 153...: . v . 123.! 95:. l . ‘1. .’!r&u¢ 13;)3r-1 ‘2; _ : r .l :x L.:x.x.§EkB .943.» “a. a, J7... ... w ‘ ‘I .. 1!... .éasrfgfi >1)» 3: 3.» r3... ‘ $§firfii¢§§z§g§ .. awn—w. ms “001/ This is to certify that the thesis entitled New Material of Trematochampsa oblita (Crocodyliformes, Trematochampsidae) from the Late Cretaceous of Madagascar, with a Review and Cladistic Analysis of the Family Trematochampsidae presented by Erin Lynn Rasmusson has been accepted towards fulfillment of the requirements for Masters degree in Geological Sciences \ Major professor Dateg M03 Zack"- 0-7639 MS U is an Affirmative Action/Equal Opportunity Institution LIBRARY Michigan State University PLACE IN RETURN BOX to remove this checkout from your record. TO AVOID FINE return on or before date due. MAY BE RECALLED with earlier due date if requested. DATE DUE DATE DUE ' DATE DUE inE 5% 2006 6/01 c:/CIRC/Dat90ue.p65-p.15 NEW MATERLAL OF IMEAMTOCHAJMPSA OBLITA (CROCODYLIFORMES, TREb-LATOCHAMPSIDAE) FROM THE LATE CRETACEOUS OF MADAGASCAR, \VITH A REVIEW AND CLADISTIC ANALYSIS OF THE FAMILY TREMATOCHAMPSIDAE By Erin Lynn Rasmusson A THESIS Submitted to Michigan State University in partial fulfillment of the requirements for the degree of MASTER OF SCIENCE Department of Geological Sciences 2002 ABSTRACT NEW MATERIAL OF 'I"Rf5i'\1."lTOCI IAA'IPS/I OBLITA (CROCODYLIFORMES, TREMATOCHAMPSIDAE) FROM THE LATE CRETACEOUS OF MADAGASCAR, WITH A REVIEW AND CLADISTIC ANALYSIS OF THE FAMILY TREMATOCHAMPSIDAE By Erin Lynn Rasmusson Recent study of new material of the trematochampsid crocodyliform T rematochampsa oblita, from the Late Cretaceous of Madagascar, has provided additional data for a Cladistic analysis of the Family Trematochampsidae, an enigmatic group of metasuchian crocodyliforms known fi'om the Cretaceous of Gondwana and the Tertiary of Europe. Among the questions addressed were: (1) Is the Family Trematochampsidae monophyletic? (2) How do trematochampsids relate to other crocodilian groups that have been hypothesized to be closely related, notably peirosaurids? (3) Is the genus T rematochampsa monophyletic with respect to other trematochampsids? and (4) What are the implications for biogeography? For the Cladistic analysis, PAUP*4.0 was employed, using a variety of search algorithms on a data matrix that included thirty unordered characters and thirteen total taxa, including two outgroups (Araripesuchus patagonicus and Notosuchus terrestris), two peirosaurids (Peirosaurus tormini and Lomasuchus palpebrosus), and the nine named trematochampsid species. Strict and Adams consensus trees were obtained with the data, one set including and another excluding the two peirosaurids. A monophyletic Trematochampsidae was recovered in both options, and the peirosaurids, when included, fall within Trematochampsidae. The genus T rematochampsa itself, however, was found to be paraphyletic, with T rematochampsa oblita closer to the European taxa on the consensus tree that does not include the peirosaurids. The two European Tertiary taxa, Bergisuchus dietrichbergi and Iberosuchus macrodon, were found to be sister taxa and to occupy a relatively crownward position in the clade. The results of the analysis support the use of a vicariance biogeography model to explain the distribution of Gondwanan trematochampsids. However, a dispersal model best explains the presence of the Tertiary forms in Europe. ACKNOWLEDGMENTS I would like to thank my advisor, Dr. Mike Gottfried, for all his assistance and guidance throughout this project, as well as my committee members: Dr. Al Holman and Dr. Danita Brandt, for their many insights. Thank you to Dr. Greg Buckley, my honorary committee member, for letting me work on his specimens. There are many people associated with the Mahajanga Basin Project that I’d like to acknowledge. First, Dr. David Krause (Stony Brook University) for his support and generosity; also, the many participants of the Mahajanga Basin expeditions for their efforts in the field; the people of Berivotra; our colleagues at the Université d’Antananarivo; and the Institute for the Conservation of Tropical Environments in Antananarivo. I would not have been able to finish this project without the help of many people. Especially,]ohn and Anita Campeau for their dedicated French translations; Sunny Wang at the MSU Museum and his staff for their computer expertise; the generous petrologists and hydrogeologists for allowing a paleontologist to invade their computer labs; and Dr. Tom V ogel for the last minute use of his digital camera. Finally, I am most grateful for the support of my fellow graduate students, who have become my very good friends, as well as that of my family and Mark. iii TABLE OF CONTENTS LIST OF TABLES . ........................................................................................................... iii LIST OF FIGURES .......................................................................................................... iv KEY TO ABBREVIATIONS ........................................................................................ vi INTRODUCTION ............................................................................................................ 1 FAMILY TREMATOCHAMPSIDAE ,4lzzargarur/Jm‘ minor ......................................................................................................... 8 [fart/rim: rampart ............................................................................................................... 8 I 'Iamadaiz/r/Jm‘ Ie/mu/z' ..................................................................................................... 1 3 anzaloc‘bawpm [aqua/1' ................................................................................................... 16 TIP/7ld/0t‘bd/II/DJZI 0/2/1711 ..................................................................................................... 16 I farm‘/Ju.rje.r//1'210i .............................................................................................................. 1 6 I Err/Izoirz/c‘bm e/ko/to/zlm .................................................................................................. 20 1/26/0511: 12115 7/10L7IN/0” ....................................................................................................... 23 Bergzlrm/mr die/n't'bbergi ................................................................................................... 23 PREVIOUS MATERLAL OF TREAL 4 T OCH/11111534 OBLIT/l ........................ 25 NEW MATERLAL OF TREM4TOCI‘L}MIPSA OBLI F1 .................................... 25 SYSTEIVIATIC PALEONTOLOGY LOCALITY AND HORIZON ............................................................................... 29 DESCRIPTION .......................................................................................................... 29 Denlagl ....................................................................................................................... 29 an/a/ ........................................................................................................................ 35 V eflebm ...................................................................................................................... 35 PHYLOGENETIC ANALYSIS ................................................................................... 39 RESULTS ........................................................................................................................... 41 DISCUSSION OF PHYLOGENY RELATIONSHIPS WITHIN TREMATOCHAMPSIDS ................................ 44 RELATIONSHIPS INCLUDING PEIROSAURIDS ...................................... 47 IMPLICATIONS FOR BIOGEOGRAPI-IY ............................................................ 47 CONCLUSION ................................................................................................................ 52 APPENDIX ....................................................................................................................... 54 REFERENCES ................................................................................................................. 58 iv LIST OF TABLES Number Page 1. Classification of the Tremtochampsidae within the Archosauromorpha ................................... 2 2. Data matrix of 13 taxa and 30 characters ........................................................................................ 40 LIST OF FIGURES Number Page 1. Skull of 1m77pctsm/Jm paiagmzzim ........................................................................................................... 4 2. Skull of Lfllllafl/t‘bfli pa/pebmmr .............................................................................................................. 6 3. Select distinguishing characteristics of the Family Trematochampsidae ..................................... 7 4. Distribution of trematochampsid species ......................................................................................... 9 5. Skull of TR’III(110(‘/}all¢>.fa [aqua/i compared with right maxilla of Amalgam/(Mr minor ................. 10 6. Right maxilla of Alizwgai‘m/my
Load more

Recommended publications
  • 8. Archosaur Phylogeny and the Relationships of the Crocodylia
    8. Archosaur phylogeny and the relationships of the Crocodylia MICHAEL J. BENTON Department of Geology, The Queen's University of Belfast, Belfast, UK JAMES M. CLARK* Department of Anatomy, University of Chicago, Chicago, Illinois, USA Abstract The Archosauria include the living crocodilians and birds, as well as the fossil dinosaurs, pterosaurs, and basal 'thecodontians'. Cladograms of the basal archosaurs and of the crocodylomorphs are given in this paper. There are three primitive archosaur groups, the Proterosuchidae, the Erythrosuchidae, and the Proterochampsidae, which fall outside the crown-group (crocodilian line plus bird line), and these have been defined as plesions to a restricted Archosauria by Gauthier. The Early Triassic Euparkeria may also fall outside this crown-group, or it may lie on the bird line. The crown-group of archosaurs divides into the Ornithosuchia (the 'bird line': Orn- ithosuchidae, Lagosuchidae, Pterosauria, Dinosauria) and the Croco- dylotarsi nov. (the 'crocodilian line': Phytosauridae, Crocodylo- morpha, Stagonolepididae, Rauisuchidae, and Poposauridae). The latter three families may form a clade (Pseudosuchia s.str.), or the Poposauridae may pair off with Crocodylomorpha. The Crocodylomorpha includes all crocodilians, as well as crocodi- lian-like Triassic and Jurassic terrestrial forms. The Crocodyliformes include the traditional 'Protosuchia', 'Mesosuchia', and Eusuchia, and they are defined by a large number of synapomorphies, particularly of the braincase and occipital regions. The 'protosuchians' (mainly Early *Present address: Department of Zoology, Storer Hall, University of California, Davis, Cali- fornia, USA. The Phylogeny and Classification of the Tetrapods, Volume 1: Amphibians, Reptiles, Birds (ed. M.J. Benton), Systematics Association Special Volume 35A . pp. 295-338. Clarendon Press, Oxford, 1988.
    [Show full text]
  • A Teleosaurid (Crocodylia, Mesosuchia) from the Toarcian of Madagascar and Its Palaeobiogeographical Significance
    Pal~iont. Z. 55 I 3/4 313-319 Stuttgart, Dezember 1981 I A teleosaurid (Crocodylia, Mesosuchia) from the Toarcian of Madagascar and its palaeobiogeographical significance ERIC BUFFETAUT, GENEVIJ~VE TERMIER & HENRI TERM1ER, Paris* With 2 figures in the text Kurzfassung: Ein Oberkieferfragment eines Krokodiliers aus dem Toarcium von NW-Madagaskar wird als Stenosaurus sp. (Familie Teleosauridae) bestimmt. Es ist der ~ilteste aus Madagaskar bekannte Kro- kodilier und deutet aunistische B eziehungen mit Westeuropa und Siidamerika an. Die Verbreitung der ober- liassischen marinen Krokodilier, die derjenigen des Ammoniten Bouleiceras iihnelt, weist auf die Existenz einer epikontinentalen Seestraf~e vonder Tethys fiber die transerythr~iische Provinz bis zum siidlichen Tell Afrikas; diese Seestral~e bildete eine Verbindung zwischen Westeuropa und Siidamerika. A b s t r a c t : A fragment of the upper jaw of a crocodilian from the Toarcian of NW Madagascar is refer- red to Steneosaurus sp. (family Teleosauridae). It is the earliest known crocodilian from Madagascar. It indi- cates faunal relationships with western Europe and South America. The distribution of late Liassic marine crocodilians, similar to that of the ammonite Bouleiceras, suggests the existence of an epicontinental seaway from the Tethyan region to the southern part of Africa via the Transerythrean Province, which provided a marine connection between western Europe and South America. R ~ s u m ~ : Un fragment de mfichoire sup~rieure de Crocodilien du Toarcien du NW de Madagascar est rapport8 ~Steneosaurus sp. (famille Teleosauridae). C'est le plus ancien Crocodilien connu ~iMadagascar. I1 indique des relations fauniques avec l'Europe occidentale et l'Am&ique du Sud.
    [Show full text]
  • 71St Annual Meeting Society of Vertebrate Paleontology Paris Las Vegas Las Vegas, Nevada, USA November 2 – 5, 2011 SESSION CONCURRENT SESSION CONCURRENT
    ISSN 1937-2809 online Journal of Supplement to the November 2011 Vertebrate Paleontology Vertebrate Society of Vertebrate Paleontology Society of Vertebrate 71st Annual Meeting Paleontology Society of Vertebrate Las Vegas Paris Nevada, USA Las Vegas, November 2 – 5, 2011 Program and Abstracts Society of Vertebrate Paleontology 71st Annual Meeting Program and Abstracts COMMITTEE MEETING ROOM POSTER SESSION/ CONCURRENT CONCURRENT SESSION EXHIBITS SESSION COMMITTEE MEETING ROOMS AUCTION EVENT REGISTRATION, CONCURRENT MERCHANDISE SESSION LOUNGE, EDUCATION & OUTREACH SPEAKER READY COMMITTEE MEETING POSTER SESSION ROOM ROOM SOCIETY OF VERTEBRATE PALEONTOLOGY ABSTRACTS OF PAPERS SEVENTY-FIRST ANNUAL MEETING PARIS LAS VEGAS HOTEL LAS VEGAS, NV, USA NOVEMBER 2–5, 2011 HOST COMMITTEE Stephen Rowland, Co-Chair; Aubrey Bonde, Co-Chair; Joshua Bonde; David Elliott; Lee Hall; Jerry Harris; Andrew Milner; Eric Roberts EXECUTIVE COMMITTEE Philip Currie, President; Blaire Van Valkenburgh, Past President; Catherine Forster, Vice President; Christopher Bell, Secretary; Ted Vlamis, Treasurer; Julia Clarke, Member at Large; Kristina Curry Rogers, Member at Large; Lars Werdelin, Member at Large SYMPOSIUM CONVENORS Roger B.J. Benson, Richard J. Butler, Nadia B. Fröbisch, Hans C.E. Larsson, Mark A. Loewen, Philip D. Mannion, Jim I. Mead, Eric M. Roberts, Scott D. Sampson, Eric D. Scott, Kathleen Springer PROGRAM COMMITTEE Jonathan Bloch, Co-Chair; Anjali Goswami, Co-Chair; Jason Anderson; Paul Barrett; Brian Beatty; Kerin Claeson; Kristina Curry Rogers; Ted Daeschler; David Evans; David Fox; Nadia B. Fröbisch; Christian Kammerer; Johannes Müller; Emily Rayfield; William Sanders; Bruce Shockey; Mary Silcox; Michelle Stocker; Rebecca Terry November 2011—PROGRAM AND ABSTRACTS 1 Members and Friends of the Society of Vertebrate Paleontology, The Host Committee cordially welcomes you to the 71st Annual Meeting of the Society of Vertebrate Paleontology in Las Vegas.
    [Show full text]
  • Baixar Este Arquivo
    DOI 10.5935/0100-929X.20120007 Revista do Instituto Geológico, São Paulo, 33 (2), 13-29, 2012 DESCRIÇÃO DE UM ESPÉCIME JUVENIL DE BAURUSUCHIDAE (CROCODYLIFORMES: MESOEUCROCODYLIA) DO GRUPO BAURU (NEOCRETÁCEO): CONSIDERAÇÕES PRELIMINARES SOBRE ONTOGENIA Caio Fabricio Cezar GEROTO Reinaldo José BERTINI RESUMO Entre os táxons de Crocodyliformes do Grupo Bauru (Neocretáceo), grande quan- tidade de morfótipos, incluindo materiais cranianos e pós-cranianos, vem sendo des- crita em associação a Baurusuchus pachecoi. Porém, a falta de estudos ontogenéticos, como os realizados para Mariliasuchus amarali, levou à atribuição de novos gêneros e espécies aos poucos crânios completos encontrados. A presente contribuição traz a descrição de um espécime juvenil de Baurusuchidae depositado no acervo do Museu de Ciências da Terra no Rio de Janeiro sob o número MCT 1724 - R. Trata-se de um rostro e mandíbula associados e em oclusão, com o lado esquerdo melhor preservado que o direito, e dentição zifodonte extremamente reduzida. O fóssil possui 125,3 mm de comprimento preservado da porção anterior do pré-maxilar até a extremidade pos- terior do dentário; 117,5 mm de comprimento preservado do pré-maxilar aos palatinos e altura lateral de 51,4 mm. Entre as informações de caráter ontogenético identificadas destacam-se: ornamentação suave composta de estrias vermiformes muito espaçadas e largas, linha ventral do maxilar mais reta, dentário levemente inclinado dorsalmente na porção mediana e sínfise mandibular menos vertical que em outros baurussúqui- dos de tamanho maior. A maioria das características rostrais e dentárias, diagnósticas para Baurusuchus pachecoi, foi identificada no exemplar MCT 1724 - R: rostro alto e comprimido lateralmente, além de dentição zifodonte com forte redução dentária, que culmina em quatro dentes pré-maxilares e cinco maxilares.
    [Show full text]
  • Craniofacial Morphology of Simosuchus Clarki (Crocodyliformes: Notosuchia) from the Late Cretaceous of Madagascar
    Society of Vertebrate Paleontology Memoir 10 Journal of Vertebrate Paleontology Volume 30, Supplement to Number 6: 13–98, November 2010 © 2010 by the Society of Vertebrate Paleontology CRANIOFACIAL MORPHOLOGY OF SIMOSUCHUS CLARKI (CROCODYLIFORMES: NOTOSUCHIA) FROM THE LATE CRETACEOUS OF MADAGASCAR NATHAN J. KLEY,*,1 JOSEPH J. W. SERTICH,1 ALAN H. TURNER,1 DAVID W. KRAUSE,1 PATRICK M. O’CONNOR,2 and JUSTIN A. GEORGI3 1Department of Anatomical Sciences, Stony Brook University, Stony Brook, New York, 11794-8081, U.S.A., [email protected]; [email protected]; [email protected]; [email protected]; 2Department of Biomedical Sciences, Ohio University College of Osteopathic Medicine, Athens, Ohio 45701, U.S.A., [email protected]; 3Department of Anatomy, Arizona College of Osteopathic Medicine, Midwestern University, Glendale, Arizona 85308, U.S.A., [email protected] ABSTRACT—Simosuchus clarki is a small, pug-nosed notosuchian crocodyliform from the Late Cretaceous of Madagascar. Originally described on the basis of a single specimen including a remarkably complete and well-preserved skull and lower jaw, S. clarki is now known from five additional specimens that preserve portions of the craniofacial skeleton. Collectively, these six specimens represent all elements of the head skeleton except the stapedes, thus making the craniofacial skeleton of S. clarki one of the best and most completely preserved among all known basal mesoeucrocodylians. In this report, we provide a detailed description of the entire head skeleton of S. clarki, including a portion of the hyobranchial apparatus. The two most complete and well-preserved specimens differ substantially in several size and shape variables (e.g., projections, angulations, and areas of ornamentation), suggestive of sexual dimorphism.
    [Show full text]
  • A New Pissarrachampsinae Specimen from the Bauru Basin, Brazil, Adds
    YCRES104969_proof ■ 29 July 2021 ■ 1/13 Cretaceous Research xxx (xxxx) xxx 55 Contents lists available at ScienceDirect 56 57 58 Cretaceous Research 59 60 journal homepage: www.elsevier.com/locate/CretRes 61 62 63 64 65 1 A new Pissarrachampsinae specimen from the Bauru Basin, Brazil, 66 2 67 3 adds data to the understanding of the Baurusuchidae 68 4 (Mesoeucrocodylia, Notosuchia) distribution in the Late Cretaceous of 69 5 70 6 Q7 South America 71 7 72 8 a, * b, c d Q6 Gustavo Darlim , Ismar de Souza Carvalho , Sandra Aparecida Simionato Tavares , 73 9 Max Cardoso Langer a 74 10 75 11 a Universidade de Sao~ Paulo, Faculdade de Filosofia, Ci^encias e Letras de Ribeirao~ Preto, Laboratorio de Paleontologia, Av. Bandeirantes, 3900, Ribeirao~ Preto, 76 12 SP, Brazil 77 b ^ 13 Q2 Universidade Federal do Rio de Janeiro, Instituto de Geociencias, CCMN, 21.910-200 Av. Athos da Silveira Ramos 273, Rio de Janeiro, RJ, Brazil c Centro de Geoci^encias da Universidade de Coimbra, Portugal 78 14 d Museu de Paleontologia “Prof. Antonio^ Celso de Arruda Campos”, Praça do Centenario, Centro, Monte Alto, SP, Brazil 79 15 80 16 81 17 article info abstract 82 18 83 19 Article history: Baurusuchidae is one of the most diverse notosuchian groups, represented by ten formally described 84 20 Received 7 January 2021 species from the Upper Cretaceous deposits of the Bauru and Neuquen basins, respectively in Brazil and 85 21 Received in revised form Argentina. Among these, recent phylogenetic analyses placed Wargosuchus australis, Campinasuchus 86 18 July 2021 22 dinizi, and Pissarrachampsa sera within Pissarrachampsinae, whereas Baurusuchinae is composed by 87 Accepted in revised form 18 July 2021 23 Aphaurosuchus escharafacies, Aplestosuchus sordidus, Baurusuchus albertoi, Baurusuchus pachecoi, Baur- Available online xxx 88 24 usuchus salgadoensis, and Stratiotosuchus maxhechti.
    [Show full text]
  • A New Species of Coloborhynchus (Pterosauria, Ornithocheiridae) from the Mid- Cretaceous of North Africa
    Accepted Manuscript A new species of Coloborhynchus (Pterosauria, Ornithocheiridae) from the mid- Cretaceous of North Africa Megan L. Jacobs, David M. Martill, Nizar Ibrahim, Nick Longrich PII: S0195-6671(18)30354-9 DOI: https://doi.org/10.1016/j.cretres.2018.10.018 Reference: YCRES 3995 To appear in: Cretaceous Research Received Date: 28 August 2018 Revised Date: 18 October 2018 Accepted Date: 21 October 2018 Please cite this article as: Jacobs, M.L., Martill, D.M., Ibrahim, N., Longrich, N., A new species of Coloborhynchus (Pterosauria, Ornithocheiridae) from the mid-Cretaceous of North Africa, Cretaceous Research (2018), doi: https://doi.org/10.1016/j.cretres.2018.10.018. This is a PDF file of an unedited manuscript that has been accepted for publication. As a service to our customers we are providing this early version of the manuscript. The manuscript will undergo copyediting, typesetting, and review of the resulting proof before it is published in its final form. Please note that during the production process errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. 1 ACCEPTED MANUSCRIPT 1 A new species of Coloborhynchus (Pterosauria, Ornithocheiridae) 2 from the mid-Cretaceous of North Africa 3 Megan L. Jacobs a* , David M. Martill a, Nizar Ibrahim a** , Nick Longrich b 4 a School of Earth and Environmental Sciences, University of Portsmouth, Portsmouth PO1 3QL, UK 5 b Department of Biology and Biochemistry and Milner Centre for Evolution, University of Bath, Bath 6 BA2 7AY, UK 7 *Corresponding author. Email address : [email protected] (M.L.
    [Show full text]
  • I – Introdução
    UNIVERSIDADE ESTADUAL PAULISTA Instituto de Geociências e Ciências Exatas Campus de Rio Claro REVISÃO SISTEMÁTICA E TAXONÔMICA DOS NOTOSUCHIA (METASUCHIA, CROCODYLOMORPHA) Marco Brandalise de Andrade Orientador: Prof. Dr. Reinaldo José Bertini Dissertação de Mestrado elaborada junto ao Programa de Pós-Graduação em Geologia - Área de concentração em Geologia Regional, para a obtenção do título de Mestre em Geociências Rio Claro (SP) 2005 Comissão Examinadora _____________________________________ Prof. Dr. Reinaldo José Bertini _____________________________________ Prof. Dr. Alexander Wilhelm Armin Kellner _____________________________________ Prof. Dr. Antonio Roberto Saad _____________________________________ Marco Brandalise de Andrade - aluno - Rio Claro, ____ de ________________ de 2005 Resultado: _____________________________________________________________ AGRADECIMENTOS Ao longo do desenvolvimento deste Mestrado, em seu caminho paradoxalmente tão longo e tão breve, contei com o apoio de um grande número de pessoas. Algumas destas contribuíram de forma direta, enquanto outras sequer estavam cientes de sua importância ao longo deste processo, em uma grande diversidade de aspectos. Agradeço em primeiro lugar ao Prof. Dr. Reinaldo José Bertini, por sua orientação e pela grande atenção e paciência a este estudante, me recebendo sem reservas na cidade de Rio Claro e na UNESP, bem como na comunidade científica. Agradeço à Universidade Estadual Paulista, ao Instituto de Geociências e Ciências Exatas e ao Departamento de Geologia Aplicada, em cujas dependências o estudo encontrou condições para o seu desenvolvimento. De fundamental importância foi o suporte fornecido pela Coordenação de Aperfeiçoamento de Pessoal de Nível Superior (CAPES), que forneceu uma Bolsa de Mestrado que em muito ajudou a ampliar o potencial e o alcance deste trabalho. Agradeço ao Prof. Dr. Antonio Roberto Saad e a Profa.
    [Show full text]
  • New Transitional Fossil from Late Jurassic of Chile Sheds Light on the Origin of Modern Crocodiles Fernando E
    www.nature.com/scientificreports OPEN New transitional fossil from late Jurassic of Chile sheds light on the origin of modern crocodiles Fernando E. Novas1,2, Federico L. Agnolin1,2,3*, Gabriel L. Lio1, Sebastián Rozadilla1,2, Manuel Suárez4, Rita de la Cruz5, Ismar de Souza Carvalho6,8, David Rubilar‑Rogers7 & Marcelo P. Isasi1,2 We describe the basal mesoeucrocodylian Burkesuchus mallingrandensis nov. gen. et sp., from the Upper Jurassic (Tithonian) Toqui Formation of southern Chile. The new taxon constitutes one of the few records of non‑pelagic Jurassic crocodyliforms for the entire South American continent. Burkesuchus was found on the same levels that yielded titanosauriform and diplodocoid sauropods and the herbivore theropod Chilesaurus diegosuarezi, thus expanding the taxonomic composition of currently poorly known Jurassic reptilian faunas from Patagonia. Burkesuchus was a small‑sized crocodyliform (estimated length 70 cm), with a cranium that is dorsoventrally depressed and transversely wide posteriorly and distinguished by a posteroventrally fexed wing‑like squamosal. A well‑defned longitudinal groove runs along the lateral edge of the postorbital and squamosal, indicative of a anteroposteriorly extensive upper earlid. Phylogenetic analysis supports Burkesuchus as a basal member of Mesoeucrocodylia. This new discovery expands the meagre record of non‑pelagic representatives of this clade for the Jurassic Period, and together with Batrachomimus, from Upper Jurassic beds of Brazil, supports the idea that South America represented a cradle for the evolution of derived crocodyliforms during the Late Jurassic. In contrast to the Cretaceous Period and Cenozoic Era, crocodyliforms from the Jurassic Period are predomi- nantly known from marine forms (e.g., thalattosuchians)1.
    [Show full text]
  • Novos Sítios Fossilíferos Da Formação Romualdo
    Anuário do Instituto de Geociências - UFRJ www.anuario.igeo.ufrj.br Novos Sítios Fossilíferos da Formação Romualdo, Cretáceo Inferior, Bacia do Araripe, Exu, Pernambuco, Nordeste do Brasil New Fossiliferous Sites of the Romualdo Formation, Lower Cretaceous, Araripe Basin, Exu, Pernambuco, Northeast of Brazil Rudah Ruano Cavalcanti Duque & Alcina Magnólia Franca Barreto Universidade Federal de Pernambuco, Centro de Tecnologia e Geociências, Departamento de Geologia, Av. Acadêmico Hélio Ramos, s/n, 50740-530, Cidade Universitária, Recife, PE, Brasil. E-mails: [email protected]; [email protected] Recebido em: 10/01/2018 Aprovado em:01/03/2018 DOI: http://dx.doi.org/10.11137/2018_1_05_14 Resumo Aqui são apresentados antigos e novos sítios fossilíferos da Formação Romualdo, Albiano da Bacia do Araripe, no município de Exu, Pernambuco, Nordeste do Brasil, com o levantamento da sua diversidade fossilífera, com ênfase em paleovertebrados. Foram estudadas sete localidades da porção sudoeste da Bacia Sedimentar do Araripe, sendo identificados 16 táxons de paleovertebrados, incluindo peixes: Vinctifer comptoni, Rhacolepis buccalis, Calamopleurus cylindricus, Cladocyclus gardneri, Neoproscinetes penalvai, Paraelops cearenses, Tharrhias araripis, Notelops brama, Araripelepdotes temnurus, Brannerion sp. e Beurlenichthys ouricuriensis; pterossauros Anhangueridae e quelônios (Araripemys barretoi). A Paleofauna de vertebrados da Formação Romualdo vem sendo estudada desde 1800, é reconhecida internacionalmente como Fossillagerstätte. Porém, a
    [Show full text]
  • Razanandrongobe Sakalavae, a Gigantic Mesoeucrocodylian from the Middle Jurassic of Madagascar, Is the Oldest Known Notosuchian
    Razanandrongobe sakalavae, a gigantic mesoeucrocodylian from the Middle Jurassic of Madagascar, is the oldest known notosuchian peerj.com /articles/3481/ Introduction A decade ago, Maganuco, Dal Sasso & Pasini (2006) described the fragmentary remains of a very large predatory archosaur from the Middle Jurassic (Bathonian) of the Mahajanga Basin, Madagascar. The material included a fragmentary right maxilla bearing three teeth, and seven peculiar isolated teeth clearly belonging to the same taxon. In spite of the scanty remains, the presence of a unique combination of features, which included a well-developed bony palate on the maxilla, mesial and lateral teeth respectively U-shaped and sub-oval in cross-section, and very large tooth denticles (1 per mm) on the carinae, allowed the authors to erect the new taxon Razanandrongobe sakalavae Maganuco, Dal Sasso & Pasini, 2006. However, the systematic position of the new species remained uncertain: indeed, besides the autapomorphic denticle size, R. sakalavae shared a mix of potential autapomorphic, synapomorphic, and homoplasic features with crocodylomorphs and theropods. Therefore, the species was referred to Archosauria incertae sedis. Here we describe new cranial material referable to R. sakalavae and consisting of an almost complete right premaxilla, the rostral half of a left dentary, a maxillary fragment with diagnostic teeth, and a very large isolated tooth crown. In addition, we tentatively refer to the same taxon five cranial fragments that were likely collected at the same locality.
    [Show full text]
  • A New Sebecid Mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of North-Western Argentina
    Zoological Journal of the Linnean Society, 2011, 163, S7–S36. With 17 figures A new sebecid mesoeucrocodylian from the Rio Loro Formation (Palaeocene) of north-western Argentina DIEGO POL1* and JAIME E. POWELL2 1CONICET, Museo Paleontológico Egidio Feruglio, Ave. Fontana 140, Trelew CP 9100, Chubut, Argentina 2CONICET, Instituto Miguel Lillo, Miguel Lillio 205, San Miguel de Tucumán CP 4000, Tucumán, Argentina Received 2 March 2010; revised 10 October 2010; accepted for publication 19 October 2010 A new basal mesoeucrocodylian, Lorosuchus nodosus gen. et sp. nov., from the Palaeocene of north-western Argentina is presented here. The new taxon is diagnosed by the presence of external nares facing dorsally, completely septated, and retracted posteriorly, elevated narial rim, sagittal crest on the anteromedial margins of both premaxillae, dorsal crests and protuberances on the anterior half of the rostrum, and anterior-most three maxillary teeth with emarginated alveolar margins. This taxon is most parsimoniously interpreted as a bizarre and highly autapomorphic basal member of Sebecidae, a position supported (amongst other characters) by the elongated bar-like pterygoid flanges, a laterally opened notch and fossa in the pterygoids located posterolaterally to the choanal opening (parachoanal fossa), base of postorbital process of jugal directed dorsally, and palatal parts of the premaxillae meeting posteriorly to the incisive foramen. Lorosuchus nodosus also shares with basal neosuchians a suite of derived characters that are interpreted as convergently acquired and possibly related to their semiaquatic lifestyle. The phylogenetic analysis used for testing the phylogenetic affinities of L. nodosus depicts Sebecidae as the sister group of Baurusuchidae, forming a monophyletic Sebecosuchia that is deeply nested within Notosuchia.
    [Show full text]